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1

Bonis, Louis de. "New genus of amphicyonid carnivoran (Mammalia, Carnivora, Amphicyonidae) from the phosphorites of Quercy (France)." Fossil Imprint 76, no. 1 (2020): 201–8. http://dx.doi.org/10.37520/fi.2020.013.

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An isolated mandible of Carnivora (Mammalia) from the phosphorites of Quercy (France) is described as a new genus. It is compared with the amphicyonid genus Cynodictis, some primitive North American amphicyonids, and with European and North American Eocene carnivoraforms. I conclude that it is a primitive amphicyonid which may be dated to the middle or late Eocene.
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2

PEIGN, STPHANE, MANUEL J. SALESA, MAURICIO ANTN, and JORGE MORALES. "A NEW AMPHICYONINE (CARNIVORA: AMPHICYONIDAE) FROM THE UPPER MIOCENE OF BATALLONES-1, MADRID, SPAIN." Palaeontology 51, no. 4 (July 2008): 943–65. http://dx.doi.org/10.1111/j.1475-4983.2008.00788.x.

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3

Peigné, Stéphane, Yaowalak Chaimanee, Chotima Yamee, Pannipa Tian, and Jean-Jacques Jaeger. "A new amphicyonid (Mammalia, Carnivora, Amphicyonidae) from the late middle Miocene of northern Thailand and a review of the amphicyonine record in Asia." Journal of Asian Earth Sciences 26, no. 5 (April 2006): 519–32. http://dx.doi.org/10.1016/j.jseaes.2004.11.003.

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4

Tomiya, Susumu, and Zhijie Jack Tseng. "Whence the beardogs? Reappraisal of the Middle to Late Eocene ‘ Miacis ’ from Texas, USA, and the origin of Amphicyonidae (Mammalia, Carnivora)." Royal Society Open Science 3, no. 10 (October 2016): 160518. http://dx.doi.org/10.1098/rsos.160518.

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The Middle to Late Eocene sediments of Texas have yielded a wealth of fossil material that offers a rare window on a diverse and highly endemic mammalian fauna from that time in the southern part of North America. These faunal data are particularly significant because the narrative of mammalian evolution in the Paleogene of North America has traditionally been dominated by taxa that are known from higher latitudes, primarily in the Rocky Mountain and northern Great Plains regions. Here we report on the affinities of two peculiar carnivoraforms from the Chambers Tuff of Trans-Pecos, Texas, that were first described 30 years ago as Miacis cognitus and M. australis . Re-examination of previously described specimens and their inclusion in a cladistic analysis revealed the two taxa to be diminutive basal amphicyonids; as such, they are assigned to new genera Gustafsonia and Angelarctocyon , respectively. These two taxa fill in some of the morphological gaps between the earliest-known amphicyonid genus, Daphoenus , and other Middle-Eocene carnivoraforms, and lend additional support for a basal caniform position of the beardogs outside the Canoidea. The amphicyonid lineage had evidently given rise to at least five rather distinct forms by the end of the Middle Eocene. Their precise geographical origin remains uncertain, but it is plausible that southern North America served as an important stage for a very early phase of amphicyonid radiation.
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Morales, Jorge, Oldřich Fejfar, Elmar Heizmann, Jan Wagner, Alberto Valenciano, and Juan Abella. "A New Thaumastocyoninae (Amphicyonidae, Carnivora) from the Early Miocene of Tuchořice, the Czech Republic." Fossil Imprint 75, no. 3-4 (December 1, 2019): 397–411. http://dx.doi.org/10.2478/if-2019-0025.

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Abstract New Amphicyonidae fossil remains from the early Miocene site of Tuchořice (the Czech Republic) confirm the presence of a new Thaumastocyoninae taxon: Peignecyon felinoides n. gen. et n. sp. It is characterized by a peculiar combination of plesiomorphic and derived morphological traits. The new genus can be defined by a long and sharp mandible diastema, loss of mesial premolars (p2–p3), p4 with an inclined distally high main cuspid, moderate sectorial carnassial teeth, m1 with relict metaconid, and talonid and trigonid of similar width, and reduced M2 and m2. In the phylogenetic analysis the Thaumastocyoninae form a monophyletic group characterized by the start of the m2/M2 reduction, still moderate in Crassidia intermedia (von Meyer, 1849), but remarkable in the other species of the clade. Peignecyon felinoides already shows the advanced features defining the Thaumastocyoninae, and constitutes the sister group of the most specialized genera Tomocyon Viret, 1929b and Thaumastocyon Sthelin et Helbing, 1925. Consequently, it can be considered an excellent link between this group and the more primitive members of the tribe Ysengrini (Ysengrinia Ginsburg, 1966 and Crassidia Heizmannn et Kordikova, 2000). Peignecyon felinoides shows that the trend towards hypercarnivory had already emerged in the European early Miocene fauna, thus helping to understand the complex evolution of the Amphicyonidae during the Miocene.
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Stefen, Clara. "ENAMEL STRUCTURE OF ARCTOID CARNIVORA: AMPHICYONIDAE, URSIDAE, PROCYONIDAE, AND MUSTELIDAE." Journal of Mammalogy 82, no. 2 (May 2001): 450–62. http://dx.doi.org/10.1644/1545-1542(2001)082<0450:esoaca>2.0.co;2.

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7

Stefen, C. "Enamel Structure of Arctoid Carnivora: Amphicyonidae, Ursidae, Procyonidae, and Mustelidae." Journal of Mammalogy 82, no. 2 (May 18, 2001): 450–62. http://dx.doi.org/10.1093/jmammal/82.2.450.

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8

Siliceo, Gema, Mauricio Antón, Jorge Morales, and Manuel J. Salesa. "Built for Strength: Functional Insights from the Thoracolumbar and Sacrocaudal Regions of the Late Miocene Amphicyonid Magericyon anceps (Carnivora, Amphicyonidae) from Batallones-1 (Madrid, Spain)." Journal of Mammalian Evolution 27, no. 3 (August 8, 2019): 497–518. http://dx.doi.org/10.1007/s10914-019-09477-6.

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9

Bonis, Louis De. "Revival of a species of the rare European Oligocene amphicyonidGoupilictisGinsburg, 1969 (Mammalia, Carnivora, Amphicyonidae)." Journal of Vertebrate Paleontology 35, no. 5 (July 24, 2015): e969401. http://dx.doi.org/10.1080/02724634.2015.969401.

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10

Siliceo, Gema, Jorge Morales, Mauricio Antón, and Manuel J. Salesa. "New fossils of Amphicyonidae (Carnivora) from the middle Miocene (MN6) site of Carpetana (Madrid, Spain)." Geodiversitas 42, no. 15 (June 18, 2020): 223. http://dx.doi.org/10.5252/geodiversitas2020v42a15.

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11

Hunt, Robert M. "Evolution of Large Carnivores During the Mid-Cenozoic of North America: The Temnocyonine Radiation (Mammalia, Amphicyonidae)." Bulletin of the American Museum of Natural History 358 (November 2011): 1–153. http://dx.doi.org/10.1206/358.1.

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12

Hunt, Robert M., and Ellen Stepleton. "A skull of the immigrant Eurasian beardogCynelos(Carnivora, Amphicyonidae) from the early Miocene of southern California." Journal of Vertebrate Paleontology 35, no. 1 (January 2, 2015): e891229. http://dx.doi.org/10.1080/02724634.2014.891229.

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13

Jiangzuo, Qigao, Chunxiao Li, Xiaoxiao Zhang, Shiqi Wang, Je Ye, and Yu Li. "Diversity of Amphicyonidae (Carnivora, Mammalia) in the Middle Miocene Halamagai formation in Ulungur River area, Xinjiang, Northwestern China." Historical Biology 32, no. 2 (May 22, 2018): 187–202. http://dx.doi.org/10.1080/08912963.2018.1477142.

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14

Siliceo, Gema, Manuel J. Salesa, Mauricio Antón, Stéphane Peigné, and Jorge Morales. "Functional anatomy of the cervical region in the late Miocene amphicyonidMagericyon anceps(Carnivora, Amphicyonidae): implications for its feeding behaviour." Palaeontology 60, no. 3 (March 6, 2017): 329–47. http://dx.doi.org/10.1111/pala.12286.

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15

Tomiya, Susumu, and Zachary S. Morris. "REIDENTIFICATION OF LATE MIDDLE EOCENE “UINTACYON” FROM THE GALISTEO FORMATION (NEW MEXICO, U.S.A.) AS AN EARLY BEARDOG (MAMMALIA, CARNIVORA, AMPHICYONIDAE)." Breviora 567, no. 1 (May 15, 2020): 1. http://dx.doi.org/10.3099/0006-9698-567.1.1.

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16

Domingo, M. Soledad, Laura Domingo, Catherine Badgley, Oscar Sanisidro, and Jorge Morales. "Resource partitioning among top predators in a Miocene food web." Proceedings of the Royal Society B: Biological Sciences 280, no. 1750 (January 7, 2013): 20122138. http://dx.doi.org/10.1098/rspb.2012.2138.

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The exceptional fossil sites of Cerro de los Batallones (Madrid Basin, Spain) contain abundant remains of Late Miocene mammals. From these fossil assemblages, we have inferred diet, resource partitioning and habitat of three sympatric carnivorous mammals based on stable isotopes. The carnivorans include three apex predators: two sabre-toothed cats (Felidae) and a bear dog (Amphicyonidae). Herbivore and carnivore carbon isotope ( δ 13 C) values from tooth enamel imply the presence of a woodland ecosystem dominated by C 3 plants. δ 13 C values and mixing-model analyses suggest that the two sabre-toothed cats, one the size of a leopard and the other the size of a tiger, consumed herbivores with similar δ 13 C values from a more wooded portion of the ecosystem. The two sabre-toothed cats probably hunted prey of different body sizes, and the smaller species could have used tree cover to avoid encounters with the larger felid. For the bear dog, δ 13 C values are higher and differ significantly from those of the sabre-toothed cats, suggesting a diet that includes prey from more open woodland. Coexistence of the sabre-toothed cats and the bear dog was likely facilitated by prey capture in different portions of the habitat. This study demonstrates the utility of stable isotope analysis for investigating the behaviour and ecology of members of past carnivoran guilds.
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17

Argot, Christine. "Morphofunctional analysis of the postcranium ofAmphicyon major(Mammalia, Carnivora, Amphicyonidae) from the Miocene of Sansan (Gers, France) compared to three extant carnivores:Ursus arctos, Panthera leo,andCanis lupus." Geodiversitas 32, no. 1 (March 2010): 65–106. http://dx.doi.org/10.5252/g2010n1a2.

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18

Werdelin, Lars, and Scott W. Simpson. "The last amphicyonid (Mammalia, Carnivora) in Africa." Geodiversitas 31, no. 4 (December 2009): 775–87. http://dx.doi.org/10.5252/g2009n4a775.

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19

Morlo, Michael, Ellen R. Miller, Katharina Bastl, Mohamed K. Abdelgawad, Mohammed Hamdan, Ahmed N. El-Barkooky, and Doris Nagel. "New Amphicyonids (Mammalia, Carnivora) from Moghra, Early Miocene, Egypt." Geodiversitas 41, no. 1 (November 7, 2019): 731. http://dx.doi.org/10.5252/geodiversitas2019v41a21.

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20

HUNT JR., R. O. B. E. R. T. M. "Small Oligocene Amphicyonids from North America (Paradaphoenus, Mammalia, Carnivora)." American Museum Novitates 3331 (April 2001): 1–20. http://dx.doi.org/10.1206/0003-0082(2001)331<0001:soafna>2.0.co;2.

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21

Van Valkenburgh, Blaire. "Iterative evolution of hypercarnivory in canids (Mammalia: Carnivora): evolutionary interactions among sympatric predators." Paleobiology 17, no. 4 (1991): 340–62. http://dx.doi.org/10.1017/s0094837300010691.

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Convergent evolution of hypercarnivorous adaptations in canids has occurred a number of times in the last 40 m.y. among distantly related taxa. The adaptations include an increase in carnassial blade length, reduction or loss of post-carnassial molars, and transformation of the talonid of the lower first molar from a basinlike depression into a trenchant, bladelike cusp. Although the diversity of these specialized canids is typically low in past and present communities, it was unusually high during the Late Oligocene of North America and the Pleistocene of South America. These two comparable events provide an opportunity for exploring possible causes of the evolution of hypercarnivory in canids. Plots of generic diversity against time for North American predators reveal a roughly inverse relationship between the number of hypercarnivorous canid taxa and the numbers of other hypercarnivores, such as creodonts, nimravids, mustelids, and amphicyonids. Similarly, the radiation of hypercarnivorous canids in South America occurred at a time of relatively low diversity of other hypercarnivores. Analysis of trophic diversity within the North American carnivore paleoguild before, during, and after the Late Oligocene reveals considerable taxonomic turnover among carnivores because of immigration and speciation. Late Oligocene hypercarnivorous canids appear to have been replaced first by amphicyonids and large mustelids, and then by felids.Despite the repeated tendency of canids to evolve adaptations for hypercarnivory, a canid has yet to appear that is completely catlike, that is, without any post-carnassial molars. This possible constraint on morphological evolution in canids is argued to have resulted, paradoxically, in increased flexibility over evolutionary time and a great potential for rapid diversification and clade survivorship. Finally, it is suggested that the iterative pattern of specialization of the lower molars for meat-slicing that is seen in all families of carnivores, past and present, is probably a result of intraspecific competition for food, perhaps among littermates. This intraspecific selective force is countered by competition among species, since there are limits on the number of sympatric hypercarnivorous species within a single community.
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22

Egi, Naoko, Takehisa Tsubamoto, and Khishigjav Tsogtbaatar. "New Amphicyonid (Mammalia: Carnivora) from the Upper Eocene Ergilin Dzo Formation, Mongolia." Paleontological Research 13, no. 3 (September 2009): 245–49. http://dx.doi.org/10.2517/1342-8144-13.3.245.

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23

HUNT, ROBERT M. "New Amphicyonid Carnivorans (Mammalia, Daphoeninae) from the Early Miocene of Southeastern Wyoming." American Museum Novitates 3385 (December 2002): 1–41. http://dx.doi.org/10.1206/0003-0082(2002)385<0001:nacmdf>2.0.co;2.

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24

Jiangzuo, Qigao, Chunxiao Li, Shiqi Wang, and Danhui Sun. "Amphicyon zhanxiangi, sp. nov., a new amphicyonid (Mammalia, Carnivora) from northern China." Journal of Vertebrate Paleontology 38, no. 6 (November 2, 2018): e1539857. http://dx.doi.org/10.1080/02724634.2018.1539857.

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25

Morlo, Michael, Katharina Bastl, Jörg Habersetzer, Thomas Engel, Bastian Lischewsky, Herbert Lutz, Axel von Berg, Renate Rabenstein, and Doris Nagel. "The apex of amphicyonid hypercarnivory: solving the riddle of Agnotherium antiquum Kaup, 1833 (Mammalia, Carnivora)." Journal of Vertebrate Paleontology 39, no. 5 (September 3, 2019): e1705848. http://dx.doi.org/10.1080/02724634.2019.1705848.

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HUNT, ROBERT M. "Intercontinental Migration of Neogene Amphicyonids (Mammalia, Carnivora): Appearance of the Eurasian Beardog Ysengrinia in North America." American Museum Novitates 3384, no. 1 (2002): 1. http://dx.doi.org/10.1206/0003-0082(2002)384<0001:imonam>2.0.co;2.

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27

Figueirido, Borja, Juan A. Pérez-Claros, Robert M. Hunt, and Paul Palmqvist. "Body Mass Estimation in Amphicyonid Carnivoran Mammals: A Multiple Regression Approach from the Skull and Skeleton." Acta Palaeontologica Polonica 56, no. 2 (June 2011): 225–46. http://dx.doi.org/10.4202/app.2010.0005.

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28

Hunt, Robert M., and Daniel A. Yatkola. "A new species of the amphicyonid carnivore Cynelos Jourdan, 1862 from the early Miocene of North America." Geodiversitas 42, no. 5 (March 5, 2020): 57. http://dx.doi.org/10.5252/geodiversitas2020v42a5.

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29

Siliceo, Gema, Manuel J. Salesa, Mauricio Antón, Juan F. Pastor, and Jorge Morales. "Comparative Anatomy of the Shoulder Region in the Late Miocene Amphicyonid Magericyon anceps (Carnivora): Functional and Paleoecological Inferences." Journal of Mammalian Evolution 22, no. 2 (August 21, 2014): 243–58. http://dx.doi.org/10.1007/s10914-014-9270-9.

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30

Gunnell, Gregg F., Philip D. Gingerich, Michele E. Morgan, and Mary Maas. "Comparative paleoecology of Paleogene and Neogene mammalian faunas: guild structure and diversity." Paleontological Society Special Publications 6 (1992): 115. http://dx.doi.org/10.1017/s2475262200006754.

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We examined guild structure and diversity in the mammalian biota of the Paleogene of Wyoming and Montana and the Neogene of Pakistan. Trophic structure was measured as frequency of generic diversity in each of the following trophic categories: insectivore, omnivore, frugivore, herbivore, and carnivore. Trophic categories were inferred from dental morphology.Results are summarized below (see graphs). In Wyoming/Montana, the Paleocene is dominated by herbivores (from the orders Condylarthra, Multituberculata, and Pantodonta) and insectivores (“Proteutheria” and Proprimates). In the early Eocene, herbivores (Perissodactyla, Artiodactyla, and Rodentia), carnivores (Carnivora, Creodonta, and Mesonychia), and insectivores (“Proteutheria,” Proprimates, and Primates) dominate. At the beginning of the Eocene, carnivores and insectivores are more diverse than herbivores, but later, herbivores are more diverse. Adapid primates become an important frugivore element in the early Eocene. Herbivores are the most common group throughout the sequence, ranging from 30% to 75% of all specimens.In the Siwaliks, herbivores (from Proboscidea, Rodentia, Perissodactyla, and Artiodactyla) dominate in generic diversity (70–80%), with artiodactyls consistently representing about 50% of herbivore genera. Circa 16 Ma, rodents appear and become an important herbivore group. At 14 Ma, bovid diversity increases dramatically. Specialized browsers (tragulids, giraffids) and grazers (suids) represent 25% to 35% of herbivore diversity throughout the sequence. Carnivores, represented by creodonts and true carnivores, range from 5% to 20% of generic diversity in the Siwaliks. At 11 Ma, carnivores increase in generic diversity with amphicyonid and hyaenid carnivores dominating the guild. Carnivores and insectivores are never as diverse in the Siwaliks as they are in the early Eocene of North America.
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31

Domingo, M. Soledad, Laura Domingo, Juan Abella, Alberto Valenciano, Catherine Badgley, and Jorge Morales. "Feeding ecology and habitat preferences of top predators from two Miocene carnivore-rich assemblages." Paleobiology 42, no. 3 (April 28, 2016): 489–507. http://dx.doi.org/10.1017/pab.2015.50.

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AbstractCarnivore-rich fossil sites are uncommon in the fossil record and, accordingly, provide valuable opportunities to study predators from vantages that are rarely applied to ancient faunas. Through stable isotopes of carbon and a Bayesian mixing model, we analyze time-successive (nearly contemporaneous), late Miocene carnivoran populations from two fossil sites (Batallones-1 and Batallones-3) from central Spain. Stable isotopes of carbon in tooth enamel provide a reliable and direct methodology to track ancient diets. These two carnivoran-dominated fossil sites display differences in the composition and abundance of the carnivoran species, with some species present at both sites and some present only at one site. This disparity has been interpreted as the consequence of habitat differences between Batallones-1, the older site, and Batallones-3, the younger site. However, carbon isotope values of carnivore and herbivore tooth enamel suggest a common habitat of C3 woodland originally present at both sites. The differences in the carnivoran faunas rather may be the consequence of the dynamics of species entrance and exit from the Madrid Basin during the time elapsed between Batallones-1 and Batallones-3 and changes in population densities due to biotic factors. We infer higher levels of interspecific competition in Batallones-3 than in Batallones-1 because of the larger number of similar-sized, sympatric predators; the clear overlap in their δ13C values (except for the amphicyonid Magericyon anceps); and similarity of their preferred prey: the hipparionine horses. Finally, carbon stable isotopic composition of Indarctos arctoides teeth implies that this ursid was a carnivorous omnivore rather than a herbivorous omnivore. This work demonstrates the insights that stable isotopes can provide in characterizing the feeding ecology and trophic interactions of ancient carnivoran taxa.
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32

Fournier, Morgane, Sandrine Ladevèze, Kévin Le Verger, Valentin Fischer, Robert P. Speijer, and Floréal Solé. "On the morphology of the astragalus and calcaneus of the amphicyonids (Carnivora, Mammalia) from the Paleogene of Europe: implications for the ecology of the European bear-dogs." Geodiversitas 42, no. 18 (July 16, 2020): 305. http://dx.doi.org/10.5252/geodiversitas2020v42a18.

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Jacobs, Louis, and Christine Janis. "Patterns of evolution in North American Neogene mammals." Paleontological Society Special Publications 6 (1992): 146. http://dx.doi.org/10.1017/s2475262200007061.

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The Neogene of North American represents a time of climatic change from an initially warm, non-arid climate to one with the development of increasing aridity, with warming temperatures through the early part and fluctuating (but basically cooler) temperatures through the later part. This reflects the classic story of a vegetational change from woodland to savanna and eventually to prairie. Note that the transition to true savanna in the Late Miocene was considerably earlier than the first savannas in the Pliocene of the Old World. The evolutionary trends in mammals reflect these climactic and vegetational changes.Some general broad trends are as follows: the replacement of terrestrial and subfossorial moles and geomyid rodents with more specialized fossorial ones; a decrease in the diversity of brachydont rodents and an increase in the diversity of hypsodont ones (including saltatorial forms), and a late Neogene diversification of microtines and deer mice; a decline in the diversity of tree squirrels and terrestrial beavers, and an increase in diversity of ground squirrels and aquatic beavers; the replacement of carnivores belong to more archaic families by more modern types; taxa and an increase in body size, leg length, and hypsodonty in most ungulate taxa, including oreodonts, protoceratids, camelids, antilocaprids, rhinos and equine horses although a couple of taxa show an apparent reversal of these trends: dromomerycids (cervoids) and some anchitherine horses show other morphological changes that suggest progressively more woodland-adapted (rather than savanna-adapted) forms. Tapirs and (to a lesser extent) peccaries seem little affected by the Neogene changes, and persist until the Recent.The Neogene was also punctuated by immigration events (primarily from Asia) and extinctions. The start of the Neogene shows surprisingly little change, with many Paleogene “holdovers”: some new forms appear as either the result of evolution in situ (e.g. equine horses and osteoborine dogs) or as immigrants (e.g. chalicotheres and hemicyonine “dog bears”). The initial major immigrations are during the late Early Miocene, marked by the Asian appearances of true felids (replacing the “false saber-tooths” or nimravids), pecoran ruminants (replacing the hypertragulids), more derived rhinos (replacing the diceratherine rhinos), neomustelids and procyonids. Archaic suoids such as anthracotheres and entelodonts become extinct at this time, and only the more derived ticholeptine oreodonts survive this period. The start of the Middle Miocene is notable for the appearance of proboscideans and deer mice. The Late Miocene sees the decline and eventual disappearance of hedgehogs, archaic carnivores (hemicyonine bears and amphicyonids), most browsing ungulates (oreodonts, protoceratids, many camelids, anchitherine horses, dromomerycids, merycodontine antilocaprids, hornless ruminants, chalicotheres, bunodont gomphotheres), and rhinos. New taxa appearing including ursine bears (immigrants), oversized camels and more derived gomphotheres (in situ evolution). The Pliocene marks a new wave of immigration: microtines, hyenas, true saber-tooths, and cervids come in from Asia; ground sloths (two families appearing in the Late Miocene), glyptodonts, armadillos and capybaras come in from South America. Most mammals that survived the end Miocene extinctions persist, but for many of them (such as horses, camels and antilocaprids) the generic diversity is greatly reduced.
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34

"Long-Legged Pursuit Carnivorans (Amphicyonidae, Daphoeninae) from the Early Miocene of North America." Bulletin of the American Museum of Natural History, 2009. http://dx.doi.org/10.1206/593.1.

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35

Jiangzuo, Qigao, Shiqi Wang, Chunxiao Li, Danhui Sun, and Xiaoxiao Zhang. "New material of Gobicyon (Carnivora, Amphicyonidae, Haplocyoninae) from northern China and a review of Aktaucyonini evolution." Papers in Palaeontology, September 25, 2019. http://dx.doi.org/10.1002/spp2.1283.

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Boardman, GS, and RM Jr Hunt. "New material and evaluation of the chronostratigraphic position of Daphoenictis tedfordi (Mammalia, Carnivora, Amphicyonidae), a cat-like carnivoran from the latest Eocene of northwestern Nebraska, USA." Palaeontologia Electronica, 2015. http://dx.doi.org/10.26879/508.

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37

Morales, Jorge, Juan Abella, Oscar Sanisidro, and Alberto Valenciano. "Ammitocyon kainos gen. et sp. nov., a chimerical amphicyonid (Mammalia, Carnivora) from the late Miocene carnivore traps of Cerro de los Batallones (Madrid, Spain)." Journal of Systematic Palaeontology, May 25, 2021, 1–23. http://dx.doi.org/10.1080/14772019.2021.1910868.

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