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1

Li, Qiang, and Ran Wei. "Comparison of Boraginales Plastomes: Insights into Codon Usage Bias, Adaptive Evolution, and Phylogenetic Relationships." Diversity 14, no. 12 (2022): 1104. http://dx.doi.org/10.3390/d14121104.

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The Boraginales (Boraginaceae a.l.) comprise more than 2450 species worldwide. However, little knowledge exists of the characteristics of the complete plastid genome. In this study, three new sequences representing the first pt genome of Heliotropiaceae and Cordiaceae were assembled and compared with other Boraginales species. The pt genome sizes of Cordia dichotoma, Heliotropium arborescens, and Tournefortia montana were 151,990 bp, 156,243 bp, and 155,891 bp, respectively. Multiple optimal codons were identified, which may provide meaningful information for enhancing the gene expression of B
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Miller, James S. "New Boraginales from tropical America 8: Nomenclatural notes on Varronia (Cordiaceae: Boraginales)." Brittonia 65, no. 3 (2013): 342–44. http://dx.doi.org/10.1007/s12228-012-9283-8.

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Miller, James S., and Marc Gottschling. "Generic Transfers in Malagasy Boraginales." Candollea 72, no. 2 (2017): 329–32. https://doi.org/10.15553/c2017v722a9.

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Miller, James S., and Holly A. Porter Morgan. "Assessing the effectiveness of Madagascar’s changing protected areas system: a case study of threatened Boraginales." Oryx 45, no. 2 (2011): 201–9. http://dx.doi.org/10.1017/s0030605310000803.

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AbstractThreat analyses of the Boraginales were conducted and used to assess the effectiveness of Madagascar’s current and proposed protected area systems in conserving the threatened species of a group of plants widespread in Madagascar. Specimen locality data for 52 species of four families of Boraginales were analysed to provisionally assign species to IUCN Red List categories. Six species were excluded from these global analyses as they are non-native and introduced. IUCN’s criterion B, analysis of geographical range, was found to be the most reliable means of estimating threat, and predic
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Luebert, Federico, Lorenzo Cecchi, Michael W. Frohlich, et al. "Familial classification of the Boraginales." Taxon 65, no. 3 (2016): 502–22. http://dx.doi.org/10.12705/653.5.

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Alawfi, Mohammad S., and Enas J. Albokhari. "Comparative Chloroplast Genomics Reveals a Unique Gene Inversion in Two Cordia Trees (Cordiaceae)." Forests 14, no. 9 (2023): 1778. http://dx.doi.org/10.3390/f14091778.

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Cordiaceae is a family comprising more than 400 species in the order Boraginales. The classification of this family has undergone changes over time, transitioning between family and subfamily status. In the present study, the complete chloroplast (cp) genomes of Cordia monoica and Cordia sinensis were sequenced, and their cp genomes were then characterized, analyzed, and compared to those of closely related taxa. The lengths of the cp genomes of C. monoica and C. sinensis were 151,813 bp and 152,050 bp, respectively. Both genomes consisted of 114 genes, divided into 4 ribosomal RNA genes, 30 t
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Alawfi, Mohammad S., Dhafer A. Alzahrani, and Enas J. Albokhari. "Complete Chloroplast Genome Sequences of Two Ehretia Trees (Ehretia cymosa and Ehretia obtusifolia): Genome Structures and Phylogenetic Analysis." Forests 14, no. 7 (2023): 1486. http://dx.doi.org/10.3390/f14071486.

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Ehretiaceae is a family in the order Boraginales. It contains more than 150 species. The Ehretiaceae classification has remained elusive and changed over time from subfamily to family, or vice versa. In this paper, we sequenced, characterized, and analyzed the complete chloroplast (cp) genomes of Ehretia cymosa and Ehretia obtusifolia, and their cp genomes were compared to those of related species. The length of the chloroplast genomes of E. cymosa was 156,328 bp, whereas that of E. obtusifolia was 155,961 bp. Each genome contained 114 genes, including 80 protein-coding genes, 4 rRNA genes, an
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Nazari, Hamid, and Farrokh Ghahremaninejad. "Micromorphological analysis of leaf trichomes in Australian species of Boraginales." Turczaninowia 28, no. 2 (2025): 101–11. https://doi.org/10.14258/turczaninowia.28.2.10.

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This study investigates the leaf trichome morphology of 27 species from 23 genera across seven families of Boraginales in Australia using scanning electron microscopy (SEM). We documented a wide diversity of trichome types, including glandular, non-glandular, bifurcate, and trifurcate forms. Trichome distribution varied considerably, ranging from sparse to densely packed across leaf surfaces. While some trichomes lacked surface ornamentation, most exhibited distinct protrusions, such as round, oval, or conical structures. Additionally, while filling a gap in the understanding of Boraginales le
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Weigend, Maximilian, and Hartmut H. Hilger. "Codonaceae-a newly required family name in Boraginales." Phytotaxa 10 (October 28, 2010): 26–30. https://doi.org/10.11646/phytotaxa.10.1.3.

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Gottschling, Marc, Federico Luebert, Hartmut H. Hilger, and James S. Miller. "Molecular delimitations in the Ehretiaceae (Boraginales)." Molecular Phylogenetics and Evolution 72 (March 2014): 1–6. http://dx.doi.org/10.1016/j.ympev.2013.12.005.

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Gottschling, Marc. "Floral Ontogeny in Bourreria (Ehretiaceae, Boraginales)." Flora - Morphology, Distribution, Functional Ecology of Plants 199, no. 5 (2004): 409–23. http://dx.doi.org/10.1078/0367-2530-00169.

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Miller, James S. "A revision of Cordiaceae (Boraginales) in Central America, Mexico, and the United States." Journal of the Botanical Research Institute of Texas 17, no. 1 (2023): 39–137. http://dx.doi.org/10.17348/jbrit.v17.i1.1290.

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A taxonomic revision is provided for Cordiaceae (Boraginales) in Mexico and Central America and the adjacent portions of the United States, including 44 species of Cordia and 17 species of Varronia. Eight of the 44 species of Cordia were described as new to science elsewhere in earlier parts of this study. Cordiaceae are distinguished from other families of Boraginales in being erect woody plants with a style that is twice-divided with four stigmas. Taxonomic history of generic delimitation in the Cordiaceae is presented, along with discussions of morphology of Cordia and Varronia, an overview
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Cecchi, Lorenzo, and Federico Selvi. "Typification of names in Boraginales described from Sicily." Natural History Sciences 2, no. 2 (2015): 97–99. https://doi.org/10.4081/nhs.2015.248.

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Cecchi, Lorenzo, Selvi, Federico (2015): Typification of names in Boraginales described from Sicily. Natural History Sciences 2 (2): 97-99, DOI: 10.4081/nhs.2015.248, URL: http://dx.doi.org/10.4081/nhs.2015.248
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Wood, John R.I., and Pablo Muñoz-Rodríguez. "A new species of Euploca (Boraginales: Heliotropiaceae) from the Inter-Andean dry valleys of Bolivia." Phytotaxa 543, no. 4 (2022): 219–32. https://doi.org/10.11646/phytotaxa.543.4.2.

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Wood, John R.I., Muñoz-Rodríguez, Pablo (2022): A new species of Euploca (Boraginales: Heliotropiaceae) from the Inter-Andean dry valleys of Bolivia. Phytotaxa 543 (4): 219-232, DOI: 10.11646/phytotaxa.543.4.2
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Cohen, James I. "Phylogenomics Of Boraginaceae and Boraginales Using Lineage-Specific and Angiosperms353 Loci." Systematic Botany 49, no. 4 (2025): 749–67. https://doi.org/10.1600/036364424X17323182682762.

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Cohen, James I. (2025): Phylogenomics Of Boraginaceae and Boraginales Using Lineage-Specific and Angiosperms353 Loci. Systematic Botany (Basel, Switzerland) 49 (4): 749-767, DOI: 10.1600/036364424X17323182682762, URL: https://doi.org/10.1600/036364424x17323182682762
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Irimia, Ramona-Elena, and Marc Gottschling. "A new species of Rochefortia (Ehretiaceae, Boraginales) from the Lesser Antilles." Phytotaxa 236, no. 1 (2015): 62–70. https://doi.org/10.11646/phytotaxa.236.1.5.

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Irimia, Ramona-Elena, Gottschling, Marc (2015): A new species of Rochefortia (Ehretiaceae, Boraginales) from the Lesser Antilles. Phytotaxa 236 (1): 62-70, DOI: 10.11646/phytotaxa.236.1.5, URL: http://dx.doi.org/10.11646/phytotaxa.236.1.5
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Miller, James S. "New Boraginales from Tropical America 11: A New Species of Cordia (Boraginales: Cordiaceae) from Colombia and Venezuela." Novon, A Journal for Botanical Nomenclature 31 (July 19, 2023): 123–25. http://dx.doi.org/10.3417/2023805.

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A new species of Cordia L. (Boraginales: Cordiaceae) is described from Amazonian Venezuela and Colombia. Cordia carnevalii J. S. Mill. is distinct in its lanceolate to elliptic-lanceolate leaves borne on short petioles, and axillary inflorescences. It is vegetatively distinct from other species of Cordia in Colombia and Venezuela, and no other species in the genus has leaves that are similar. The species occurs in a poorly known region of Amazonian Colombia and Venezuela, and is presently known from only three collections made in a very restricted area, so the species is clearly of conservatio
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Melo, José Iranildo Miranda De, Renan Da Cruz Paulino, Regina Célia De Oliveira, and Diego Daltro Vieira. "Flora of Rio Grande do Norte, Brazil: Boraginales." Phytotaxa 357, no. 4 (2018): 235–60. https://doi.org/10.11646/phytotaxa.357.4.1.

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Melo, José Iranildo Miranda De, Paulino, Renan Da Cruz, Oliveira, Regina Célia De, Vieira, Diego Daltro (2018): Flora of Rio Grande do Norte, Brazil: Boraginales. Phytotaxa 357 (4): 235-260, DOI: 10.11646/phytotaxa.357.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.357.4.1
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Melo, José Iranildo Miranda De, and Diego Daltro Vieira. "A new species of Varronia (Cordiaceae) and a checklist of Boraginales for the State of Sergipe, Brazil." Phytotaxa 231, no. 2 (2015): 145–55. https://doi.org/10.11646/phytotaxa.231.2.3.

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Melo, José Iranildo Miranda De, Vieira, Diego Daltro (2015): A new species of Varronia (Cordiaceae) and a checklist of Boraginales for the State of Sergipe, Brazil. Phytotaxa 231 (2): 145-155, DOI: 10.11646/phytotaxa.231.2.3, URL: http://dx.doi.org/10.11646/phytotaxa.231.2.3
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Gottschling, Marc, and Ramona-Elena Irimia. "Taxonomic revision of Rochefortia Sw. (Ehretiaceae, Boraginales)." Biodiversity Data Journal 4 (June 8, 2016): e7720. http://dx.doi.org/10.3897/bdj.4.e7720.

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Kottaimuthu, R. "A new name forEhretia ovalifoliaWight (Ehretiaceae: Boraginales)." Webbia 71, no. 2 (2016): 239–40. http://dx.doi.org/10.1080/00837792.2016.1221175.

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DIANE, N., H. H. HILGER, and M. GOTTSCHLING. "Transfer cells in the seeds of Boraginales." Botanical Journal of the Linnean Society 140, no. 2 (2002): 155–64. http://dx.doi.org/10.1046/j.1095-8339.2002.00085.x.

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Irimia, Ramona-Elena, and Marc Gottschling. "Taxonomic revision of Rochefortia Sw. (Ehretiaceae, Boraginales)." Biodiversity Data Journal 4 (June 8, 2016): e7720. https://doi.org/10.3897/BDJ.4.e7720.

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<i>Rochefortia</i> is a small taxon of woody plants in the Ehretiaceae (Boraginales) exhibiting coriaceous leaves with cystoliths, small whitish flowers and drupaceous fruits containing four pyrenes. It shares the dioecious sex distribution with its sister group <i>Lepidocordia</i> and can be delimited from the latter (and all other Ehretiaceae) by the presence of thorns. Neotropical <i>Rochefortia</i> is distributed over most Caribbean islands, Central America and northern South America. Twenty-eight validly published names (corresponding to twenty-one typified taxa at the species level and b
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Pätzold, Janin, Benedikt Feyrer, Johanna Saumweber, Hartmut H. Hilger, and Marc Gottschling. "Observations on flower and fruit anatomy in dioecious species of Cordia (Cordiaceae, Boraginales) with evolutionary interpretations." Organisms Diversity & Evolution 23, no. 1 (2022): 77–90. https://doi.org/10.1007/s13127-022-00576-6.

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Pätzold, Janin, Feyrer, Benedikt, Saumweber, Johanna, Hilger, Hartmut H., Gottschling, Marc (2023): Observations on flower and fruit anatomy in dioecious species of Cordia (Cordiaceae, Boraginales) with evolutionary interpretations. Organisms Diversity &amp; Evolution 23 (1): 77-90, DOI: 10.1007/s13127-022-00576-6, URL: http://dx.doi.org/10.1007/s13127-022-00576-6
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Heigl, Heidi M.L., Juliane Kretschmann, Hartmut H. Hilger, and Marc Gottschling. "Flower and fruit anatomy of Cordia nodosa Lam. and Varronia bonplandii Desv. (Cordiaceae, Boraginales) with phylogenetic implications." Organisms Diversity & Evolution 20, no. 1 (2019): 9–24. https://doi.org/10.1007/s13127-019-00420-4.

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Heigl, Heidi M.L., Kretschmann, Juliane, Hilger, Hartmut H., Gottschling, Marc (2020): Flower and fruit anatomy of Cordia nodosa Lam. and Varronia bonplandii Desv. (Cordiaceae, Boraginales) with phylogenetic implications. Organisms Diversity &amp; Evolution (New York, N.Y.) 20 (1): 9-24, DOI: 10.1007/s13127-019-00420-4, URL: http://dx.doi.org/10.1007/s13127-019-00420-4
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Cecchi, Lorenzo, and Federico Selvi. "Typification of names in Boraginales described from Sicily." Natural History Sciences 2, no. 2 (2015): 97. http://dx.doi.org/10.4081/nhs.2015.248.

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Eight names in &lt;em&gt;Boraginales&lt;/em&gt; (&lt;em&gt;Boraginaceae&lt;/em&gt; s.l.) described from Sicily between 1814 and 1919 are typified in the framework of the&lt;em&gt; Flora Critica d’Italia&lt;/em&gt; and &lt;em&gt;Loci classici&lt;/em&gt; project. Some critical aspects are briefly discussed to clarify the circumstances that led to the choice of the lectotypes and the current taxonomic status of the taxa.
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MIRANDA DE MELO, JOSÉ IRANILDO, RENAN DA CRUZ PAULINO, REGINA CÉLIA DE OLIVEIRA, and DIEGO DALTRO VIEIRA. "Flora of Rio Grande do Norte, Brazil: Boraginales." Phytotaxa 357, no. 4 (2018): 235. http://dx.doi.org/10.11646/phytotaxa.357.4.1.

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A taxonomic treatment of the Boraginales of Rio Grande do Norte, Brazil, is presented. Twenty-one species in five genera and two families have been recorded: Cordiaceae, with two genera (Cordia (seven species) and Varronia (four)) and Heliotropiaceae, with three genera (Euploca (five), Heliotropium (two) and Myriopus (three)). Cordia oncocalyx and Euploca paradoxa represents new occurrence records for the flora of Rio Grande do Norte, the latter being found for the first time also in the littoral vegetation. Keys to the families and respective genera and species, descriptions and illustrations
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Gottschling, Marc, Sebastian Nagelmüller, and Hartmut H. Hilger. "Generative ontogeny inTiquilia(Ehretiaceae: Boraginales) and phylogenetic implications." Biological Journal of the Linnean Society 112, no. 3 (2014): 520–34. http://dx.doi.org/10.1111/bij.12266.

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Weigend, M., and H. H. Hilger. "Codonaceae-a newly required family name in Boraginales." Phytotaxa 10, no. 1 (2014): 26. http://dx.doi.org/10.11646/phytotaxa.10.1.3.

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Walden, Genevieve K., Laura M. Garrison, and Robert Patterson. "Nomenclatural Adjustments in Phacelia sect. Glandulosae (Hydrophyllaceae, Boraginales)." Western North American Naturalist 76, no. 3 (2016): 287–90. http://dx.doi.org/10.3398/064.076.0305.

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Melo, José Iranildo Miranda de, Guilherme M. Antar, Luan Pedro da Silva, and Thaynara de Sousa Silva. "New records in Brazil reveal a disjunct distribution for Cordia weddellii (Cordiaceae)." Darwiniana, nueva serie 9, no. 2 (2021): 320–28. http://dx.doi.org/10.14522/darwiniana.2021.92.964.

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Cordia weddellii, a species of Cordia sect. Cordia (Cordiaceae, Boraginales) endemic to South America is recorded for the first time in Brazil, growing in Cerrado vegetation in Bahia and Tocantins states. A distribution map, photographs, comments on the habitat and phenology, and a preliminary conservation assessment are provided for the species. Additionally, we provide a lectotype for C. weddellii.
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Tang, Cheng‐Yi, Song Li, Yun‐Tong Wang, and Xi Wang. "Comparative genome/transcriptome analysis probes Boraginales' phylogenetic position, WGDs in Boraginales, and key enzyme genes in the alkannin/shikonin core pathway." Molecular Ecology Resources 20, no. 1 (2019): 228–41. http://dx.doi.org/10.1111/1755-0998.13104.

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Gottschling, Marc, and James S. Miller. "A REVISION OF BOURRERIA (BORAGINALES, EHRETIACEAE) IN SOUTH AMERICA1." Annals of the Missouri Botanical Garden 94, no. 4 (2007): 734–44. http://dx.doi.org/10.3417/0026-6493(2007)94[734:arobbe]2.0.co;2.

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de Stapf, María Natividad Sánchez, Neusa Taroda Ranga, and Tânia Reginados Santos Silva. "A New Species of Cordia (Cordiaceae, Boraginales) from Brazil." Novon: A Journal for Botanical Nomenclature 20, no. 2 (2010): 212–14. http://dx.doi.org/10.3417/2007099.

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de Sousa Silva, Thaynara, Luan Pedro da Silva, and José Iranildo Miranda de Melo. "Nomenclatural Updates in Varronia (Cordiaceae, Boraginales) from South America." Harvard Papers in Botany 24, no. 1 (2019): 25. http://dx.doi.org/10.3100/hpib.v24iss1.2019.n4.

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Rueangsawang, Kanokorn, Manop Poopath, and Pranom Chantaranothai. "A new record of Ehretia (Ehretiaceae, Boraginales) for Thailand." Thai Forest Bulletin (Botany) 47 (2019): 34–37. http://dx.doi.org/10.20531/tfb.2019.47.1.07.

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Weigend, Maximilian, Federico Luebert, Marc Gottschling, Thomas L. P. Couvreur, Hartmut H. Hilger, and James S. Miller. "From capsules to nutlets-phylogenetic relationships in the Boraginales." Cladistics 30, no. 5 (2013): 508–18. http://dx.doi.org/10.1111/cla.12061.

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Véliz Pérez, Mario Esteban, Ma Goreti Campos, and J. S. Miller. "Especie nueva del género Bourreria (Ehretiaceae, Boraginales) de Mesoamérica." Brittonia 61, no. 3 (2009): 237–40. http://dx.doi.org/10.1007/s12228-009-9080-1.

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Miller, James S., Colin Robinson, and Victoria Patrick. "New Boraginales from Tropical America 10: New Species of Cordia from Colombia, Ecuador, and Peru." Novon, A Journal for Botanical Nomenclature 31 (July 18, 2023): 118–22. http://dx.doi.org/10.3417/2023799.

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Two new species of Cordia L. (Boraginales: Cordiaceae), C. caudata J. S. Mill. and C. caucana J. S. Mill., are described from Colombia, Ecuador, and Peru. Both species are quite distinctive morphologically, but their relationships are not evident, so they are compared with the two other species known from Colombia with which they are most likely to be confused. Both new species are apparently uncommon and have geographically restricted ranges and are of conservation concern.
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Miranda de Melo, José Iranildo, and Diego Daltro Vieira. "A new species of Varronia (Cordiaceae) and a checklist of Boraginales for the State of Sergipe, Brazil." Phytotaxa 231, no. 2 (2015): 145. http://dx.doi.org/10.11646/phytotaxa.231.2.3.

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Varronia johnstoniana, a new species of Cordiaceae is described for Sergipe State, Brazil. This new taxon is characterized principally by having sessile, shiny leaves, corolla with rufescent indumentum, and verrucose fruits. Illustrations, data concerning its habitats, distribution, reproductive phenology, and comments about taxonomic affinities based on morphology are presented. A checklist of Boraginales in Sergipe State is presented, comprising three families, seven genera, and 28 species, of which ten species constituting new occurrences, with their respective vegetation types.
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Feuillet, Christian. "Lectotypifications of six taxa in the Boraginales (Cordiaceae and Heliotropiaceae)." PhytoKeys 62 (April 15, 2016): 95–100. http://dx.doi.org/10.3897/phytokeys.62.6259.

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Celestino de Oliveira Chagas, Earl, and James Lucas da Costa-Lima. "A New Species of Varronia (Cordiaceae, Boraginales) from Northeastern Brazil." Systematic Botany 43, no. 4 (2018): 1026–29. http://dx.doi.org/10.1600/036364418x697760.

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SILVA, THAYNARA DE SOUSA, and JOSÉ IRANILDO MIRANDA DE MELO. "New synonym, new combination and typifications in Varronia (Cordiaceae, Boraginales)." Phytotaxa 411, no. 4 (2019): 293–300. http://dx.doi.org/10.11646/phytotaxa.411.4.4.

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Due to their recent reestablishment at the generic level, some species of the genus Varronia need to be (re-)allocated and their names and respective nomenclature types reviewed. One new combination (V. serratifolia) and a new synonym (V. intonsa) are proposed here. To promote nomenclatural stability, the historical of the names of widely distribute species is discussed and a lectotype is assigned to V. globosa, a lectotype and an epitype for V. curassavica. The names V. hermanniifolia, C. hermanniifolia var. calycina, Cordia sessilifolia var. macrantha, and V. villicaulis are also lectotypifi
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de Stapf, María Natividad Sánchez, and Tânia Regina dos Santos Silva. "Four new species of genus Cordia (Cordiaceae, Boraginales) from Brazil." Brittonia 65, no. 2 (2013): 191–99. http://dx.doi.org/10.1007/s12228-012-9268-7.

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Carlquist, Sherwin, and C. Matt Guilliams. "Distinctive wood anatomy of the root-parasitic family Lennoaceae (Boraginales)." IAWA Journal 38, no. 1 (2017): 3–12. http://dx.doi.org/10.1163/22941932-20170153.

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The four species of Lennoaceae have strands of primary plus secondary xylem in a background of starch-rich parenchyma. These strands constitute a cylinder with large primary rays. The wood within these strands is markedly different from that of other families in the crown group of Boraginales such as Cordiaceae and Ehretiaceae, most of which are woody. Lennoaceae differ because they lack fibrous cells (libriform fibers), lack rays within the vascular strands, and have markedly elliptical vessel-to-vessel pits without vestures. Lennoaceae have secondary xylem with short, wide vessel elements wi
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46

Feuillet, Christian. "Lectotypifications of six taxa in the Boraginales (Cordiaceae and Heliotropiaceae)." PhytoKeys 62 (April 15, 2016): 95–100. https://doi.org/10.3897/phytokeys.62.6259.

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A large number of specimens used as original material for the description of new species were destroyed in the bombing of the Berlin-Dahlem herbarium, B, in 1943. Six lectotypes are designated here for Cordia discolor Cham., Cordia multispicata Cham., Cordia tobagensis Urb. and its variety broadwayi Urb. in the Cordiaceae and for Tournefortia paniculata Cham. and Tournefortia ulei Vaupel in the Heliotropiaceae.
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Hilger, Hartmut H., and Marc Gottschling. "Phylogenetic analysis and character evolution of Ehretia and Bourreria (Ehretiaceae, Boraginales) - correction." Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 124, no. 2 (2002): 149–50. http://dx.doi.org/10.1127/0006-8152/2002/0124-0149.

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48

Mirra, Fabiana, Marc Gottschling, and Pablo Moroni. "Revisión taxonómica del género Cortesia (Ehretiaceae, Boraginales), un género endémico de la Argentina." Darwiniana, nueva serie 10, no. 1 (2022): 279–87. http://dx.doi.org/10.14522/darwiniana.2022.101.1034.

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Cortesia is a halophytic and monotypic genus endemic to Argentina that belongs to Ehretiaceae (Boraginales). It is morphologically distinct from the other members of the family in having cuneate leaves with a 3-dentate apex, an involucrum of fused bracts, spatulate calyx lobes, and white drupes with a two-parted endocarp. Its single species, Cortesia cuneifolia, is here treated and illustrated based on the study of numerous herbarium collections. Additionally, a lectotype is designated for the name to assure its reliable application. Furthermore, the species is recorded for the first time in T
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49

Cohen, James I. "Phylogenomics Of Boraginaceae and Boraginales Using Lineage-Specific and Angiosperms353 Loci." Systematic Botany 49, no. 4 (2025): 749–67. https://doi.org/10.1600/036364424x17323182682762.

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Abstract— During the past 20 yr, the phylogenetics of Boraginaceae has taken shape using plastid DNA regions and the nuclear ribosomal internal transcribed spacer (ITS), but these regions only reflect a limited understanding of the evolutionary history of the family. Using hybridization-enrichment sequencing, 531 nuclear regions from lineage-specific and Angiosperms353 loci were sequenced and aligned for 49 species from across Boraginaceae. Additionally, the Angiosperms353 loci were incorporated with a broader dataset of the same loci from 115 accessions of Boraginales and relatives. Based on
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Gottschling, Marc, and Hartmut H. Hilger. "Characterisation of a Novel Fruit Type Found in Ehretia (Ehretiaceae, Boraginales)." Blumea - Biodiversity, Evolution and Biogeography of Plants 49, no. 1 (2004): 145–53. http://dx.doi.org/10.3767/000651904x486250.

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