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Journal articles on the topic 'Bembidion'

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Published: 3 June 2025
Last updated: 19 July 2025

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1

Neri, Paolo, and Luca Toledano. "Geographic and taxonomic notes, addenda and corrigenda on the subtribe Bembidiina Stephens, 1827 of the 2017 ‘Catalogue of Palaearctic Coleoptera’ (Coleoptera, Carabidae, Bembidiina)." ZooKeys 1044 (June 16, 2021): 563–87. http://dx.doi.org/10.3897/zookeys.1044.62593.

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Some corrections to the section of subtribe Bembidiina of the Catalogue of Palaearctic Coleoptera, Vol. 1, together with geographic, systematic, and synonymic updates are reported and commented upon. The following five new synonymies are proposed (with junior synonym listed first): Bembidion (Peryphanes) dostali Kirschenhofer, 1984 = Bembidion (Peryphanes) sanatum Bates, 1883 syn. nov.; Bembidion (Terminophanes) pseudoconsumatum Kirschenhofer, 1984 = Bembidion (Terminophanes) sjoelanderi Jedlička, 1965 syn. nov.; Bembidion (Asioperyphus) sapporense Jedlička, 1951 = Bembidion (Politophanes) chloreum Bates, 1873 syn. nov.; Bembidion (Peryphus) torosiense Jedlička, 1961 = Bembidion (Peryphus) subcostatum vau Netolitzky, 1913 syn. nov.; Sinechostictus (Sinechostictus) multisulcatus cariniger Korge, 1971 = Sinechostictus (Sinechostictus) multisulcatus (Reitter, 1890) syn. nov. Furthermore we confirm the following synonimies: Asaphidion weiratheri Netolitzky, 1935 = Asaphidion ganglbaueri J.Müller, 1921; Sinechostictus (Sinechostictus) effluviorum (Peyron, 1858) = Sinechostictus (Sinechostictus) tarsicus (Peyron, 1858). The following nine new combinations are proposed: Bembidion (Euperyphus) dimidiatum Ménétriés, 1832 comb. nov.; Bembidion (Peryphus) psuchrum Andrewes, 1922 comb. nov.; Bembidion (Plataphus) pseudolucillum Netolitzky, 1938 comb. nov.; Bembidion (Politophanes) chloreum Bates, 1873 comb. nov.; Bembidion (Politophanes) gotoense Habu, 1973 comb. nov.; Bembidion (Politophanes) shunichii Habu, 1973 comb. nov.; Bembidion (Politophanes) umeyai Habu, 1959 comb. nov.; Bembidion (Politophanes) yoshidai Morita, 2009 comb. nov.; Bembidion (Terminophanes) sjoelanderi Jedlička, 1965 comb. nov. The species Bembidion psuchrum Andrewes, 1922 and Bembidion sanatum Bates, 1883 are here redescribed.
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2

Neri, Paolo, and Luca Toledano. "Geographic and taxonomic notes, addenda and corrigenda on the subtribe Bembidiina Stephens, 1827 of the 2017 'Catalogue of Palaearctic Coleoptera' (Coleoptera, Carabidae, Bembidiina)." ZooKeys 1044 (June 16, 2021): 563–87. https://doi.org/10.3897/zookeys.1044.62593.

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Some corrections to the section of subtribe Bembidiina of the Catalogue of Palaearctic Coleoptera, Vol. 1, together with geographic, systematic, and synonymic updates are reported and commented upon. The following five new synonymies are proposed (with junior synonym listed first): Bembidion (Peryphanes) dostali Kirschenhofer, 1984 = Bembidion (Peryphanes) sanatum Bates, 1883 syn. nov.; Bembidion (Terminophanes) pseudoconsumatum Kirschenhofer, 1984 = Bembidion (Terminophanes) sjoelanderi Jedlička, 1965 syn. nov.; Bembidion (Asioperyphus) sapporense Jedlička, 1951 = Bembidion (Politophanes) chloreum Bates, 1873 syn. nov.; Bembidion (Peryphus) torosiense Jedlička, 1961 = Bembidion (Peryphus) subcostatum vau Netolitzky, 1913 syn. nov.; Sinechostictus (Sinechostictus) multisulcatus cariniger Korge, 1971 = Sinechostictus (Sinechostictus) multisulcatus (Reitter, 1890) syn. nov. Furthermore we confirm the following synonimies: Asaphidion weiratheri Netolitzky, 1935 = Asaphidion ganglbaueri J.Müller, 1921; Sinechostictus (Sinechostictus) effluviorum (Peyron, 1858) = Sinechostictus (Sinechostictus) tarsicus (Peyron, 1858). The following nine new combinations are proposed: Bembidion (Euperyphus) dimidiatum Ménétriés, 1832 comb. nov.; Bembidion (Peryphus) psuchrum Andrewes, 1922 comb. nov.; Bembidion (Plataphus) pseudolucillum Netolitzky, 1938 comb. nov.; Bembidion (Politophanes) chloreum Bates, 1873 comb. nov.; Bembidion (Politophanes) gotoense Habu, 1973 comb. nov.; Bembidion (Politophanes) shunichii Habu, 1973 comb. nov.; Bembidion (Politophanes) umeyai Habu, 1959 comb. nov.; Bembidion (Politophanes) yoshidai Morita, 2009 comb. nov.; Bembidion (Terminophanes) sjoelanderi Jedlička, 1965 comb. nov. The species Bembidion psuchrum Andrewes, 1922 and Bembidion sanatum Bates, 1883 are here redescribed.
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3

SERGIO ROIG-JUÑENT, SERGIO, and ERICA SCHEIBLER. "A new Argentinian species of Bembidion (Chilioperyphus) (Coleoptera: Carabidae: Bembidiini) with accessory setae." Zootaxa 782, no. 1 (2004): 1. http://dx.doi.org/10.11646/zootaxa.782.1.1.

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The new species, Bembidion (Chilioperyphus) cuyanus (type locality ARGENTINA, San Luis, Quebrada Lopez, 7 km SE San Francisco) is described, based on external structural features (extra setae in the supraorbital region, pronotum, elytral margin and intervals 3, 5 and 7) and those of the male genitalia (details of the endophallic flagellum). Line drawings illustrate diagnostic features and a key distinguishes among the species of subgenus Chilioperyphus Jeannel. This species lives along the border of rivers and creeks in the arid region of western Argentina.KEY WORDS. Carabidae. Bembidiini. Bembidion (Chilioperyphus) cuyanus n. sp., ArgentinaSe describe una nueva especie del g nero Bembidion Latreille (Coleoptera: Carabidae: Bembidiini) del centro oeste de la Argentina. Esta ha sido encontrada en las orillas de arroyos y r os de esta regi n semides rtica. La morfolog a externa y las caracter sticas de las estructuras de la genitalia masculina hacen considerar que esta nueva especie pertenece al g nero Bembidion Latreille, y algunos caracteres del ed ago muestran que pertenece al subg nero Chilioperyphus Jeannel. La estructura del flagelo y la presencia de numerosas setas, tanto en la regi n supraorbital, pronoto, borde del litro e interesar as 3, 5 y 7, permiten reconocer a esta como una nueva especie, Bembidion (Chilioperyphus) cuyanus nov. sp. Se provee la descripci n e ilustraciones de esta nueva especie, como as tambi n una clave para diferenciarla de las restantes especies del subg nero Chilioperyphus.PALABRAS CLAVE. Carabidae. Bembidiini. Bembidion (Chilioperyphus) cuyanus n. sp., Argentina
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4

Maddison, David R. "Phylogeny of Bembidion and related ground beetles (Coleoptera: Carabidae: Trechinae: Bembidiini: Bembidiina)." Molecular Phylogenetics and Evolution 63, no. 3 (2012): 533–76. http://dx.doi.org/10.1016/j.ympev.2012.01.015.

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5

Maddison, David R. "Shards, sequences, and shorelines: two new species of Bembidion from North America (Coleoptera, Carabidae)." ZooKeys 1007 (December 30, 2020): 85–128. http://dx.doi.org/10.3897/zookeys.1007.60012.

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Two new species of Bembidion are described from river shores in North America. One, Bembidion mimbressp. nov., from the Gila River watershed in the lands of the Mimbres culture in New Mexico and Arizona, is closely related to the widespread Bembidion levigatum. DNA sequences from several linkage groups and morphology provide evidence of the distinctiveness of B. mimbres. The second, Bembidion corgenomasp. nov., has been the subject of recent genomic and transcriptomic studies. It belongs in the Bembidion transversale subgroup, and occurs from California north to British Columbia, east to Montana and Nevada. The B. transversale subgroup as a whole is reviewed, and morphological characters that distinguish B. corgenoma from the similar and sympatric B. transversale and B. erosum are described and illustrated. DNA sequences of these three species show no consistent differences in 28S, COI, CAD, and Topoisomerase, and a coalescent species delimitation analysis reveals no notable structure within the complex. This is the first known trio of species within Bembidion for which those genes provide no clear signal of species boundaries. A neotype is designated for the one name in the group that lacks a primary type, Bembidium haplogonum Chaudoir. Chromosomes of the new species and their relatives are as is typical for Bembidion, with eleven pairs of autosomes and an XY/XX sex chromosome system.
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6

Maddison, David R. "Shards, sequences, and shorelines: two new species of Bembidion from North America (Coleoptera, Carabidae)." ZooKeys 1007 (December 30, 2020): 85–128. https://doi.org/10.3897/zookeys.1007.60012.

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Two new species of Bembidion are described from river shores in North America. One, Bembidion mimbres sp. nov., from the Gila River watershed in the lands of the Mimbres culture in New Mexico and Arizona, is closely related to the widespread Bembidion levigatum. DNA sequences from several linkage groups and morphology provide evidence of the distinctiveness of B. mimbres. The second, Bembidion corgenoma sp. nov., has been the subject of recent genomic and transcriptomic studies. It belongs in the Bembidion transversale subgroup, and occurs from California north to British Columbia, east to Montana and Nevada. The B. transversale subgroup as a whole is reviewed, and morphological characters that distinguish B. corgenoma from the similar and sympatric B. transversale and B. erosum are described and illustrated. DNA sequences of these three species show no consistent differences in 28S, COI, CAD, and Topoisomerase, and a coalescent species delimitation analysis reveals no notable structure within the complex. This is the first known trio of species within Bembidion for which those genes provide no clear signal of species boundaries. A neotype is designated for the one name in the group that lacks a primary type, Bembidium haplogonum Chaudoir. Chromosomes of the new species and their relatives are as is typical for Bembidion, with eleven pairs of autosomes and an XY/XX sex chromosome system.
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7

Maddison, David R., John S. Sproul, and Howard Mendel. "Origin and adaptive radiation of the exceptional and threatened bembidiine beetle fauna of St Helena (Coleoptera: Carabidae)." Zoological Journal of the Linnean Society 189, no. 4 (2019): 1155–75. http://dx.doi.org/10.1093/zoolinnean/zlz150.

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Abstract The central peaks of the isolated island of St Helena (south Atlantic Ocean) are home to an extraordinary set of ground beetles of the tribe Bembidiini, which belong to three endemic genus-group taxa. These beetles are strikingly different in overall body form from the many bembidiines found elsewhere in the world. At least some of the St Helena species are likely to be extinct, and all are threatened by habitat destruction and invasive species. Through next-generation sequencing of historical museum specimens, we examine the phylogenetic relationships of the St Helena fauna. We find that, in spite of their morphological disparities, the endemic bembidiines of St Helena form a clade of genetically similar species, with their sister group being Bembidion alsium from the Indian Ocean island of La Réunion, and the sister group of this pair being the African subgenus Omotaphus. We propose that the St Helena Peaks Bembidion are an adaptive radiation that arose from a single dispersal event to St Helena from a now-extinct African lineage (sister to Omotaphus) and that this extinct lineage also served as the ancestral source of B. alsium. Given that the St Helena Peaks Bembidion are deeply nested in the genus Bembidion, we move the three taxa back in that genus as subgenera and provide a new name (Bembidion shepherdae) for the now-homonymous Bembidion wollastoni.
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8

Toledano, Luca, and Oto Nakládal. "Bembidion (Plataphodes) kmecoi sp. nov. from China, Sichuan (Coleoptera: Carabidae: Bembidiina)." Zootaxa 2882 (December 31, 2011): 64–68. https://doi.org/10.5281/zenodo.277588.

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9

TOLEDANO, LUCA, and OTO NAKLÁDAL. "Bembidion (Plataphodes) kmecoi sp. nov. from China, Sichuan (Coleoptera: Carabidae: Bembidiina)." Zootaxa 2882, no. 1 (2011): 64. http://dx.doi.org/10.11646/zootaxa.2882.1.7.

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Bembidion kmecoi sp. nov. herewith described, sister species of Bembidion farkaci Toledano & Sciaky, 1998, is the third species of the subgenus Plataphodes Ganglbauer, 1891 known from China. A key for the identification of these species is provided. Another species, B. elatum Andrewes, 1930, formerly known as Plataphodes, is transferred to Blepharoplataphus Netolitzky, 1920. Nomenclatorial acts proposed in this paper: Description of Bembidion (Plataphodes) kmecoi sp. nov. (China); Species transferred from Bembidion subg. Plataphodes Ganglbauer, 1891 to Bembidion subg. Blepharoplataphus Netolitzky, 1920: B. elatum Andrewes, 1930.
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10

Roig-Juñent, Sergio, and Erica Scheibler. "A new Argentinian species of Bembidion (Chilioperyphus) (Coleoptera: Carabidae: Bembidiini) with accessory setae." Zootaxa 782 (December 31, 2004): 1–10. https://doi.org/10.5281/zenodo.158016.

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Roig-Juñent, Sergio, Scheibler, Erica (2004): A new Argentinian species of Bembidion (Chilioperyphus) (Coleoptera: Carabidae: Bembidiini) with accessory setae. Zootaxa 782: 1-10, DOI: 10.5281/zenodo.158016
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11

Larochelle, André, Zeperyphodes, Broun, and Marie-Claude Larivière. "Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini)." Insecta Mundi 2015, no. 415 (2015): 1–78. https://doi.org/10.5281/zenodo.5181756.

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Larochelle, André, Zeperyphodes, Broun, Larivière, Marie-Claude (2015): Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 2015 (415): 1-78, DOI: 10.5281/zenodo.5181756
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12

Avgin, Sakine Serap. "A Synopsis of the SubgenusTestedium Motschulsky, 1864 (Coleoptera: Carabidae: Bembidiini: Bembidion Latreille) in Turkey, with One New Country Record." Coleopterists Bulletin 65, no. 1 (2011): 39–42. https://doi.org/10.1649/0010-065x-65.1.39.

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Avgin, Sakine Serap (2011): A Synopsis of the SubgenusTestedium Motschulsky, 1864 (Coleoptera: Carabidae: Bembidiini: Bembidion Latreille) in Turkey, with One New Country Record. The Coleopterists Bulletin 65 (1): 39-42, DOI: 10.1649/0010-065x-65.1.39
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13

Maddison, David R., Luca Toledano, Soledad Sallenave, and Sergio Roig-Juñent. "Phylogenetic relationships of the South American ground beetle subgenus Chilioperyphus Jeannel (Coleoptera: Carabidae: Trechinae: Bembidiini: Bembidion Latreille)." Zootaxa 3636, no. 4 (2013): 547–60. https://doi.org/10.11646/zootaxa.3636.4.3.

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Maddison, David R., Toledano, Luca, Sallenave, Soledad, Roig-Juñent, Sergio (2013): Phylogenetic relationships of the South American ground beetle subgenus Chilioperyphus Jeannel (Coleoptera: Carabidae: Trechinae: Bembidiini: Bembidion Latreille). Zootaxa 3636 (4): 547-560, DOI: 10.11646/zootaxa.3636.4.3
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14

Maddison, David, and Karen Ober. "Phylogeny of minute carabid beetles and their relatives based upon DNA sequence data (Coleoptera, Carabidae, Trechitae)." ZooKeys 147 (November 16, 2011): 229–60. https://doi.org/10.3897/zookeys.147.1871.

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The phylogeny of ground beetles of supertribe Trechitae is inferred using DNA sequences of genes that code for 28S ribosomal RNA, 18S ribosomal RNA, and <i>wingless</i>. Within the outgroups, austral psydrines are inferred to be monophyletic, and separate from the three genera of true Psydrina (<i>Psydrus, Nomius, Laccocenus</i>); the austral psydrines are formally removed from Psydrini and are treated herein as their own tribe, Moriomorphini Sloane. All three genes place <i>Gehringia</i> with Psydrina. Trechitae is inferred to be monophyletic, and sister to Patrobini.Within trechites, evidence is presented that <i>Tasmanitachoides</i> is not a tachyine, but is instead a member of Trechini. <i>Perileptus</i> is a member of subtribe Trechodina. Against Erwin’s hypothesis of anillines as a polyphyletic lineage derived from the tachyine genus <i>Paratachys</i>, the anillines sampled are monophyletic, and not related to <i>Paratachys</i>. Zolini, Pogonini, Tachyina, and Xystosomina are all monophyletic, with the latter two being sister groups. The relationships of the subtribe Bembidiina were studied in greater detail. <i>Phrypeus</i> is only distantly related to <i>Bembidion</i>, and there is no evidence from sequence data that it belongs within Bembidiina. Three groups that have been recently considered to be outside of the large genus <i>Bembidion</i> are shown to be derived members of <i>Bembidion</i>, related to subgroups: <i>Cillenus</i> is related to the <i>Ocydromus</i> complex of <i>Bembidion</i>, <i>Zecillenus</i> is related to the New Zealand subgenus <i>Zeplataphus</i>, and <i>Hydrium</i> is close to subgenus <i>Metallina</i>. The relationships among major lineages of Trechitae are not, however, resolved with these data.
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Toledano, Luca, and Borislav Guéorguiev. "Systematic, synonymic and geographic notes on Bembidion (Terminophanes) pulcherrimum (Motschulsky, 1850) and its closest relatives (Coleoptera: Carabidae: Bembidiini)." Ecologica Montenegrina 2, no. 4 (2015): 295–301. https://doi.org/10.37828/em.2015.2.35.

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Toledano, Luca, Guéorguiev, Borislav (2015): Systematic, synonymic and geographic notes on Bembidion (Terminophanes) pulcherrimum (Motschulsky, 1850) and its closest relatives (Coleoptera: Carabidae: Bembidiini). Ecologica Montenegrina 2 (4): 295-301, DOI: 10.37828/em.2015.2.35, URL: https://www.biotaxa.org/em/article/view/em.2015.2.35
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16

Schmidt, Joachim, and Peter Michalik. "The ground beetle genus Bembidion Latreille in Baltic amber: Review of preserved specimens and first 3D reconstruction of endophallic structures using X-ray microscopy (Coleoptera, Carabidae, Bembidiini)." ZooKeys 662 (March 22, 2017): 101–26. https://doi.org/10.3897/zookeys.662.12124.

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The ground beetle genus Bembidion is a highly diverse group of small predators with more than 1.200 described extant species. In contrast, only two representatives of Bembidion are known from the amber fossil record and their position within this mega-diverse genus is dubious. Here, we address the taxonomic position of these two extinct Bembidion species (B. succini Giebel, 1856 and B. christelae Ortuño &amp; Arillo, 2010). Based on the insufficient description and the missing type specimen, B. succini, nomen dubium, cannot be assigned to the genus Bembidion and/or to the tribe Bembidiini with certainty. The subgenus Archaeophilochthus Ortuño &amp; Arillo, 2010 was erected for the second extinct species, B. christelae, based on external characters. However, this species seems indistinguishable to members of the earlier described subgenus Philochthemphanes Netolitzky, 1943 which comprises about extant 10 species distributed in East and Southeast Asia. Furthermore, we describe two new species, B. bukejsi sp. n. and B. alekseevi sp. n., from the Eocene Baltic amber using X-ray microscopy. Based on external and genital morphology including endophallic structures, we erected the monotypic subgenus Eodontium subgen. n. for B. bukejsi sp. n., which is probably related to the subgenera Andrewesa Netolitzky, 1931, the Hydrium complex, or the Odontium series sensu Maddison (2012). On the other hand, B. alekseevi sp. n. can be assigned to the subgenus Eupetedromus Netolitzky, 1911. The occurrence of representatives of at least two species groups adapted to a temperate climate suggests the presence of at least locally temperate climates in Baltic amber forests.
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17

Neri, Paolo, and Luca Toledano. "Bembidion (Ocydromus) terryerwini sp. nov. from Iran (Coleoptera, Carabidae, Bembidiina)." ZooKeys 1044 (June 16, 2021): 221–28. http://dx.doi.org/10.3897/zookeys.1044.63607.

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Bembidion (Ocydromus) terryerwinisp. nov. from Central Iran (Kerman and Yazd Provinces), described here, belongs to the decorum species group (Ocydromus Clairville, 1806). The new species is compared with the other taxa of this species group occurring in Iran and neighboring regions (Georgia, Armenia, Azerbaijan, Turkey, Iraq, Iran, South Pakistan, Afghanistan, and Turkmenistan).
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18

Neri, Paolo, and Luca Toledano. "Bembidion (Ocydromus) terryerwini sp. nov. from Iran (Coleoptera, Carabidae, Bembidiina)." ZooKeys 1044 (June 16, 2021): 221–28. https://doi.org/10.3897/zookeys.1044.63607.

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Bembidion (Ocydromus) terryerwini sp. nov. from Central Iran (Kerman and Yazd Provinces), described here, belongs to the decorum species group (Ocydromus Clairville, 1806). The new species is compared with the other taxa of this species group occurring in Iran and neighboring regions (Georgia, Armenia, Azerbaijan, Turkey, Iraq, Iran, South Pakistan, Afghanistan, and Turkmenistan).
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19

Neri, Paolo, and Luca Toledano. "Notes on taxonomy and geography of Bembidion (Neja ) leucoscelis Chaudoir and Bembidion (Neja ) curtulum Duval (Coleoptera: Carabidae: Bembidiina)." Entomologist's Monthly Magazine 160, no. 1 (2024): 53–66. http://dx.doi.org/10.31184/m00138908.1601.4208.

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The systematic relationship of Bembidion ( Neja ) leucoscelis Chaudoir, 1850, and Bembidion ( Neja ) curtulum Duval, 1851, is examined. A new subspecies of B. curtulum from the Middle East is described and the subspecies tripolitanum Schatzmayr, 1937, is assigned to Bembidion curtulum (comb. n). The distribution of the species and subspecies is re-examined and the main diagnostic characters emphasized, including the importance of aedeagal structure and elytral microsculpture. The species Bembidion ( Chlorodium ) almum almum Sahlberg, 1900, is reported for Armenia and the Altaj Republic (Russia). A key to the species of Bembidion subgenus Neja Motschulsky, 1864, with elytra not microsculptured or with microsculpture restricted only at the extreme of elytral apex, is provided.
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20

Raupach, Michael, Karsten Hannig, Jerome Moriniere, and Lars Hendrich. "A DNA barcode library for ground beetles (Insecta, Coleoptera, Carabidae) of Germany: The genus Bembidion Latreille, 1802 and allied taxa." ZooKeys 592 (May 25, 2016): 121–41. https://doi.org/10.3897/zookeys.592.8316.

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As molecular identification method, DNA barcoding based on partial cytochrome c oxidase subunit 1 (COI) sequences has been proven to be a useful tool for species determination in many insect taxa including ground beetles. In this study we tested the effectiveness of DNA barcodes to discriminate species of the ground beetle genus Bembidion and some closely related taxa of Germany. DNA barcodes were obtained from 819 individuals and 78 species, including sequences from previous studies as well as more than 300 new generated DNA barcodes. We found a 1:1 correspondence between BIN and traditionally recognized species for 69 species (89%). Low interspecific distances with maximum pairwise K2P values below 2.2% were found for three species pairs, including two species pairs with haplotype sharing (Bembidion atrocaeruleum/Bembidion varicolor and Bembidion guttula/Bembidion mannerheimii). In contrast to this, deep intraspecific sequence divergences with distinct lineages were revealed for two species (Bembidion geniculatum/Ocys harpaloides). Our study emphasizes the use of DNA barcodes for the identification of the analyzed ground beetles species and represents an important step in building-up a comprehensive barcode library for the Carabidae in Germany and Central Europe as well.
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21

MADDISON, DAVID R., and A. ELIZABETH ARNOLD. "A review of the Bembidion (Odontium) aenulum subgroup (Coleoptera: Carabidae), with description of a new species." Zootaxa 2214, no. 1 (2009): 45–61. http://dx.doi.org/10.11646/zootaxa.2214.1.3.

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The Bembidion aenulum subgroup of the subgenus Odontium is reviewed. This subgroup, widespread in the United States east of the Rocky Mountains, previously has been considered to consist of only one species, Bembidion aenulum Hayward. However, morphological studies coupled with analyses of 28S ribosomal RNA, cytochrome oxidase I, CAD, and wingless genes reveal that eastern members of this group belong to a new species herein named Bembidion paraenulum new species. This species ranges from New Hampshire in the northeast, through Virginia and North Carolina, south to Florida and west to Mississippi. Both species are described and illustrated.
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22

Makarov, Kirill Vladimirovich, Yurii Nikolaevich Sundukov, and Andrey Vladimirovich Matalin. "Ground beetles (Coleoptera, Carabidae) in fumarole fields of Kunashir Island, Kuril Archipelago, Russia." Acta Zoologica Academiae Scientiarum Hungaricae 66, Suppl. (2020): 97–146. http://dx.doi.org/10.17109/azh.66.suppl.97.2020.

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Five species of ground beetles are permanent inhabitants of the fumarola fields on Kunashir Island: Cicindela (Cicindela) sachalinensis A. Morawitz, 1862; Cylindera (Eugrapha) elisae (Motschulsky, 1859); Bembidion (Ocydromus) dolorosum (Motschulsky, 1860); B. (Peryphanes) sanatum Bates, 1883, and Poecilus (Poecilus) samurai (Lutshnik, 1916). These species respond differently to extreme conditions. In some species, the size is decreased (C. elisae, B. dolorosum), but is increased in P. samurai; in B. dolorosum, the pigmentation is decreased, while increased in others (C. sachalinensis, C. elisae, P. samurai). The degree of these variations depends neither on taxonomic relations nor the adaptation time. The areas of moderate thermal activity of Kunashir volcanoes could have served as refugia during the colder climatic periods. Based on data on the variability and barcoding of B. dolorosum, the following new synonymy is established: Bembidion (Ocydromus) dolorosum (Motschulsky, 1860) = Bembidion (Ocydromus) negrei Habu, 1958, syn. nov. = Bembidion (Peryphus) kuznetsovi Lafer, 2002, syn. nov.
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23

Hildebrandt, Drew, and David Maddison. "A new species of Bembidion Latrielle 1802 from the Ozarks, with a review of the North American species of subgenus Trichoplataphus Netolitzky 1914 (Coleoptera, Carabidae, Bembidiini)." ZooKeys 147 (November 16, 2011): 261–75. https://doi.org/10.3897/zookeys.147.1872.

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A new species of <i>Bembidion</i> (<i>Trichoplataphus </i>Netolitzky) from the Ozark Plateau of Missouri and Arkansas is described (<i>Bembidion ozarkense</i> Maddison and Hildebrandt). It is distinguishable from the closely related species, <i>B. rolandi</i> Fall, by characteristics of the male genitalia, and sequences of the genes cytochrome oxidase I and 28S ribosomal DNA. A brief review of the North American species of <i>Trichoplataphus</i> is presented, including a key to species.
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24

SASAKAWA, KÔJI. "Taxonomic studies on the Bembidion (‘Cillenus’) complex (Coleoptera: Carabidae): A revision of the subgeneric taxonomy and description of a new species from Japan." Zootaxa 1575, no. 1 (2007): 35–45. http://dx.doi.org/10.11646/zootaxa.1575.1.2.

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The supraspecific taxonomy of the coastal carabid Bembidion (‘Cillenus’) complex is controversial because a single ‘synapomorphy,’ a large head with long mandibles, may be the result of functional convergence. This study examined the phylogeny of B. (‘Cillenus’) complex based on morphological characters to revise the complicated taxonomy of this group. All type species of the genera concerned were included, and membranous parts of the genitalia (male endophallus and female vagina and spermatheca) were examined for the first time. Because all species are treated under the genus Bembidion Latreille (s. l.), Bembidion umi nom. nov. is proposed as a new name for Sakagutia marina Uéno. In the resultant tree, reciprocal monophyly of most traditional taxa was not supported, whereas a monophyletic group of the following seven species was relatively well supported: B. seticorne Lindroth, B. yokohamae (Bates), B. sumaoi Morita, B. aestuarii (Uéno &amp; Habu), B. tokunoshimanum (Nakane), B. tsutsuii (Uéno), and B. yakushimanum sp. nov. Based on these results, the subgenus Desarmatocillenus Netolitzky (type species: B. yokohamae) was redefined, and two subgenera were synonymised with it: Novicillenus Uéno &amp; Habu (type species: B. aestuarii) syn. nov., and Corallicillenus Uéno (type species: B. tsutsuii) syn. nov.. Bembidion (Desarmatocillenus) yakushimanum sp. nov. is described from Japan.
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25

Liebherr, James K., and David R. Maddison. "Colonisation of the Pacific by Bembidion beetles (Coleoptera : Carabidae), with description of Bembidion tahitiense, sp. nov. from Tahiti, French Polynesia." Invertebrate Systematics 27, no. 4 (2013): 439. http://dx.doi.org/10.1071/is13003.

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Bembidion (Sloanephila) tahitiense, sp. nov. is described from Mont Mauru, an isolated massif of Tahiti Nui volcano. Based on evidence from seven genes (four nuclear protein-coding, one mitochondrial protein-coding, two nuclear ribosomal), its sister group is the Australian B. jacksoniense Guérin-Méneville, with which it shares a synapomorphic spur on the ostium of the male genitalia. In contrast to B. jacksoniense, B. tahitiense is brachypterous, with rounded humeri, constricted posterior pronotal margins and convex body form. Examination of the seven genes in two species of the Hawaiian subgenus Nesocidium Sharp reveals that the sister group of Nesocidium is subgenus Zecillenus Lindroth from New Zealand. These two subgenera belong to the Ananotaphus complex, a clade inhabiting Australia, New Zealand and Hawaii. The relationships of the second Hawaiian subgenus, Gnatholymnaeum Sharp, are less clear, although Gnatholymnaeum belongs to the Bembidion series (along with Sloanephila and the Ananotaphus complex). Bembidion beetles colonised the Society and Hawaiian islands independently from source areas in the south-west Pacific. Based on parsimonious reconstructions of flight-wing configuration, the Tahitian and Hawaiian colonisations involved winged individuals. Colonisation of the Society and Hawaiian islands by carabid beetles of two other tribes – Platynini and Moriomorphini – follow the dispersal patterns hypothesised for Bembidion.
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26

Maddison, David. "An unexpected clade of South American ground beetles (Coleoptera, Carabidae, Bembidion)." ZooKeys 416 (June 17, 2014): 113–55. https://doi.org/10.3897/zookeys.416.7706.

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Phylogenetic relationships of the <em>Antiperyphanes </em>Complex of the genus <em>Bembidion</em> are inferred using DNA sequences from seven genes (two nuclear ribosomal, four nuclear protein coding, and one mitochondrial protein coding). Redefined subgenera within the complex are each well-supported as monophyletic. Most striking was the discovery that a small set of morphologically and ecologically heterogeneous species formed a clade, here called subgenus <em>Nothonepha</em>. This unexpected result was corroborated by the discovery of deep pits in the lateral body wall (in the mesepisternum) of all <em>Nothonepha</em>, a trait unique within <em>Bembidion</em>. These pits are filled with a waxy substance in ethanol-preserved specimens. In one newly discovered species (<em>Bembidion tetrapholeon</em> <strong>sp. n.</strong>, described here), these pits are so deep that their projections into the body cavity from the two sides touch each other internally. These structures in <em>Bembidion</em> (<em>Nothonepha</em>) are compared to very similar mesepisternal pits which have convergently evolved in two other groups of carabid beetles. The function of these thoracic pits is unknown. Most members of subgenus <em>Nothonepha</em> have in addition similar but smaller pits in the abdomen. A revised classification is proposed for the <em>Antiperyphanes</em> Complex.
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27

Sundukov, Yu N. "NEW OR LITTLE-KNOWN SPECIES OF GROUND BEETLES (COLEOPTERA, CARABIDAE) FROM THE SAKHALIN ISLAND." Зоологический журнал 102, no. 3 (2023): 291–99. http://dx.doi.org/10.31857/s0044513423030145.

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Information concerning 20 new or little-known species of ground beetles found in the south of Sakhalin is presented. Of these, two species (Bembidion pliculatum Bates, 1883, Apristus grandis Andrewes 1937) are new to the fauna of Russia, while further five species (Dyschirius tristis Stephens 1827, Bembidion pseudolucillum Netolitzky 1938, Trechus nigricornis arsenjevi (Jeannel 1962), Harpalus nigrans A. Morawitz 1862, Lebia bifenestrata A. Morawitz 1862) represent the first formal records from Sakhalin Island. The earlier records of 13 species from this island are supported by additional material: Carabus canaliculatus sichotensis Born 1914, Dyschirius aeneus ovicollis Putzeys 1873, Eobroscus lutshniki (Roubal 1928), Bembidion hiogoense Bates 1873, B. lucillum lucillum Bates 1883, B. varium (Olivier 1795), Pterostichus longinquus Bates 1873, Agonum consimile (Gyllenhal 1810), Amara erratica (Duftschmid 1812), A. hicksi Lindroth, 1968, Harpalus platynotus Bates 1873, Chlaenius inops inops Chaudoir 1856, and Parena tripunctata (Bates 1873).
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28

Avgin, Serap. "THE TRIBE BEMBIDIINI IN SOUTHERN TURKEY, WITH A NEW RECORD OF BEMBIDION (COLEOPTERA: CARABIDAE)." Entomological News 117, no. 1 (2006): 109–14. http://dx.doi.org/10.3157/0013-872x(2006)117[109:ttbist]2.0.co;2.

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29

Maddison, David, and Luca Toledano. "A new species of Bembidion (Ecuadion) from Ecuador (Coleoptera, Carabidae, Bembidiini), with a key to members of the georgeballi species group." ZooKeys 249 (December 10, 2012): 51–60. https://doi.org/10.3897/zookeys.249.4149.

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A new species of ground beetle, Bembidion ricei, is described from the Andes mountains of Ecuador east of Quito. It belongs to the georgeballi species group of subgenus Ecuadion, and is most similar to B. georgeballi. A key to the species of the group is provided.
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30

Maddison, David R., and Roy Anderson. "Hidden species within the genus Ocys Stephens: the widespread species O. harpaloides (Audinet-Serville) and O. tachysoides (Antoine) (Coleoptera, Carabidae, Bembidiini)." Deutsche Entomologische Zeitschrift 63, no. (2) (2016): 287–301. https://doi.org/10.3897/dez.63.10748.

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Beetles previously considered to be Ocys harpaloides (Audinet-Serville) from northern Africa, Spain, France, the United Kingdom, France, and Belgium belong to two species. These species can be distinguished using DNA sequences of 28S rDNA, 18S rDNA, COI, CAD, and topoisomerase I. A key, diagnoses, and images are provided to enable identification of specimens based upon characteristics of male and female genitalia, as well as microsculpture and other external structures. Through examination of the holotype of Bembidium harpaloides v. tachysoides Antoine, as well as designation of lectotypes of Bembidion harpaloides Audinet-Serville and Ocys melanocephalus Stephens, and designation of a neotype for Tachis rufescens Guérin-Ménéville, the valid names of the two species were determined to be Ocys harpaloides and Ocys tachysoides (Antoine).
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31

Maddison, David R., John S. Sproul, and Kipling Will. "Re-collected after 55 years: a new species of Bembidion (Coleoptera, Carabidae) from California." ZooKeys 1156 (March 27, 2023): 87–106. https://doi.org/10.3897/zookeys.1156.101072.

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A new species of the carabid beetle genus Bembidion Latreille is described from the Central Valley, Los Angeles Basin, and surrounding areas of California. Bembidion brownorum sp. nov. is a distinctive species, a relatively large member of the subgenus Notaphus Dejean, and within Notaphus a member of the B. obtusangulum LeConte species group. It has faint spots on the elytra and a large, convex, rounded prothorax. Of the 22 specimens from 11 localities, all but one were collected more than 55 years ago. Although the collection of the holotype in 2021 at UV light suggest the species is still extant, the lack of other recent specimens suggests the species may have a more restricted distribution than in the past, and its populations may be in decline.
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32

Rozek, Maria. "C-bands and NORs on Chromosomes of Bembidion lampros (Herbst) and Bembidion properans (Steph.) (Coleoptera, Carabidae)." CYTOLOGIA 63, no. 3 (1998): 317–21. http://dx.doi.org/10.1508/cytologia.63.317.

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33

Guseva, Olga G., and Alexander G. Koval. "Distribution of ground beetles of the genus Bembidion (Coleoptera, Carabidae) in the agricultural landscape in Northwestern Russia." Acta Biologica Sibirica 7 (September 6, 2021): 227–36. http://dx.doi.org/10.3897/abs.7.e70229.

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We have observed seven species of predator beetles of the genus Bembidion in the agricultural landscape of the Leningrad Region (Northwestern Russia) between the years 2008 and 2018. These species reach their highest abundance in sun-exposed areas, especially mesophilic B. quadrimaculatum Linnaeus, 1761, B. properans (Stephens, 1828), B. lampros (Herbst, 1784), and B. femoratum Sturm, 1825. A few mesohygrophilic B. guttula (Fabricius, 1792) and B. gilvipes (Sturm, 1825) live primarily in areas of dense vegetation. Finally, the hygrophilic B. bruxellense (Wesmael, 1835) was only observed on the most humid soils. Assemblages of Bembidion ground beetles were separated in the fields, field boundaries, and adjacent habitats.
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34

BOUSQUET, YVES, and REGINALD WEBSTER. "Descriptions of three new species of Bembidion Latreille (Coleoptera: Carabidae) occurring in Canada." Zootaxa 1297, no. 1 (2006): 23. http://dx.doi.org/10.11646/zootaxa.1297.1.3.

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Three new species of Bembidion Latreille occurring in Canada are described: B. (Semicampa) nigrivestis Bousquet, sp. nov., type locality: Sainte-Catherine-de-Hatley (45°15'N, 72°03'W), Quebec; B. (Eupetedromus) iridipenne Bousquet and Webster, sp. nov., type locality: Lincoln (45° 49'N, 66°35'W), New Brunswick; and B. (Neobembidion) nitidicolle Bousquet, sp. nov., type locality: Rosefield (49°05'N, 107°32'W), Saskatchewan. A new subgenus, Neobembidion Bousquet, subgen. nov. (type species: Bembidion constricticolle Hayward, 1897), is proposed for the species of the constricticolle group as defined by Lindroth. Keys are included to distinguish the new species from their North American relatives.
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35

Maddison, David, Kipling Will, Sarah Crews, and James LaBonte. "Bembidion ambiguum (Coleoptera: Carabidae) is established in California." Biodiversity Data Journal 6 (December 4, 2018): e30763. https://doi.org/10.3897/BDJ.6.e30763.

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The ground beetle Bembidion (Neja) ambiguum Dejean is native to Europe and north Africa, in the Mediterranean region. We report it from North America for the first time, from five localities around San Francisco Bay, California. The earliest record is from 2012.
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36

Halimov, Fazlitdin. "Seasonal dynamics of dominant species of soil predators (Coleoptera: Carabidae, Staphylinidae) in agrolandscapes and their potential gluttony." Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 63, no. 2 (2020): 175–87. http://dx.doi.org/10.3897/travaux.63.e54116.

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We studied the seasonal change in the number of dominant species of predatory beetles on vegetable crops and determined their potential gluttony. A total of 1,472 beetles were collected, belonging to 22 species of ground beetles (Carabidae) and 27 species of rove beetles (Staphylinidae). The dominant species are: Harpalus rufipes (De Geer, 1774) (17.6% dominance), Amara fulva (De Geer) (13.28%), Bembidion properans (Stephens, 1828) (10.39%), Trechus quadristriatus (Schrank, 1781) (6.20%), Calathus melanocephalus (Linnaeus, 1758) (5.39%), Poecilus cupreus Linnaeus, 1758 (5.3%),&amp;nbsp;Bembidion femoratum Sturm, 1825 (5.10%), Aleochara bilineata, (Gyllenhaal, 1810) (17.6%), Aloconota gregagia (Erichson, 1839) (10.21%), Amischa analis (Gravenhorst, 1802) (6.01%), Amischa bifoveotata&amp;nbsp;(Mannerheim, 1830) (5.41%). During the season, there is a change in dominant species. At the beginning of the season, smaller species dominate, while larger species dominate in the second half of the plant vegetation. The maximum number of predators are observed in June and August. The periods of the maximum abundance of ground beetles and rove beetles do not coincide in time, which is the evolutionary adaptation of two groups of predators that coexist. In laboratory experiments, when pest eggs were offered as food, Aleochara bilineata and Bembidion femoratum were the most voracious. Larger predator species showed high voracity when larvae of flies were offered as food.
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37

Bousquet, Yves, and Reginald Webster. "Descriptions of three new species of Bembidion Latreille (Coleoptera: Carabidae) occurring in Canada." Zootaxa 1297 (December 31, 2006): 23–35. https://doi.org/10.5281/zenodo.173556.

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38

Avgin, Sakine Serap, and Owe Nodmar. "Expansion of Distribution Records for Nine Subfamilies Of Carabidae (Coleoptera) With Two New Records in Turkey." Journal of Entomological Science 44, no. 4 (2009): 323–40. http://dx.doi.org/10.18474/0749-8004-44.4.323.

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Carabid (Coleoptera: Carabidae) specimens from various environs in Turkey were examined. Eighty-one species/subspecies representing 20 genera in 9 subfamilies were identified. The subfamilies represented were Brachininae (5 species/subspecies), Broscinae (1 species), Carabinae (24 species/subspecies), Elaphrinae (1 species), Nebriinae (6 species/subspecies), Omophorninae (1 species), Paussinae (1 species), Scartinae (5 species/subspecies), and Trechinae (37 species/subspecies). All but a few of these species/subspecies are new records of distribution in several Turkish provinces. Furthermore, identification of Bembidion (Be-mbidionetolitzkya) cyaneum Chaudoir, 1846 and Bembidion (Bembidionetolitzkya) coeruleum Audinet-Serville, 1821 (Coleoptera: Carabidae: Trechinae) are the first recorded occurrences of these species in Turkey. Previously published records from Turkey, geographical distribution and chorotype information are provided for each species.
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39

Andersen, Johan, and Arne Skorping. "Parasites of carabid beetles: prevalence depends on habitat selection of the host." Canadian Journal of Zoology 69, no. 5 (1991): 1216–20. http://dx.doi.org/10.1139/z91-173.

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The prevalence of endoparasitic nematodes (Mermithoidea and Heterotylenchus sp.) and the ectoparasitic fungi Laboulbenia spp. was examined in 14 riparian species of carabid beetles of the genus Bembidion. The values for Laboulbenia spp. and nematodes (Heterotylenchus sp) were 41.1 and 11.4%, respectively, in host species living in protected microhabitats (silty, more or less vegetated, often shady sites). In Bembidion species living in open microhabitats (open silty, sandy, or gravelly–stony sites with little or no vegetation) the prevalence of both parasites was less than 1%. The open microhabitats are more subject to floodwater scouring and erosion than the protected ones. This may explain the observed difference in prevalence, since the beetles are infected with the parasites through contact with contaminated soil.
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40

Maddison, David R., John S. Sproul, and Kipling Will. "Re-collected after 55 years: a new species of Bembidion (Coleoptera, Carabidae) from California." ZooKeys 1156 (March 27, 2023): 87–106. http://dx.doi.org/10.3897/zookeys.1156.101072.

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A new species of the carabid beetle genus Bembidion Latreille is described from the Central Valley, Los Angeles Basin, and surrounding areas of California. Bembidion brownorumsp. nov. is a distinctive species, a relatively large member of the subgenus Notaphus Dejean, and within Notaphus a member of the B. obtusangulum LeConte species group. It has faint spots on the elytra and a large, convex, rounded prothorax. Of the 22 specimens from 11 localities, all but one were collected more than 55 years ago. Although the collection of the holotype in 2021 at UV light suggest the species is still extant, the lack of other recent specimens suggests the species may have a more restricted distribution than in the past, and its populations may be in decline.
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41

Guseva, Olga G., and Alexander G. Koval. "Distribution of ground beetles of the genus Bembidion (Coleoptera, Carabidae) in the agricultural landscape in Northwestern Russia." Acta Biologica Sibirica 7 (September 6, 2021): 227–36. https://doi.org/10.3897/abs.7.e70229.

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We have observed seven species of predator beetles of the genus <em>Bembidion</em> in the agricultural landscape of the Leningrad Region (Northwestern Russia) between the years 2008 and 2018. These species reach their highest abundance in sun-exposed areas, especially mesophilic <em>B. quadrimaculatum</em> Linnaeus, 1761, <em>B. properans</em> (Stephens, 1828), <em>B. lampros</em> (Herbst, 1784), and <em>B. femoratum</em> Sturm, 1825. A few mesohygrophilic <em>B. guttula</em> (Fabricius, 1792) and <em>B. gilvipes</em> (Sturm, 1825) live primarily in areas of dense vegetation. Finally, the hygrophilic <em>B. bruxellense</em> (Wesmael, 1835) was only observed on the most humid soils. Assemblages of <em>Bembidion</em> ground beetles were separated in the fields, field boundaries, and adjacent habitats.
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42

Rutschmann, Florin, Jürg Schlegel, and Werner A. Marggi. "Bembidion (Peryphanes) italicum De Monte, 1943 - Erstnachweis nördlich der Schweizer Alpen (Coleoptera: Carabidae)." Entomo Helvetica 5 (December 31, 2012): 89–94. https://doi.org/10.5169/seals-986124.

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Rutschmann, Florin, Schlegel, Jürg, Marggi, Werner A. (2012): Bembidion (Peryphanes) italicum De Monte, 1943 - Erstnachweis nördlich der Schweizer Alpen (Coleoptera: Carabidae). Entomo Helvetica 5: 89-94, DOI: 10.5169/seals-986124
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43

Maddison, David R., and Elizabeth Arnold. "A review of the Bembidion (Odontium) aenulum subgroup (Coleoptera: Carabidae), with description of a new species." Zootaxa 2214 (December 31, 2009): 45–61. https://doi.org/10.5281/zenodo.189922.

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Maddison, David R., Arnold, Elizabeth (2009): A review of the Bembidion (Odontium) aenulum subgroup (Coleoptera: Carabidae), with description of a new species. Zootaxa 2214: 45-61, DOI: 10.5281/zenodo.189922
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44

Ruchin, Alexander, Sergey Alekseev, Leonid Egorov, Oleg Artaev, Gennadiy Semishin, and Mikhail Esin. "Ground beetle fauna (Coleoptera, Carabidae) of Mordovia State Nature Reserve (Russia)." Biodiversity Data Journal 9 (October 25, 2021): e69807. https://doi.org/10.3897/BDJ.9.e69807.

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Protected areas are organised in different climatic zones, which usually include typical ecosystems characteristic of certain climatic zones. In most cases, protected areas are biodiversity hotspots. These areas are benchmarks in terms of nature conservation and to determine their biological diversity is becoming an important task. It is important to investigate the carabid family of protected areas within the framework of understanding the overall biological diversity of these systems. In addition, ground beetles, as one of the largest groups of ground-based inhabitants, are indicators of the state of ecosystems and serve as markers of their well-being.We present 2,969 new occurrence records comprising 226 species of Carabidae, belonging to eight subfamilies, from the Mordovia State Nature Reserve (central Russia). Ten species are listed for the first time for the Mordovia State Nature Reserve fauna after previous research: <i>Cicindela maritima</i>, <i>Bembidion striatum</i>, <i>Dyschirius angustatus</i>, <i>Dyschirius arenosus</i>, <i>Notiophilus aestuans</i>, <i>Bembidion argenteolum</i>, <i>Bembidion velox</i>, <i>Bradycellus caucasicus</i>, <i>Cymindis angularis</i> and <i>Syntomus truncatellus</i>, five of which were first recorded for the Republic of Mordovia (Egorov et al. 2020). Previously, this information was not published anywhere and we wanted to make it available to everyone by embedding it in the global database on biodiversity (GBIF).
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45

Davidson, Robert, and Jessica Ryyken. "Rediscovery of Bembidion (Lymnaeum) nigropiceum (Marsham) (= puritanum Hayward) in Massachusetts, with remarks on biology and habitat (Coleoptera, Carabidae, Bembidiini)." ZooKeys 147 (November 16, 2011): 487–96. https://doi.org/10.3897/zookeys.147.2105.

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<i>Bembidion (Lymnaeum) nigropiceum </i>(Marsham) (=<i>puritanum </i>Hayward), a European species introduced into Massachusetts but presumed not to have become established, has been rediscovered during the Boston Harbor Islands All Taxa Biodiversity Inventory undertaken by the Museum of Comparative Zoology and the National Park Service. A summary is presented of treatment of this species in North America. Data on specimens collected are presented, along with observations on habitat and biology. Some speculations are presented about its highly specialized habitat in the gravel pushed up by high tide, which may act as a food-trapping sieve. A few words are included about future actions needed to resolve questions of distribution and behavior.
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46

Halimov, Fazlitdin. "Seasonal dynamics of dominant species of soil predators (Coleoptera: Carabidae, Staphylinidae) in agrolandscapes and their potential gluttony." Travaux du Muséum National d'Histoire Naturelle "Grigore Antipa" 63, no. (2) (2020): 175–87. https://doi.org/10.3897/travaux.63.e54116.

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We studied the seasonal change in the number of dominant species of predatory beetles on vegetable crops and determined their potential gluttony. A total of 1,472 beetles were collected, belonging to 22 species of ground beetles (Carabidae) and 27 species of rove beetles (Staphylinidae). The dominant species are: <em>Harpalus rufipes </em>(De Geer, 1774) (17.6% dominance), <em>Amara fulva </em>(De Geer) (13.28%), <em>Bembidion properans </em>(Stephens, 1828) (10.39%), <em>Trechus quadristriatus </em>(Schrank, 1781) (6.20%), <em>Calathus melanocephalus </em>(Linnaeus, 1758) (5.39%), <em>Poecilus cupreus </em>Linnaeus, 1758 (5.3%), <em>Bembidion femoratum </em>Sturm, 1825 (5.10%), <em>Aleochara bilineata, </em>(Gyllenhaal, 1810) (17.6%), <em>Aloconota gregagia </em>(Erichson, 1839) (10.21%), <em>Amischa analis </em>(Gravenhorst, 1802) (6.01%), <em>Amischa bifoveotata</em> (Mannerheim, 1830) (5.41%). During the season, there is a change in dominant species. At the beginning of the season, smaller species dominate, while larger species dominate in the second half of the plant vegetation. The maximum number of predators are observed in June and August. The periods of the maximum abundance of ground beetles and rove beetles do not coincide in time, which is the evolutionary adaptation of two groups of predators that coexist. In laboratory experiments, when pest eggs were offered as food, <em>Aleochara bilineata </em>and <em>Bembidion femoratum </em>were the most voracious. Larger predator species showed high voracity when larvae of flies were offered as food.
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47

Makarov, K.V., and Yu.N. Sundukov. "A new subspecies of Bembidion sanatum (Coleoptera: Carabidae) endemic to the Mendeleev Volcano (Kunashir Island, Russia)." Far Eastern Entomologist 440 (November 2, 2021): 13–24. https://doi.org/10.25221/fee.440.2.

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Abstract:
Makarov, K.V., Sundukov, Yu.N. (2021): A new subspecies of Bembidion sanatum (Coleoptera: Carabidae) endemic to the Mendeleev Volcano (Kunashir Island, Russia). Far Eastern Entomologist 440: 13-24, DOI: 10.25221/fee.440.2, URL: http://dx.doi.org/10.25221/fee.440.2
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48

Maddison, David R. "A New Subgenus of Bembidion Latreille from México and Guatemala, with Descriptions of Two New Species (Coleoptera: Carabidae)." Proceedings of the California Academy of Sciences 67, no. 19 (2023): 433–50. https://doi.org/10.5281/zenodo.11067242.

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Maddison, David R. (2023): A New Subgenus of Bembidion Latreille from México and Guatemala, with Descriptions of Two New Species (Coleoptera: Carabidae). Proceedings of the California Academy of Sciences 67 (19): 433-450, DOI: 10.5281/zenodo.11067242
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49

Makarov, Kirill V., and Yurii N. Sundukov. "A revision of the subgenus Limnaeoperyphus Nakane, 1963 (Coleoptera: Carabidae, Bembidion), with the description of a new species." Zootaxa 5493, no. 4 (2024): 345–70. https://doi.org/10.11646/zootaxa.5493.4.3.

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Makarov, Kirill V., Sundukov, Yurii N. (2024): A revision of the subgenus Limnaeoperyphus Nakane, 1963 (Coleoptera: Carabidae, Bembidion), with the description of a new species. Zootaxa 5493 (4): 345-370, DOI: 10.11646/zootaxa.5493.4.3, URL: http://dx.doi.org/10.11646/zootaxa.5493.4.3
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50

Sasakawa, Kôji. "Taxonomic studies on the Bembidion ('Cillenus') complex (Coleoptera: Carabidae): A revision of the subgeneric taxonomy and description of a new species from Japan." Zootaxa 1575, no. 1 (2007): 35–45. https://doi.org/10.11646/zootaxa.1575.1.2.

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Sasakawa, Kôji (2007): Taxonomic studies on the Bembidion ('Cillenus') complex (Coleoptera: Carabidae): A revision of the subgeneric taxonomy and description of a new species from Japan. Zootaxa 1575 (1): 35-45, DOI: 10.11646/zootaxa.1575.1.2, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1575.1.2
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