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1

Gavilán, Brenda, Elena Perea-Atienza, and Pedro Martínez. "Xenacoelomorpha: a case of independent nervous system centralization?" Philosophical Transactions of the Royal Society B: Biological Sciences 371, no. 1685 (2016): 20150039. http://dx.doi.org/10.1098/rstb.2015.0039.

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Centralized nervous systems (NSs) and complex brains are among the most important innovations in the history of life on our planet. In this context, two related questions have been formulated: How did complex NSs arise in evolution, and how many times did this occur? As a step towards finding an answer, we describe the NS of several representatives of the Xenacoelomorpha, a clade whose members show different degrees of NS complexity. This enigmatic clade is composed of three major taxa: acoels, nemertodermatids and xenoturbellids. Interestingly, while the xenoturbellids seem to have a rather ‘
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Arroyo, Alicia S., David López-Escardó, Colomban de Vargas, and Iñaki Ruiz-Trillo. "Hidden diversity of Acoelomorpha revealed through metabarcoding." Biology Letters 12, no. 9 (2016): 20160674. http://dx.doi.org/10.1098/rsbl.2016.0674.

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Animals with bilateral symmetry comprise the majority of the described species within Metazoa. However, the nature of the first bilaterian animal remains unknown. As most recent molecular phylogenies point to Xenacoelomorpha as the sister group to the rest of Bilateria, understanding their biology, ecology and diversity is key to reconstructing the nature of the last common bilaterian ancestor (Urbilateria). To date, sampling efforts have focused mainly on coastal areas, leaving potential gaps in our understanding of the full diversity of xenacoelomorphs. We therefore analysed 18S rDNA metabar
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Cannon, Johanna Taylor, Bruno Cossermelli Vellutini, Julian Smith, Fredrik Ronquist, Ulf Jondelius, and Andreas Hejnol. "Xenacoelomorpha is the sister group to Nephrozoa." Nature 530, no. 7588 (2016): 89–93. http://dx.doi.org/10.1038/nature16520.

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Haszprunar, Gerhard. "Review of data for a morphological look on Xenacoelomorpha (Bilateria incertae sedis)." Organisms Diversity & Evolution 16, no. 2 (2015): 363–89. https://doi.org/10.1007/s13127-015-0249-z.

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Haszprunar, Gerhard (2016): Review of data for a morphological look on Xenacoelomorpha (Bilateria incertae sedis). Organisms Diversity & Evolution (New York, N.Y.) 16 (2): 363-389, DOI: 10.1007/s13127-015-0249-z, URL: http://dx.doi.org/10.1007/s13127-015-0249-z
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Ceccolini, Filippo, and Fabio Cianferoni. "A replacement name for Pelophila Dörjes, 1968 (Xenacoelomorpha: Acoela, Convolutidae)." Graellsia 78, no. 1 (2022): e165. http://dx.doi.org/10.3989/graellsia.2022.v78.339.

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Se ha detectado un homónimo más moderno entre los Acoela (Xenacoelomorpha, Acoelomorpha) y se propone el siguiente nombre sustitutivo: Acoelopelophila Ceccolini & Cianferoni nom. nov. pro Pelophila Dörjes, 1968 nec Dejean, 1821. Se da también la siguiente nueva combinación: Acoelopelophila lutheri (Westblad, 1946) comb. nov.
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TYLER, SETH, and STEPHEN SCHILLING. "Phylum Xenacoelomorpha Philippe, et al., 2011. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness." Zootaxa 3148, no. 1 (2011): 24–25. https://doi.org/10.11646/zootaxa.3148.1.6.

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TYLER, SETH, SCHILLING, STEPHEN (2011): Phylum Xenacoelomorpha Philippe, et al., 2011. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148 (1): 24-25, DOI: 10.11646/zootaxa.3148.1.6, URL: http://dx.doi.org/10.11646/zootaxa.3148.1.6
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Çinar, Melih Ertan. "Checklist of the phyla Platyhelminthes, Xenacoelomorpha, Nematoda, Acanthocephala, Myxozoa, Tardigrada, Cephalorhyncha, Nemertea, Echiura, Brachiopoda, Phoronida, Chaetognatha, and Chordata (Tunicata, Cephalochordata,." Turkish Journal of Zoology 38, no. 6 (2014): 698–722. https://doi.org/10.3906/zoo-1405-70.

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Çinar, Melih Ertan (2014): Checklist of the phyla Platyhelminthes, Xenacoelomorpha, Nematoda, Acanthocephala, Myxozoa, Tardigrada, Cephalorhyncha, Nemertea, Echiura, Brachiopoda, Phoronida, Chaetognatha, and Chordata (Tunicata, Cephalochordata,. Turkish Journal of Zoology 38 (6): 698-722, DOI: 10.3906/zoo-1405-70, URL: http://dx.doi.org/10.3906/zoo-1405-70
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Kapli, Paschalia, and Maximilian J. Telford. "Topology-dependent asymmetry in systematic errors affects phylogenetic placement of Ctenophora and Xenacoelomorpha." Science Advances 6, no. 50 (2020): eabc5162. http://dx.doi.org/10.1126/sciadv.abc5162.

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The evolutionary relationships of two animal phyla, Ctenophora and Xenacoelomorpha, have proved highly contentious. Ctenophora have been proposed as the most distant relatives of all other animals (Ctenophora-first rather than the traditional Porifera-first). Xenacoelomorpha may be primitively simple relatives of all other bilaterally symmetrical animals (Nephrozoa) or simplified relatives of echinoderms and hemichordates (Xenambulacraria). In both cases, one of the alternative topologies must be a result of errors in tree reconstruction. Here, using empirical data and simulations, we show tha
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Perea-Atienza, E., B. Gavilan, M. Chiodin, et al. "The nervous system of Xenacoelomorpha: a genomic perspective." Journal of Experimental Biology 218, no. 4 (2015): 618–28. http://dx.doi.org/10.1242/jeb.110379.

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Ceccolini, Filippo, and Fabio Cianferoni. "A replacement name for Pelophila Dörjes, 1968 (Xenacoelomorpha: Acoela, Convolutidae)." Graellsia 78, no. 1 (2022): e165 [2 pp.]. https://doi.org/10.3989/graellsia.2022.v78.339.

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The following replacement name in Acoela (Xenacoelomorpha, Acoelomorpha) is proposed: <em>Acoelopelophila</em> Ceccolini &amp; Cianferoni <strong>nom. nov.</strong> pro <em>Pelophila</em> D&ouml;rjes, 1968 nec Dejean, 1821. Further, the following new combination is also established: <em>Acoelopelophila lutheri</em> (Westblad, 1946) <strong>comb. nov.</strong>
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Querido, Marcelo C., Arthur Z. Güth, Amana G. Garrido, et al. "The photosymbiotic acoel Convolutriloba retrogemma (Xenacoelomorpha) is sensitive to thermal stress." Journal of Experimental Marine Biology and Ecology 582 (January 2025): 152079. https://doi.org/10.1016/j.jembe.2024.152079.

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12

Rouse, Greg W., Nerida G. Wilson, Jose I. Carvajal, and Robert C. Vrijenhoek. "New deep-sea species of Xenoturbella and the position of Xenacoelomorpha." Nature 530, no. 7588 (2016): 94–97. http://dx.doi.org/10.1038/nature16545.

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Ono, Rintaro, and Hiroshi Kajihara. "Description of a new species of Amphiscolops (Xenacoelomorpha: Convolutidae)." Journal of Natural History 59, no. 21-24 (2025): 1611–24. https://doi.org/10.1080/00222933.2025.2489502.

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14

Haszprunar, Gerhard. "Review of data for a morphological look on Xenacoelomorpha (Bilateria incertae sedis)." Organisms Diversity & Evolution 16, no. 2 (2015): 363–89. http://dx.doi.org/10.1007/s13127-015-0249-z.

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Buckland-Nicks, John, Kennet Lundin, and Andreas Wallberg. "The sperm of Xenacoelomorpha revisited: implications for the evolution of early bilaterians." Zoomorphology 138, no. 1 (2018): 13–27. http://dx.doi.org/10.1007/s00435-018-0425-8.

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Dittmann, Isabel L., Thomas Zauchner, Lucy M. Nevard, Maximilian J. Telford, and Bernhard Egger. "SALMFamide2 and serotonin immunoreactivity in the nervous system of some acoels (Xenacoelomorpha)." Journal of Morphology 279, no. 5 (2018): 589–97. http://dx.doi.org/10.1002/jmor.20794.

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Martinez, P., E. Perea-Atienza, B. Gavilán, C. Fernandez, and S. Sprecher. "The study of xenacoelomorph nervous systems. Molecular and morphological perspectives." Invertebrate Zoology 14, no. 1 (2017): 32–44. https://doi.org/10.15298/invertzool.14.1.06.

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Martinez, P., Perea-Atienza, E., Gavilán, B., Fernandez, C., Sprecher, S. (2017): The study of xenacoelomorph nervous systems. Molecular and morphological perspectives. Invertebrate Zoology 14 (1): 32-44, DOI: 10.15298/invertzool.14.1.06, URL: https://doi.org/10.15298/invertzool.14.1.06
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Philippe, Hervé, Albert J. Poustka, Marta Chiodin, et al. "Mitigating Anticipated Effects of Systematic Errors Supports Sister-Group Relationship between Xenacoelomorpha and Ambulacraria." Current Biology 29, no. 11 (2019): 1818–26. http://dx.doi.org/10.1016/j.cub.2019.04.009.

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Ono, Rintaro, and Hiroshi Kajihara. "Phylogenetic position of the acoel <i>Oxyposthia praedator</i> (Xenacoelomorpha: Convolutidae)." Plankton and Benthos Research 20, no. 1 (2025): 101–6. https://doi.org/10.3800/pbr.20.101.

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Brauchle, Michael, Adem Bilican, Claudia Eyer, et al. "Xenacoelomorpha Survey Reveals That All 11 Animal Homeobox Gene Classes Were Present in the First Bilaterians." Genome Biology and Evolution 10, no. 9 (2018): 2205–17. http://dx.doi.org/10.1093/gbe/evy170.

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Juravel, Ksenia, Luis Porras, Sebastian Höhna, Davide Pisani, and Gert Wörheide. "Exploring genome gene content and morphological analysis to test recalcitrant nodes in the animal phylogeny." PLOS ONE 18, no. 3 (2023): e0282444. http://dx.doi.org/10.1371/journal.pone.0282444.

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An accurate phylogeny of animals is needed to clarify their evolution, ecology, and impact on shaping the biosphere. Although datasets of several hundred thousand amino acids are nowadays routinely used to test phylogenetic hypotheses, key deep nodes in the metazoan tree remain unresolved: the root of animals, the root of Bilateria, and the monophyly of Deuterostomia. Instead of using the standard approach of amino acid datasets, we performed analyses of newly assembled genome gene content and morphological datasets to investigate these recalcitrant nodes in the phylogeny of animals. We explor
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Hulett, Ryan E., Deirdre Potter, and Mansi Srivastava. "Neural architecture and regeneration in the acoel Hofstenia miamia." Proceedings of the Royal Society B: Biological Sciences 287, no. 1931 (2020): 20201198. http://dx.doi.org/10.1098/rspb.2020.1198.

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The origin of bilateral symmetry, a major transition in animal evolution, coincided with the evolution of organized nervous systems that show regionalization along major body axes. Studies of Xenacoelomorpha, the likely outgroup lineage to all other animals with bilateral symmetry, can inform the evolutionary history of animal nervous systems. Here, we characterized the neural anatomy of the acoel Hofstenia miamia . Our analysis of transcriptomic data uncovered orthologues of enzymes for all major neurotransmitter synthesis pathways. Expression patterns of these enzymes revealed the presence o
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Himmel, Nathaniel J., Thomas R. Gray, and Daniel N. Cox. "Phylogenetics Identifies Two Eumetazoan TRPM Clades and an Eighth TRP Family, TRP Soromelastatin (TRPS)." Molecular Biology and Evolution 37, no. 7 (2020): 2034–44. http://dx.doi.org/10.1093/molbev/msaa065.

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Abstract Transient receptor potential melastatins (TRPMs) are most well known as cold and menthol sensors, but are in fact broadly critical for life, from ion homeostasis to reproduction. Yet, the evolutionary relationship between TRPM channels remains largely unresolved, particularly with respect to the placement of several highly divergent members. To characterize the evolution of TRPM and like channels, we performed a large-scale phylogenetic analysis of &amp;gt;1,300 TRPM-like sequences from 14 phyla (Annelida, Arthropoda, Brachiopoda, Chordata, Cnidaria, Echinodermata, Hemichordata, Mollu
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Hayes, Matthew J., Anne-C. Zakrzewski, Timothy P. Levine, and Maximilian J. Telford. "Nucleus–Plasma Membrane Contact Sites Are Formed During Spermiogenesis in the Acoel Symsagittifera roscoffensis." Contact 3 (January 2020): 251525642092635. http://dx.doi.org/10.1177/2515256420926354.

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Symsagittifera roscoffensis is a small marine worm found in the intertidal zone of sandy beaches around the European shores of the Atlantic. S. roscoffensis is a member of the Acoelomorpha, a group of flatworms formerly classified with the Platyhelminthes, but now recognized as Xenacoelomorpha, a separate phylum of disputed affinity. We have used electron microscopy to examine the process of spermiogenesis (the final stage of spermatogenesis) in S. roscoffensis, by which spermatids form highly elongated spermatozoa. Their nuclei are long and thread-like, running most of the cell’s length, and
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Achatz, Johannes, and Pedro Martinez. "The nervous system of Isodiametra pulchra (Acoela) with a discussion on the neuroanatomy of the Xenacoelomorpha and its evolutionary implications." Frontiers in Zoology 9, no. 1 (2012): 27. http://dx.doi.org/10.1186/1742-9994-9-27.

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Atherton, Sarah, and Ulf Jondelius. "Diversity in the family Isodiametridae (Acoela): New species bring back old problems." Zootaxa 5169, no. 5 (2022): 401–24. https://doi.org/10.11646/zootaxa.5169.5.1.

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Atherton, Sarah, and Ulf Jondelius. "Phylogenetic assessment and systematic revision of the acoel family Isodiametridae." Zoological Journal of the Linnean Society 194 (December 31, 2022): 736–60. https://doi.org/10.1093/zoolinnean/zlab050.

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Atherton, Sarah, Jondelius, Ulf (2022): Phylogenetic assessment and systematic revision of the acoel family Isodiametridae. Zoological Journal of the Linnean Society 194: 736-760, DOI: 10.1093/zoolinnean/zlab050
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Graciano, Miguel Luis. "Renal Intercalated Cells: Alien Cells Inside Us?" Biology 14, no. 6 (2025): 607. https://doi.org/10.3390/biology14060607.

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Mammalian renal intercalated cells are known for their role in acid secretion and maintaining acid–base balance. Herein, we discuss the theoretical reasons behind their development based on published data, focusing on the unique characteristics of renal intercalated cell biology that distinguish them from other mammalian cell types, while simultaneously attempting to explain the persistence of cells similar to intercalated cells throughout evolution. In addition, we traced these characteristics phylogenetically back to the simplest organisms. Intercalated cells have several functions and attri
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TYLER, SETH, and STEPHEN SCHILLING. "Phylum Xenacoelomorpha Philippe, et al., 2011. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness." Zootaxa 3148, no. 1 (2011): 24. http://dx.doi.org/10.11646/zootaxa.3148.1.6.

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Gaunt, Stephen J. "Hox cluster genes and collinearities throughout the tree of animal life." International Journal of Developmental Biology 62, no. 11-12 (2018): 673–83. http://dx.doi.org/10.1387/ijdb.180162sg.

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The discovery of Hox gene clusters, first in Drosophila (a protostome) and then as homologues in vertebrates (deuterostomes), was a major step in our understanding of both developmental and evolutionary biology. Hox genes in both species perform the same overall function: that is, organization of the body along its head-tail axis. The conclusion is that the protostome-deuterostome ancestor, founder of 99% of all described animal species, must already have had this same basic Hox cluster, and that it probably used it in the same way to establish its body plan. A striking feature of Hox genes is
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Achatz, Johannes G., Robert Gschwentner, and Reinhard Rieger. "Symsagittifera smaragdina sp. nov.: A new acoel (Acoela: Acoelomorpha) from the Mediterranean Sea." Zootaxa 1085 (December 31, 2005): 33–45. https://doi.org/10.5281/zenodo.170467.

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Achatz, Johannes G., Gschwentner, Robert, Rieger, Reinhard (2005): Symsagittifera smaragdina sp. nov.: A new acoel (Acoela: Acoelomorpha) from the Mediterranean Sea. Zootaxa 1085: 33-45, DOI: 10.5281/zenodo.170467
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Achatz, Johannes G., and Matthew D. Hooge. "Convolutidae (Acoela) from Tanzania." Zootaxa 1362 (December 31, 2006): 1–21. https://doi.org/10.5281/zenodo.174702.

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Iii, Thomas Shannon, and Johannes G. Achatz. "Convolutriloba macropyga sp. nov., an uncommonly fecund acoel (Acoelomorpha) discovered in tropical aquaria." Zootaxa 1525 (December 31, 2007): 1–17. https://doi.org/10.5281/zenodo.177524.

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Iii, Thomas Shannon, Achatz, Johannes G. (2007): Convolutriloba macropyga sp. nov., an uncommonly fecund acoel (Acoelomorpha) discovered in tropical aquaria. Zootaxa 1525: 1-17, DOI: 10.5281/zenodo.177524
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Doweld, Alexander B. "Leleshusia, a new replacement name for Granulina Leleshus, 1975 (Anthozoa: Heliolitoidea) nec Jousseaume, 1888 (Gastropoda: Neogastropoda: Marginellidae)." Zootaxa 3986, no. 5 (2015): 588–90. https://doi.org/10.11646/zootaxa.3986.5.6.

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Doweld, Alexander B. (2015): Leleshusia, a new replacement name for Granulina Leleshus, 1975 (Anthozoa: Heliolitoidea) nec Jousseaume, 1888 (Gastropoda: Neogastropoda: Marginellidae). Zootaxa 3986 (5): 588-590, DOI: 10.11646/zootaxa.3986.5.6
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Hooge, Matthew D., and Seth Tyler. "Two new acoels (Acoelomorpha) of the genus Haplogonaria from the northwest Atlantic." Zootaxa 4013, no. 1 (2015): 111–19. https://doi.org/10.11646/zootaxa.4013.1.8.

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Achatz, Johannes G. "Convolutidae (Acoela) from the Andaman Sea." Zootaxa 1824 (December 31, 2008): 1–16. https://doi.org/10.5281/zenodo.274392.

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Ogunlana, Maxina V., Matthew D. Hooge, Yonas I. Tekle, Yehuda Benayahu, Orit Barneah, and Seth Tyler. "Waminoa brickneri n. sp. (Acoela: Acoelomorpha) associated with corals in the Red Sea." Zootaxa 1008, no. 1 (2005): 1–11. https://doi.org/10.11646/zootaxa.1008.1.1.

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Ogunlana, Maxina V., Hooge, Matthew D., Tekle, Yonas I., Benayahu, Yehuda, Barneah, Orit, Tyler, Seth (2005): Waminoa brickneri n. sp. (Acoela: Acoelomorpha) associated with corals in the Red Sea. Zootaxa 1008 (1): 1-11, DOI: 10.11646/zootaxa.1008.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1008.1.1
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Raikova, Olga I., Inga Meyer-Wachsmuth, and Ulf Jondelius. "The plastic nervous system of Nemertodermatida." Organisms Diversity & Evolution 16, no. 1 (2015): 85–104. https://doi.org/10.1007/s13127-015-0248-0.

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Raikova, Olga I., Meyer-Wachsmuth, Inga, Jondelius, Ulf (2016): The plastic nervous system of Nemertodermatida. Organisms Diversity &amp; Evolution (New York, N.Y.) 16 (1): 85-104, DOI: 10.1007/s13127-015-0248-0, URL: http://dx.doi.org/10.1007/s13127-015-0248-0
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ÇINAR, MELİH ERTAN, DERYA ÜRKMEZ, and BAKİ YOKEŞ. "Diversity of Platyhelminthes, Xenacoelomorpha, Nematoda, Acanthocephala, Brachiopoda, Kinorhyncha, Nemertea, Chaetognatha, Tardigrada, Gastrotricha, Rotifera, Phoronida, Echinodermata and Chordata (Tunicata, Cephalochordata and Hemichordata) from the coasts of Türkiye." Turkish Journal of Zoology 48, no. 6 (2024): 379–412. http://dx.doi.org/10.55730/1300-0179.3192.

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Hooge, Matthew D., and Seth Tyler. "Two new acoels (Acoela, Platyhelminthes) from the central coast of California." Zootaxa 131 (December 31, 2003): 1–14. https://doi.org/10.5281/zenodo.157080.

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Kånneby, Tobias, and Ulf Jondelius. "Four new species of Acoela from Chile." Zootaxa 3736, no. 5 (2013): 471–85. https://doi.org/10.11646/zootaxa.3736.5.3.

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Hooge, Matthew D., and Neil Eppinger. "New species of Acoela (Acoelomorpha) from the Gulf of California." Zootaxa 1009, no. 1 (2005): 1–14. https://doi.org/10.11646/zootaxa.1009.1.1.

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Hooge, Matthew D., Eppinger, Neil (2005): New species of Acoela (Acoelomorpha) from the Gulf of California. Zootaxa 1009 (1): 1-14, DOI: 10.11646/zootaxa.1009.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1009.1.1
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ÇINAR, Melih Ertan. "Checklist of the phyla Platyhelminthes, Xenacoelomorpha, Nematoda, Acanthocephala, Myxozoa, Tardigrada, Cephalorhyncha, Nemertea, Echiura, Brachiopoda, Phoronida, Chaetognatha, and Chordata (Tunicata, Cephalochordata, and Hemichordata) from the coasts of Turkey." TURKISH JOURNAL OF ZOOLOGY 38 (2014): 698–722. http://dx.doi.org/10.3906/zoo-1405-70.

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Achatz, Johannes G., Matthew D. Hooge, and Seth Tyler. "Convolutidae (Acoela) from Belize." Zootaxa 1479 (December 31, 2007): 35–66. https://doi.org/10.5281/zenodo.176820.

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Gavilán, B., S. G. Sprecher, V. Hartenstein, and P. Martinez. "The digestive system of xenacoelomorphs." Cell and Tissue Research 377, no. 3 (2019): 369–82. http://dx.doi.org/10.1007/s00441-019-03038-2.

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Haenel, Quiterie, Oleksandr Holovachov, Ulf Jondelius, Per Sundberg, and Sarah Bourlat. "NGS-based biodiversity and community structure analysis of meiofaunal eukaryotes in shell sand from Hållö island, Smögen, and soft mud from Gullmarn Fjord, Sweden." Biodiversity Data Journal 5 (June 8, 2017): e12731. https://doi.org/10.3897/BDJ.5.e12731.

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Aim: The aim of this study was to assess the biodiversity and community structure of Swedish meiofaunal eukaryotes using metabarcoding. To validate the reliability of the metabarcoding approach, we compare the taxonomic resolution obtained using the mitochondrial cytochrome oxidase 1 (COI) 'mini-barcode' and nuclear 18S small ribosomal subunit (18S) V1-V2 region, with traditional morphology-based identification of Xenacoelomorpha and Nematoda. Location: 30 samples were analysed from two ecologically distinct locations along the west coast of Sweden. 18 replicate samples of coarse shell sand we
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Nilsson, Karin Sara, Andreas Wallberg, and Ulf Jondelius. "New species of Acoela from the Mediterranean, the Red Sea, and the South Pacific." Zootaxa 2867 (December 31, 2011): 1–31. https://doi.org/10.5281/zenodo.277458.

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Hooge, Matthew D., and Seth Tyler. "Acoela (Acoelomorpha) from Belize." Zootaxa 1479, no. 1 (2007): 21–33. https://doi.org/10.11646/zootaxa.1479.1.3.

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Hejnol, Andreas, and Kevin Pang. "Xenacoelomorpha's significance for understanding bilaterian evolution." Current Opinion in Genetics & Development 39 (August 2016): 48–54. http://dx.doi.org/10.1016/j.gde.2016.05.019.

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Ezhova, О. V., A. I. Lukinykh, and V. V. Malakhov. "Nemertodermatid endosymbionts of deep-sea acorn worms (<i>Hemichordata, Torquaratoridae</i>)." Доклады Российской академии наук. Науки о жизни 515, no. 2 (2024): 60–63. http://dx.doi.org/10.31857/s2686738924020119.

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Worm-like endosymbionts were found in the hepatic region of a deep-sea acorn worm, a representative of the family Torquaratoridae Quatuoralisia malakhovi [Ezhova et Lukinykh, 2022] from the Bering Sea. Histological study of the symbionts allows us to attribute them to the taxon Nemertodermatida. Torquaratorids are similar in type of feeding to holothuroids, in which the xenacoelomorph endosymbionts have also been found.
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