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1

Arbour, Victoria M., Derek Larson, Matthew Vavrek, Lisa Buckley, and David Evans. "An ankylosaurian dinosaur from the Cenomanian Dunvegan Formation of northeastern British Columbia, Canada." Fossil Record 23, no. 2 (2020): 179–89. http://dx.doi.org/10.5194/fr-23-179-2020.

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Abstract. Fragmentary but associated dinosaur bones collected in 1930 from the Pine River of northeastern British Columbia are identified here as originating from an ankylosaur. The specimen represents only the second occurrence of dinosaur skeletal material from the Cenomanian Dunvegan Formation and the first from Dunvegan outcrops in the province of British Columbia. Nodosaurid ankylosaur footprints are common ichnofossils in the formation, but the skeletal material described here is too fragmentary to confidently assign to either a nodosaurid or ankylosaurid ankylosaur. The Cenomanian is a time of major terrestrial faunal transitions in North America, but many localities of this age are located in the southern United States; the discovery of skeletal fossils from the Pine River demonstrates the potential for the Dunvegan Formation to produce terrestrial vertebrate fossils that may provide important new data on this significant transitional period during the Cretaceous.
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2

Arbour, Victoria M., and Jordan C. Mallon. "Unusual cranial and postcranial anatomy in the archetypal ankylosaur Ankylosaurus magniventris." FACETS 2, no. 2 (2017): 764–94. http://dx.doi.org/10.1139/facets-2017-0063.

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Ankylosaurus magniventris is an iconic dinosaur species often depicted in popular media. It is known from relatively fragmentary remains compared with its earlier and smaller relatives such as Euoplocephalus and Anodontosaurus. Nevertheless, the known fossils of Ankylosaurus indicate that it had diverged significantly in cranial and postcranial anatomy compared with other Laramidian ankylosaurines. In particular, the dentition, narial region, tail club, and overall body size differ substantially from other Campanian–Maastrichtian ankylosaurines. We review the anatomy of this unusual ankylosaur using data from historic and newly identified material and discuss its palaeoecological implications.
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3

Norman, David B., and Tamsin Faiers. "On the first partial skull of an ankylosaurian dinosaur from the Lower Cretaceous of the Isle of Wight, southern England." Geological Magazine 133, no. 3 (1996): 299–310. http://dx.doi.org/10.1017/s0016756800009031.

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AbstractThe specimen is identified as the partial cranium of a nodosaurid ankylosaur (Ornithischia: Thyreophora) on the basis of the presence of bone which is fused to the dorsal surface of the skull and has secondarily closed the upper temporal fenestrae. The only unequivocally nodosaurid material recovered from the Isle of Wight to date comes from Wealden facies, and has been referred to the genus Polacanthus; it is considered highly probable that this new skull is referable to the same genus. Despite having undergone abrasion, through post-emergence water-rolling, the skull and cranial walls have proved to be relatively informative of the general anatomy of the braincase and the neural and vascular anatomy of this part of the head. The anatomy of the braincase of most ankylosaurs (with the notable exception of the juvenile specimens of the ankylosaurid Pinacosaurus) is surprisingly poorly known, despite the relative abundance of cranial material in North American and Asian collections.The cranial neural and vascular anatomy is well shown in this specimen and enables the first detailed description of nodosaurid endocranial structures. The general form of the brain can be outlined from the structure of the endocast and the principal lobes can be identified; the majority of the cranial nerves have been identified, and a significant component of the associated vascular system is also visible. In most respects the endocast shows a neural anatomy which is common to that known in most dinosaurs. When compared to their nearest relatives, the ankylosaurid ankylosaurs (Euoplocephalus), the nodosaurid endocranial cast shows a more pronounced cerebral flexure, a forebrain which is broader and more expanded dorsally, and a more prominent cerebellum (although there is no evidence for a floccular lobe); there are minor differences in the arrangement of the cranial nerves, and the dorsal portions of the vascular system are better shown.Because of erosion, the olfactory lobes of this specimen of cf. Polacanthus are not preserved, and cannot be compared to those of ankylosaurid ankylosaurs; the latter are unusual in the strong separation of the lobes (reflected in the divergent olfactory stalks); this feature may be associated with the very complex passages within the nasal region of the skull, which are lacking in the nodosaurids described to date.
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4

Apesteguía, Sebastián, and Pablo A. Gallina. "Tunasniyoj, a dinosaur tracksite from the Jurassic-Cretaceous boundary of Bolivia." Anais da Academia Brasileira de Ciências 83, no. 1 (2011): 267–77. http://dx.doi.org/10.1590/s0001-37652011000100015.

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Here we report a superbly preserved and profusely represented five-ichnotaxa dinosaur track assemblage near Icla village, 100 km southeast of Sucre, Bolivia. As preserved in reddish Jurassic-Cretaceous boundary aeolian sandstones, this rich and uncommon assemblage is, additionally, the oldest dinosaur tracksite for Bolivia. Four trackmakers were identified in the area: three quadrupedal and one bipedal, all of them with tracks of around 35 cm in lenght. One of the quadrupedals is represented by no less than five adult individuals (ichnotaxon A), and four purported juveniles (ichnotaxon B) walking in association. The other two quadrupedals (ichnotaxa C and D) involve four trackways, and the last, the bipedal trackmaker (ichnotaxon E), is represented by one trackway. The five ichnotaxa represented in the "Palmar de Tunasniyoj" could be tentatively assigned to the following trackmakers: Ichnotaxa A and B are assigned to basal stegosaurians; ichnotaxon C to a basal tyreophoran, perhaps related to the ankylosaur lineage; ichnotaxon D to the Ankylosauria, and ichnotaxon E to Theropoda. The Tunasniyoj assemblage, the oldest dinosaur tracksite for Bolivia, includes the oldest known evidence assigned to ankylosaurs and stegosaurs for South America.
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5

Arbour, Victoria M., Michael E. Burns, and Philip J. Currie. "A review of pelvic shield morphology in ankylosaurs (Dinosauria: Ornithischia)." Journal of Paleontology 85, no. 2 (2011): 298–302. http://dx.doi.org/10.1666/10-071.1.

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The pelvic shield of ankylosaurian dinosaurs refers to an area of osteoderms lacking differentiated transverse bands over the pelvic region and it is used as a diagnostic character for various ankylosaur groups. The pelvic shield character varies across ankylosaur taxa but is typically coded as a binary character or is excluded from phylogenetic analyses, which obscures evolutionary trends and relationships. This study investigates for the first time pelvic shield morphology in a stratigraphic and geographic context. This paper comprehensively reviews pelvic shield morphology with firsthand observations of specimens, and proposes three categories of pelvic shield morphology. Category 1 pelvic shields have un-fused but tightly interlocking osteoderms. Category 2 pelvic shields have fused osteoderms forming rosettes and are restricted to the Late Jurassic to mid Cretaceous of North America and Europe. Category 3 pelvic shields have fused polygonal osteoderms of similar size, and are found in the mid- to Late Cretaceous of North America. Although the pelvic shield is used to characterize the Polacanthidae, an interpretation supported by this review, the validity of such a clade is dependent upon a global parsimony analysis incorporating this character. Future analyses of the Ankylosauria should incorporate a more detailed treatment of the pelvic shield to determine its diagnostic value within the group.
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6

Xu, Xing, Xiao-Lin Wang, and Hai-Lu You. "A juvenile ankylosaur from China." Naturwissenschaften 88, no. 7 (2001): 297–300. http://dx.doi.org/10.1007/s001140100233.

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7

Parsons, William L., and Kristen M. Parsons. "A new ankylosaur (Dinosauria: Ankylosauria) from the Lower Cretaceous Cloverly Formation of central Montana." Canadian Journal of Earth Sciences 46, no. 10 (2009): 721–38. http://dx.doi.org/10.1139/e09-045.

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A cranium and other associated skeletal elements representing a new ankylosaurid dinosaur, Tatankacephalus cooneyorum gen. et sp. nov. possess several diagnostic features that indicate that this new taxon differs from the only other known ankylosaur from the Cloverly Formation, Sauropelta edwardsorum . These features include a frontoparietal dome, an enlarged nuchal ridge that obscures the occipital region, a circular orbit, ventral curvature in the posterolaterally directed paroccipital processes, a posteroventrally directed foramen magnum, and a number of features on the braincase. The phylogenetic analysis positions Tatankacephalus with Ankylosauridae based on its sharing of several characters with other members of this clade, including an enlarged nuchal segment that obscures the occiput in dorsal view, a ventrally curving lateral profile of the cranium anterior to the orbit, pyramidal postorbital boss, laterally projecting pyramidal quadratojugal boss, the presence of a postocular shelf, the presence of paranasal sinuses, and the lack of a cingulum on a maxillary (or dentary) tooth. It is considered a basal member of Ankylosauridae because it retains premaxillary teeth and a visible lateral temporal fenestra, in contrast to the absence of premaxillary teeth and an obscured lateral temporal fenestra in younger members of this clade.
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8

Carpenter, Kenneth, Tony DiCroce, Billy Kinneer, and Robert Simon. "Pelvis of Gargoyleosaurus (Dinosauria: Ankylosauria) and the Origin and Evolution of the Ankylosaur Pelvis." PLoS ONE 8, no. 11 (2013): e79887. http://dx.doi.org/10.1371/journal.pone.0079887.

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9

Rodríguez-de la Rosa, Rubén A., María Patricia Velasco-de León, Javier Arellano-Gil, and Diego Enrique Lozano-Carmona. "Middle Jurassic ankylosaur tracks from Mexico." Boletín de la Sociedad Geológica Mexicana 70, no. 2 (2018): 379–95. http://dx.doi.org/10.18268/bsgm2018v70n2a8.

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10

Molnar, Ralph E., and H. Trevor Clifford. "Gut contents of a small ankylosaur." Journal of Vertebrate Paleontology 20, no. 1 (2000): 194–96. http://dx.doi.org/10.1671/0272-4634(2000)020[0194:gcoasa]2.0.co;2.

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11

Carpenter, Kenneth, Clifford Miles, and Karen Cloward. "Skull of a Jurassic ankylosaur (Dinosauria)." Nature 393, no. 6687 (1998): 782–83. http://dx.doi.org/10.1038/31684.

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12

Matthias, Angela E., Lorrie A. McWhinney, and Kenneth Carpenter. "Pathological pitting in ankylosaur (Dinosauria) osteoderms." International Journal of Paleopathology 13 (June 2016): 82–90. http://dx.doi.org/10.1016/j.ijpp.2016.02.006.

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13

Hayashi, Shoji, Kenneth Carpenter, Torsten M. Scheyer, Mahito Watabe, and Daisuke Suzuki. "Function and Evolution of Ankylosaur Dermal Armor." Acta Palaeontologica Polonica 55, no. 2 (2010): 213–28. http://dx.doi.org/10.4202/app.2009.0103.

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14

Hawakaya, Hiroshi, Makoto Manabe, and Kenneth Carpenter. "Nodosaurid Ankylosaur from the Cenomanian of Japan." Journal of Vertebrate Paleontology 25, no. 1 (2005): 240–45. http://dx.doi.org/10.1671/0272-4634(2005)025[0240:naftco]2.0.co;2.

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15

Vickaryous, Matthew K., Anthony P. Russell, Philip J. Currie, and Xi-Jin Zhao. "A new ankylosaurid (Dinosauria: Ankylosauria) from the Lower Cretaceous of China, with comments on ankylosaurian relationships." Canadian Journal of Earth Sciences 38, no. 12 (2001): 1767–80. http://dx.doi.org/10.1139/e01-051.

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Amongst the fossil material collected by the Sino-Soviet Expeditions (1959–1960) to the Alshan Desert, China, was a large, virtually complete ankylosaur skeleton. Gobisaurus domoculus gen. et sp. nov. closely resembles Shamosaurus scutatus, but is distinct in having an unfused basipterygoid–pterygoid contact and elongate premaxillary processes of the vomers. Although it is difficult to make a definitive taxonomic assignment without considering postcranial material, a preliminary phylogenetic analysis places Gobisaurus as the sister taxon of Shamosaurus, clustered as one of several successive outgroups of the Ankylosaurinae.
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16

Paulina-Carabajal, Ariana, Francisco T. Barrios, Ariel H. Méndez, Ignacio A. Cerda, and Yuong-Nam Lee. "A Late Cretaceous dinosaur and crocodyliform faunal association–based on isolate teeth and osteoderms–at Cerro Fortaleza Formation (Campanian-Maastrichtian) type locality, Santa Cruz, Argentina." PLOS ONE 16, no. 9 (2021): e0256233. http://dx.doi.org/10.1371/journal.pone.0256233.

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The Late Cretaceous dinosaur record in southern South America has been improved recently; particularly with findings from Chorrillo and Cerro Fortaleza formations, both bearing ankylosaur remains, a clade that was not previously recorded in the Austral Basin. The dinosaur fauna of the type locality of Cerro Fortaleza Formation is known from -and biased to- large-sized sauropod remains and a single described taxon, the titanosaur Dreadnoughtus schrani. Here, we report the taxonomic composition of a site preserving thirteen isolated teeth and several osteoderms belonging to three dinosaur clades (Abelisauridae, Titanosauria, and Nodosauridae), and at least one clade of notosuchian crocodyliforms (Peirosauridae). They come from sediments positioned at the mid-section of the Cerro Fortaleza Formation, which is Campanian-Maastrichtian in age, adding valuable information to the abundance and biodiversity of this Cretaceous ecosystem. Since non-titanosaur dinosaur bones are almost absent in the locality, the teeth presented here provide a window onto the archosaur biodiversity of the Late Cretaceous in southern Patagonia. The nodosaurid tooth and small armor ossicles represent the first record of ankylosaurs for this stratigraphic unit. The peirosaurid material also represents the most austral record of the clade in South America.
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17

Kubo, Tai, Wenjie Zheng, Mugino O. Kubo, and Xingsheng Jin. "Dental microwear of a basal ankylosaurine dinosaur, Jinyunpelta and its implication on evolution of chewing mechanism in ankylosaurs." PLOS ONE 16, no. 3 (2021): e0247969. http://dx.doi.org/10.1371/journal.pone.0247969.

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Jinyunpelta sinensis is a basal ankylosaurine dinosaur excavated from the mid Cretaceous Liangtoutang Formation of Jinyun County, Zhejiang Province, China. In the present study, its dental microwear was observed using a confocal laser microscope. Jinyunpelta had steep wear facets that covered most of buccal surfaces of posterior dentary teeth. Observation of dental microwear on the wear facet revealed that scratch orientation varied according to its location within the wear facet: vertically (i.e. apicobasally) oriented scratches were dominant in the upper half of the wear facet, and horizontally (i.e. mesiolaterally) oriented ones were in the bottom of the facet. These findings indicated that Jinyunpelta adopted precise tooth occlusion and biphasal jaw movement (orthal closure and palinal lower jaw movement). The biphasal jaw movement was widely observed among nodosaurids, among ankylosaurids, it was previously only known from the Late Cretaceous North American taxa, and not known among Asian ankylosaurids. The finding of biphasal jaw movement in Jinyunpelta showed sophisticate feeding adaptations emerged among ankylosaurids much earlier (during Albian or Cenomanian) than previously thought (during Campanian). The Evolution of the biphasal jaw mechanism that contemporaneously occurred among two lineages of ankylosaurs, ankylosaurids and nodosaurids, showed high evolutionary plasticity of ankylosaur jaw mechanics.
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18

Carpenter, Kenneth, and Michael J. Everhart. "Skull of the ankylosaur Niobrarasaurus coleu (Ankylosauria: Nodosauridae) from the Smoky Hill Chalk (Coniacian) of western Kansas." Transactions of the Kansas Academy of Science 110, no. 1 & 2 (2007): 1–9. http://dx.doi.org/10.1660/0022-8443(2007)110[1:sotanc]2.0.co;2.

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19

Scheyer, Torsten M., and P. Martin Sander. "Histology of ankylosaur osteoderms: implications for systematics and function." Journal of Vertebrate Paleontology 24, no. 4 (2004): 874–93. http://dx.doi.org/10.1671/0272-4634(2004)024[0874:hoaoif]2.0.co;2.

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20

Coombs, Walter P. "A juvenile ankylosaur referable to the genusEuoplocephalus(Reptilia, Ornithischia)." Journal of Vertebrate Paleontology 6, no. 2 (1986): 162–73. http://dx.doi.org/10.1080/02724634.1986.10011608.

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21

Gangloff, Roland A. "EdmontoniaSp., The First Record of an Ankylosaur from Alaska." Journal of Vertebrate Paleontology 15, no. 1 (1995): 195–200. http://dx.doi.org/10.1080/02724634.1995.10011218.

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22

Barrett, Paul M., and Paul Upchurch. "Regnosaurus northamptoni, a stegosaurian dinosaur from the Lower Cretaceous of southern England." Geological Magazine 132, no. 2 (1995): 213–22. http://dx.doi.org/10.1017/s0016756800011754.

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AbstractThe type specimen ofRegnosaurus northamptoniMantell, a partial right mandible, is redescribed and compared with a wide variety of dinosaurian taxa. We suggest thatRegnosaurusis not an ankylosaur or a sauropod as previously thought, but represents the fragmentary remains of a ‘primitive’ stegosaur.RegnosaurusandHuayangosaurusshare two derived characters, suggesting that these two genera should be provisionally regarded as sister taxa. The implications of these findings for stegosaur evolution and biogeography are discussed briefly.
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23

Hooker, J. J., A. C. Milner, and S. E. K. Sequeira. "An ornithopod dinosaur from the Late Cretaceous of West Antarctica." Antarctic Science 3, no. 3 (1991): 331–32. http://dx.doi.org/10.1017/s0954102091000391.

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In February 1989, the partial skeleton of an ornithopod dinosaur was discovered during a British Antarctic Survey (BAS) expedition, supported by RRS John Biscoe, to the James Ross Island area, east of the Antarctic Peninsula. This was only the second dinosaur to be found in the continent of Antarctica, the first being an ankylosaur collected three years earlier (Olivero et al. 1986, Gasparini et al. 1987, Gasparini 1988, Gasparini & Olivero 1989, Olivero et al. 1991).
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24

Godfrey, Stephen J., and Philip J. Currie. "A xiphisternal from the Dinosaur Park Formation (Campanian, Upper Cretaceous) of Alberta, Canada." Canadian Journal of Earth Sciences 31, no. 11 (1994): 1661–63. http://dx.doi.org/10.1139/e94-148.

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Xiphisternal elements of dinosaurs are only rarely recovered, probably because they seldom ossified and those that did were fragile and easily destroyed. An isolated but relatively complete, right xiphisternal element was collected from the Dinosaur Park Formation of Dinosaur Provincial Park, Alberta. It apparently contacted the left xiphisternal, the sternum, and at least three costal cartilages. Similar in overall morphology to xiphisternals of Edmontonia, Nodosaurus, and Panoplosaurus, it can be referred to the ankylosaur family Nodosauridae.
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25

Sachs, Sven, and Jahn J. Hornung. "Ankylosaur Remains from the Early Cretaceous (Valanginian) of Northwestern Germany." PLoS ONE 8, no. 4 (2013): e60571. http://dx.doi.org/10.1371/journal.pone.0060571.

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26

Burns, Michael E., Tatiana A. Tumanova, and Philip J. Currie. "Postcrania of juvenile Pinacosaurus grangeri (Ornithischia: Ankylosauria) from the Upper Cretaceous Alagteeg Formation, Alag Teeg, Mongolia: implications for ontogenetic allometry in ankylosaurs." Journal of Paleontology 89, no. 1 (2015): 168–82. http://dx.doi.org/10.1017/jpa.2014.14.

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AbstractThe ankylosaurine Pinacosaurus is one of the best known ankylosaur to date in terms of the number and preservational quality of specimens. Juvenile to sub-adult postcrania collected by the Soviet-Mongolian Paleontological Expedition from the Upper Cretaceous Alagteeg Formation at Alag Teeg, Mongolia can be assigned to Pinacosaurus grangeri based on discrete cranial characters. One individual is significantly larger than the others and demonstrates delayed fusion of postcranial elements with the earliest occurring between dorsal ribs and vertebrae. The robustness of forelimb elements is positively allometric with respect to their length, indicating weight-bearing relationship. Such length-dependent correlations are not seen in the hind limbs. Finally, incipient cervical half rings suggest a developmental pathway of outgrowths from the underlying band combined with fusion of overlying osteoderms.
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27

Long, John. "The Enigmatic Dinosaur Faunas of Australia." Paleontological Society Special Publications 7 (1994): 397–410. http://dx.doi.org/10.1017/s2475262200009679.

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Australia has a scant but significant record of dinosaurs which includes a prosauropod, two or three sauropods, at least four theropods (including carnosaurs and an ornithomimosaur), four hypsilophodontids, an aberrant iguanodont, a primitive thyreophoran (possibly an ankylosaur), and a possible primitive neoceratopsian. Footprint fossils indicate higher taxonomic diversity, including small bipedal ornithopods and coelurosaurs, very large sauropods, large carnosaurs, and a stegosaur. The most diverse dinosaur assemblage, from the Early Cretaceous of Victoria, represents a polar dinosaur community that lived within the Cretaceous Antarctic Circle.
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Murray, Alejandro, Facundo Riguetti, and Sebastián Rozadilla. "New ankylosaur (Thyreophora, ornithischia) remains from the Upper Cretaceous of Patagonia." Journal of South American Earth Sciences 96 (December 2019): 102320. http://dx.doi.org/10.1016/j.jsames.2019.102320.

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29

Coombs, Walter P. "A Nodosaurid Ankylosaur (Dinosauria: Ornithischia) from the Lower Cretaceous of Texas." Journal of Vertebrate Paleontology 15, no. 2 (1995): 298–312. http://dx.doi.org/10.1080/02724634.1995.10011231.

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Augustin, Felix J., Andreas T. Matzke, Michael W. Maisch, and Hans-Ulrich Pfretzschner. "First evidence of an ankylosaur (Dinosauria, Ornithischia) from the Jurassic Qigu Formation (Junggar Basin, NW China) and the early fossil record of Ankylosauria." Geobios 61 (August 2020): 1–10. http://dx.doi.org/10.1016/j.geobios.2020.06.005.

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Arbour, Victoria M., and Philip J. Currie. "Analyzing Taphonomic Deformation of Ankylosaur Skulls Using Retrodeformation and Finite Element Analysis." PLoS ONE 7, no. 6 (2012): e39323. http://dx.doi.org/10.1371/journal.pone.0039323.

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Ősi, Attila. "Hungarosaurus tormai, a new ankylosaur (Dinosauria) from the Upper Cretaceous of Hungary." Journal of Vertebrate Paleontology 25, no. 2 (2005): 370–83. http://dx.doi.org/10.1671/0272-4634(2005)025[0370:htanad]2.0.co;2.

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Burns, Michael E. "Taxonomic utility of ankylosaur (Dinosauria, Ornithischia) osteoderms:Glyptodontopelta mimusFord, 2000: a test case." Journal of Vertebrate Paleontology 28, no. 4 (2008): 1102–9. http://dx.doi.org/10.1671/0272-4634-28.4.1102.

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Kilbourne, Brandon, and Kenneth Carpenter. "Redescription of Gargoyleosaurus parkpinorum, a polacanthid ankylosaur from the Upper Jurassic of Albany County, Wyoming." Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 235, no. 1 (2005): 111–60. http://dx.doi.org/10.1127/njgpa/235/2005/111.

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Kilbourne, Brandon, and Kenneth Carpenter. "Redescription of Gargoyleosaurus parkpinorum, a polacanthid ankylosaur from the Upper Jurassic of Albany County, Wyoming." Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 237, no. 1 (2005): 111–60. http://dx.doi.org/10.1127/njgpa/237/2005/111.

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36

Coombs, Walter P., and Thomas A. Deméré. "A Late Cretaceous nodosaurid ankylosaur (Dinosauria: Ornithischia) from marine sediments of coastal California." Journal of Paleontology 70, no. 2 (1996): 311–26. http://dx.doi.org/10.1017/s0022336000023404.

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A partial nodosaurid ankylosaur skeleton, consisting primarily of the ilia, hindlimbs, posterior dorsal armor, plus partial forelimb elements and additional armor, was recovered from the marine Point Loma Formation, late Campanian age, north of San Diego, California. The specimen is similar to contemporaneous species of Panoplosaurus and Edmontonia from terrestrial sediments of the western interior, but there are also similarities to the armor of Stegopelta landerensis from marine sediments of earliest Cenomanian age from Wyoming. Skeletal elements critical for generic determination are not preserved, and the specimen is identified as Nodosauridae, incertae sedis. An associated marine invertebrate fauna and nannoplankton flora have revealed some discrepencies in the correlation systems used for Upper Cretaceous marine rocks of coastal California. Hollow limb bones of the specimen are interpreted as a preservational artifact. Nodosaurids had broad ecological tolerances, and visited riparian and coastal environments more frequently than other dinosaurs. However, a review of morphologic and distributional evidence fails to support a theory of amphibious or aquatic habits for nodosaurids.
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37

Rongjun, CHEN, ZHENG Wenjie, Yoichi AZUMA, et al. "A New Nodosaurid Ankylosaur from the Chaochuan Formation of Dongyang, Zhejiang Province, China." Acta Geologica Sinica - English Edition 87, no. 3 (2013): 658–71. http://dx.doi.org/10.1111/1755-6724.12077.

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38

Burns, Michael E., and Matthew J. Vavrek. "Probable Ankylosaur Ossicles from the Middle Cenomanian Dunvegan Formation of Northwestern Alberta, Canada." PLoS ONE 9, no. 5 (2014): e96075. http://dx.doi.org/10.1371/journal.pone.0096075.

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Burns, Michael E., and Philip J. Currie. "External and internal structure of ankylosaur (Dinosauria, Ornithischia) osteoderms and their systematic relevance." Journal of Vertebrate Paleontology 34, no. 4 (2014): 835–51. http://dx.doi.org/10.1080/02724634.2014.840309.

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JI, Shu-an, Lijun ZHANG, Shudong ZHANG, Lijun ZHANG, and Shan HANG. "Large-Sized Ankylosaur (Dinosauria) from the Lower Cretaceous Jiufotang Formation of Western Liaoning, China." Acta Geologica Sinica - English Edition 88, no. 4 (2014): 1060–65. http://dx.doi.org/10.1111/1755-6724.12273.

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Weishampel, David B. "The evolution of ornithischian dinosaurs during the Cretaceous: jaws, plants, and evolutionary metrics revisited." Paleontological Society Special Publications 6 (1992): 308. http://dx.doi.org/10.1017/s2475262200008686.

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Coevolutionary links between plants and herbivores, often cited as examples of adaptive response of one group of organisms to another, have been much studied from both neontological and paleontological perspectives. The most commonly cited case of coevolution from the latter viewpoint is the radiation of grassland grasses and grazing mammals during the mid-Tertiary. Other promising examples are also beginning to emerge, among them the radiation of ornithischian dinosaurs and early angiosperms during the Cretaceous.Preliminary studies (Weishampel and Norman 1989) analyzed the temporal distribution of trophic groups among Ornithischia (and other herbivorous tetrapods) across the Mesozoic. Trophic groups were based on a mixture of monophyletic and paraphyletic taxa assigned to assorted taxonomic rank. Speciation, extinction, and turn-over rates were calculated from stratigraphic data to identify co-evolutionary “hot spots” between herbivores and contemporary plants: times of evolutionary perturbation between these two groups of organisms within the Mesozoic.The present study reanalyzes Ornithischia from the perspective of ecosystem richness and phylogeny. As in the previous study, trophic categories are assessed on the basis of tooth morphology, occlusal patterns, and jaw construction. All ornithischians appear to orally process food in ways ranging from orthal pulping (ankylosaurs, most pachycephalosaurs) to orthal slicing and transverse grinding (euornithopodans, ceratopsians, some pachycephalosaurs).During the Early Cretaceous, relative species-level trophic diversity (as expressed as percentages) for ornithischians consists of a subequal mixture of herbivores with a transverse power-stroke (euornithopodans) and orthal pulpers (ankylosaurs, rare pachycephalosaurs). Toward the end of the Early Cretaceous, orthal slicing herbivores (ceratopsians) have their earliest record. At the end of the Cretaceous (Santonian-Maastrichtian), the transverse-chewing euornithopodans contribute somewhat less toward relative diversity, while the orthal slicing ceratopsians diversify and orthal pulpers decline (mostly due to reduced ankylosaur diversity).The pattern of acquisition of feeding/chewing styles among Cretaceous ornithischians is determined by mapping trophic categories on the phylogeny of the clade. With this in mind, our knowledge of the phylogeny of ornithischian tropic organization is in part a product of the completeness of the fossil record of these animals. Errors that may accrue because of a patchy record can be partially corrected by combining phylogenetic and stratigraphic information. This approach calls for the identification of ghost lineages, as well as their calibration using minimal divergence times (MDTs). Thus, diversity counts based on monophyly and evolutionary continuing can be augmented for yet-to-be-discovered species. Using ghost lineages and MDTs, a fuller picture of the pattern of acquisition of different jaw systems (and hence trophic organization) among ornithischian will become available.
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McDonald, Andrew T., and Douglas G. Wolfe. "A new nodosaurid ankylosaur (Dinosauria: Thyreophora) from the Upper Cretaceous Menefee Formation of New Mexico." PeerJ 6 (August 24, 2018): e5435. http://dx.doi.org/10.7717/peerj.5435.

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Nodosauridae is a clade of armored dinosaurs with a rich fossil record and long history of study in North America. Nodosaurid fossils have been collected throughout the western United States and Canada. Here, we report three new nodosaurid specimens from the Upper Cretaceous (lower Campanian) Allison Member of the Menefee Formation, San Juan Basin, northwestern New Mexico. The three specimens belong to a new genus and species,Invictarx zephyri, characterized by a unique combination of features pertaining to the morphology of the osteoderms. Among the three specimens there are representative cervical/pectoral and thoracic osteoderms, as well as components of a probable co-ossified pelvic shield. The new tax on is most similar toGlyptodontopelta mimusfrom the Maastrichtian of New Mexico.
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Lee, Yuong-Nam. "A new nodosaurid ankylosaur (Dinosauria: Ornithischia) from the Paw Paw Formation (Late Albian) of Texas." Journal of Vertebrate Paleontology 16, no. 2 (1996): 232–45. http://dx.doi.org/10.1080/02724634.1996.10011311.

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Arbour, Victoria M., and David C. Evans. "A new ankylosaurine dinosaur from the Judith River Formation of Montana, USA, based on an exceptional skeleton with soft tissue preservation." Royal Society Open Science 4, no. 5 (2017): 161086. http://dx.doi.org/10.1098/rsos.161086.

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The terrestrial Judith River Formation of northern Montana was deposited over an approximately 4 Myr interval during the Campanian (Late Cretaceous). Despite having been prospected and collected continuously by palaeontologists for over a century, few relatively complete dinosaur skeletons have been recovered from this unit to date. Here we describe a new genus and species of ankylosaurine dinosaur, Zuul crurivastator , from the Coal Ridge Member of the Judith River Formation, based on an exceptionally complete and well-preserved skeleton (ROM 75860). This is the first ankylosaurin skeleton known with a complete skull and tail club, and it is the most complete ankylosaurid ever found in North America. The presence of abundant soft tissue preservation across the skeleton, including in situ osteoderms, skin impressions and dark films that probably represent preserved keratin, make this exceptional skeleton an important reference for understanding the evolution of dermal and epidermal structures in this clade. Phylogenetic analysis recovers Zuul as an ankylosaurin ankylosaurid within a clade of Dyoplosaurus and Scolosaurus , with Euoplocephalus being more distantly related within Ankylosaurini. The occurrence of Z. crurivastator from the upper Judith River Formation fills a gap in the ankylosaurine stratigraphic and geographical record in North America, and further highlights that Campanian ankylosaurines were undergoing rapid evolution and stratigraphic succession of taxa as observed for Laramidian ceratopsids, hadrosaurids, pachycephalosaurids and tyrannosaurids.
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Wiersma, Jelle P., and Randall B. Irmis. "A new southern Laramidian ankylosaurid, Akainacephalus johnsoni gen. et sp. nov., from the upper Campanian Kaiparowits Formation of southern Utah, USA." PeerJ 6 (July 19, 2018): e5016. http://dx.doi.org/10.7717/peerj.5016.

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A partial ankylosaurid skeleton from the upper Campanian Kaiparowits Formation of southern Utah is recognized as a new taxon, Akainacephalus johnsoni, gen. et sp. nov. The new taxon documents the first record of an associated ankylosaurid skull and postcranial skeleton from the Kaiparowits Formation. Preserved material includes a complete skull, much of the vertebral column, including a complete tail club, a nearly complete synsacrum, several fore- and hind limb elements, and a suite of postcranial osteoderms, making Akainacephalus johnsoni the most complete ankylosaurid from the Late Cretaceous of southern Laramidia. Arrangement and morphology of cranial ornamentation in Akainacephalus johnsoni is strikingly similar to Nodocephalosaurus kirtlandensis and some Asian ankylosaurids (e.g., Saichania chulsanensis, Pinacosaurus grangeri, and Minotaurasaurus ramachandrani); the cranium is densely ornamented with symmetrically arranged and distinctly raised ossified caputegulae which are predominantly distributed across the dorsal and dorsolateral regions of the nasals, frontals, and orbitals. Cranial caputegulae display smooth surface textures with minor pitting and possess a distinct conical to pyramidal morphology which terminates in a sharp apex. Character analysis suggests a close phylogenetic relationship with N. kirtlandensis, M. ramachandrani, Tarchia teresae, and S. chulsanensis, rather than with Late Cretaceous northern Laramidian ankylosaurids (e.g., Euoplocephalus tutus, Anodontosaurus lambei, and Ankylosaurus magniventris). These new data are consistent with evidence for distinct northern and southern biogeographic provinces in Laramidia during the late Campanian. The addition of this new ankylosaurid taxon from southern Utah enhances our understanding of ankylosaurid diversity and evolutionary relationships. Potential implications for the geographical distribution of Late Cretaceous ankylosaurid dinosaurs throughout the Western Interior suggest multiple time-transgressive biogeographic dispersal events from Asia into Laramidia.
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姬, 书安. "Discovery of the Early Cretaceous Ankylosaur Gastroliths and Scale Impressions from the Ordos Basin, Inner Mongolia, China." Advances in Geosciences 06, no. 05 (2016): 355–60. http://dx.doi.org/10.12677/ag.2016.65037.

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Ősi, Attila, János Magyar, Károly Rosta, and Matthew Vickaryous. "Cranial ornamentation in the Late Cretaceous nodosaurid ankylosaurHungarosaurus." PeerJ 9 (March 3, 2021): e11010. http://dx.doi.org/10.7717/peerj.11010.

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Bony cranial ornamentation is developed by many groups of vertebrates, including ankylosaur dinosaurs. To date, the morphology and ontogenetic origin of ankylosaurian cranial ornamentation has primarily focused on a limited number of species from only one of the two major lineages, Ankylosauridae. For members of the sister group Nodosauridae, less is known. Here, we provide new details of the cranial anatomy of the nodosauridHungarosaurusfrom the Santonian of Europe. Based on a number of previously described and newly identified fragmentary skulls and skull elements, we recognize three different size classes ofHungarosaurus. We interpret these size classes as representing different stages of ontogeny. Cranial ornamentation is already well-developed in the earliest ontogenetic stage represented herein, suggesting that the presence of outgrowths may have played a role in intra- and interspecific recognition. We find no evidence that cranial ornamentation inHungarosaurusinvolves the contribution of coossified osteoderms. Instead, available evidence indicates that cranial ornamentation forms as a result of the elaboration of individual elements. Although individual differences and sexual dimorphism cannot be excluded, the observed variation inHungarosauruscranial ornamentation appears to be associated with ontogeny.
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Raven, Thomas J., Paul M. Barrett, Stuart B. Pond, and Susannah C. R. Maidment. "Osteology and Taxonomy of British Wealden Supergroup (Berriasian–aptian) Ankylosaurs (Ornithischia, Ankylosauria)." Journal of Vertebrate Paleontology 40, no. 4 (2020): e1826956. http://dx.doi.org/10.1080/02724634.2020.1826956.

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Hornung, Jahn J., and Mike Reich. "Metatetrapous valdensisNopcsa, 1923 and the Presence of Ankylosaur Tracks (Dinosauria: Thyreophora) in the Berriasian (Early Cretaceous) of Northwestern Germany." Ichnos 21, no. 1 (2014): 1–18. http://dx.doi.org/10.1080/10420940.2013.873720.

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Petti, Fabio Massimo, Simone D’Orazi Porchetti, Eva Sacchi, and Umberto Nicosia. "A new purported ankylosaur trackway in the Lower Cretaceous (lower Aptian) shallow-marine carbonate deposits of Puglia, southern Italy." Cretaceous Research 31, no. 6 (2010): 546–52. http://dx.doi.org/10.1016/j.cretres.2010.07.004.

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