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1

JONES, HUGH D., and WOLFGANG STERRER. "Terrestrial planarians (Platyhelminthes, with three new species) and nemertines of Bermuda." Zootaxa 1001, no. 1 (2005): 31. http://dx.doi.org/10.11646/zootaxa.1001.1.3.

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Records of terrestrial planarians and nemertines found on Bermuda are summarized. Three new species of terrestrial planarian are described though they certainly originated elsewhere. These are Amaga expatria n. sp., Gigantea gouvernoni n. sp. (both Geoplanidae and originating from South or Central America) and Bipalium vagum n. sp. (Bipaliidae, probably originating from Asia). Records of other planarians (Bipalium kewense, Anisorhynchodemus sp., Dolichoplana striata and Rhynchodemus sylvaticus) as well as nemertines (Pantinonemertes agricola and Geonemertes pelaensis) are discussed with regard to origin, distribution and mode of dispersal.
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2

Perez-Gelabert, Daniel. "Primera cita de la planaria terrestre cosmopolita Bipalium kewense Moseley, 1878 (Turbellaria: Tricladida: Terricola) para la República Dominicana." Novitates Caribaea, no. 3 (October 1, 2010): 81–82. http://dx.doi.org/10.33800/nc.v0i3.177.

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3

Trimen, Roland. "7. On Bipalium kewense at the Cape." Proceedings of the Zoological Society of London 55, no. 3 (2009): 548–50. http://dx.doi.org/10.1111/j.1096-3642.1887.tb03015.x.

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4

Juan Antonio, Pujol, and Ubero-Pascal Nicolás. "Non-indigenous terrestrial flatworms in Torrevieja (Alicante): presence of Bipalium cf. kewense Mosely, 1878 (Platyhelminthes, Tricladida, Geoplanidae)." Anales de Biología, no. 41 (October 3, 2019): 69–72. http://dx.doi.org/10.6018/analesbio.41.09.

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Se cita por primera vez en la provincia de Alicante (península ibérica) la especie de planaria terrestre exótica Bipalium cf. kewense Moseley, 1878. Con esta presencia aumenta el área de distribución de la especie en la Comunidad Valenciana, siendo la más meridional. Dado que la especie está considerada en esta comunidad autónoma como especie exótica invasora, se valora el riesgo potencial de colonización de ambientes naturales limítrofes a la zona de observación (jardín privado en zona urbanizada). The non-indigenous terrestrial flatworm Bipalium cf. kewense Moseley, 1878 is reported in Alicante province (Iberian Peninsula) for the first time. This occurrence is the most meridional from the Comunidad Valenciana, enlarging its distribution in this area. As the species is considered an invasive alien species in this region, the potential risk of colonizing natural environments closer to the observation area is evaluated.
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5

JUSTINE, JEAN-LOU, THOMAS THÉRY, DELPHINE GEY, and LEIGH WINSOR. "First record of the invasive land flatworm Bipalium adventitium (Platyhelminthes, Geoplanidae) in Canada." Zootaxa 4656, no. 3 (2019): 591–95. http://dx.doi.org/10.11646/zootaxa.4656.3.13.

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Specimens of Bipalium adventitium (Platyhelminthes, Geoplanidae) were found in Montréal, Québec, Canada. The specimens showed the typical colour pattern of the species and barcoding (Cytochrome Oxidase I) demonstrated near-identity with a sequence of the same species from the USA. This is the first record of the species in Canada.
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6

Stokes, Amber N., Peter K. Ducey, Lorin Neuman-Lee, et al. "Confirmation and Distribution of Tetrodotoxin for the First Time in Terrestrial Invertebrates: Two Terrestrial Flatworm Species (Bipalium adventitium and Bipalium kewense)." PLoS ONE 9, no. 6 (2014): e100718. http://dx.doi.org/10.1371/journal.pone.0100718.

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7

Selvamurugan, Selvaraj, and N. Santhana Bharathi. "New locality record of Bipalium univittatum Grube, 1866 (Tricladida: Geoplanidae: Bipaliinae) from India." International Journal of Agricultural and Applied Sciences 1, no. 2 (2020): 100–102. http://dx.doi.org/10.52804/ijaas2020.1217.

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Land planarians (Platyhelminthes) are likely important components of the soil cryptofauna, although relevant aspects of their ecology such as their density remain largely unstudied. The land planarian Bipalium univittatum Grube,1866 (Tricladida: Geoplanidae) is thought to be native from Southeast Asia. This species is found in Periyakallar, Valparai, Coimbatore district, Tamil Nadu state, India. This note gives details, distribution, description and conservation status
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8

Gastineau, Romain, Jean-Lou Justine, Claude Lemieux, Monique Turmel, and Andrzej Witkowski. "Complete mitogenome of the giant invasive hammerhead flatworm Bipalium kewense." Mitochondrial DNA Part B 4, no. 1 (2019): 1343–44. http://dx.doi.org/10.1080/23802359.2019.1596768.

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9

Ducey, Peter K., Jamie Cerqua, Lori-Jeanne West, and Monica Warner. "RARE EGG CAPSULE PRODUCTION IN THE INVASIVE TERRESTRIAL PLANARIAN BIPALIUM KEWENSE." Southwestern Naturalist 51, no. 2 (2006): 252–55. http://dx.doi.org/10.1894/0038-4909(2006)51[252:recpit]2.0.co;2.

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10

Ducey, Peter K., and Stacey Noce. "Successful Invasion of New York State by the Terrestrial Flatworm, Bipalium adventitium." Northeastern Naturalist 5, no. 3 (1998): 199. http://dx.doi.org/10.2307/3858619.

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11

Oki, Iwashiro, Sachiko Tamura, Robert E. Ogren, and Masaharu Kawakatsu. "Karyology of four land-planarian species of the genus Bipalium from Japan." Hydrobiologia 227, no. 1 (1991): 163–67. http://dx.doi.org/10.1007/bf00027597.

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12

SLUYS, RONALD, MATTHIAS NEUMANN, RICARDO F. DE LIMA, and ROBERT C. DREWES. "Land flatworms (Platyhelminthes, Geoplanidae) of São Tomé: a first account on their diversity, with the description of five new species." Zootaxa 4221, no. 3 (2017): 291. http://dx.doi.org/10.11646/zootaxa.4221.3.2.

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The present contribution provides the first faunistic and taxonomic account of six species of land flatworm from the island of São Tomé, including five new species of the genus Othelosoma Gray, 1869 and the introduced Bipalium kewense Moseley, 1878. One of the new species represents the first African land flatworm that has specks on its dorsal body surface, instead of stripes or a more or less uniform colouration. At least two of the new species were observed to prey on snails. The study details the fourth record of a sclerotic spermatophore in a species of land flatworm, and discusses the definition and homology of double female genital canals in African and Indian species of the genus Othelosoma.
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13

FIORE, C., J. TULL, S. ZEHNER, and P. DUCEY. "Tracking and predation on earthworms by the invasive terrestrial planarian Bipalium adventitium (Tricladida, Platyhelminthes)." Behavioural Processes 67, no. 3 (2004): 327–34. http://dx.doi.org/10.1016/s0376-6357(04)00138-x.

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14

Ducey, Peter K., Lori-Jeanne West, Gina Shaw, and Jacquelyn De Lisle. "Reproductive ecology and evolution in the invasive terrestrial planarian Bipalium adventitium across North America." Pedobiologia 49, no. 4 (2005): 367–77. http://dx.doi.org/10.1016/j.pedobi.2005.04.002.

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15

Baptista, Vanessa dos Anjos, Simone Machado de Oliveira, and Ana Maria Leal-Zanchet. "Inventário de planárias terrestres (Platyhelminthes, Tricladida) em remanescente de Floresta Estacional Decidual do Sul do Brasil." Biota Neotropica 10, no. 2 (2010): 247–52. http://dx.doi.org/10.1590/s1676-06032010000200027.

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Os tricladidos terrestres são predadores de topo de cadeia no seu microhábitat, alimentando-se de outros invertebrados de solo. A riqueza de espécies de tricladidos pode indicar a diversidade de outros grupos. A composição das comunidades de planárias terrestres da Floresta Estacional Decidual, que ocupa um quarto da cobertura vegetal do Estado do Rio Grande do Sul, é pouco conhecida. O presente trabalho teve como área de estudo o maior remanescente dessa formação florestal no Rio Grande do Sul, apresentando os seguintes objetivos: inventariar as espécies de planárias terrestres no Parque Estadual do Turvo (PE-Turvo) e comparar as comunidades desse Parque com as de outras áreas do estado com inventários desses tricladidos. Na área de estudo, foram inventariadas 26 espécies, sendo duas pertencentes às subfamílias Rhynchodeminae e Bipaliinae, e as demais, à subfamília Geoplaninae. Além dos gêneros Rhynchodemus Leidy 1851 e Bipalium Stimpson, 1857, foram registrados os seguintes gêneros de Geoplaninae: Geoplana Stimpson 1857, Choeradoplana Graff 1896, Pasipha Ogren & Kawakatsu 1990 e Notogynaphallia Ogren & Kawakatsu 1990, além do grupo coletivo Pseudogeoplana Ogren e Kawakatsu, 1990. O maior número de espécies foi registrado para o gênero Geoplana (10), seguido pelo gênero Pasipha (4). Geoplana rubidolineata Baptista & Leal-Zanchet, 2005 havia sido registrada anteriormente apenas para sua localidade-tipo, no nordeste do estado. Bipalium kewense Moseley, 1878, de distribuição cosmopolita, foi encontrada apenas em local submetido a alto impacto antrópico, não tendo sido registrada em áreas de floresta. Para comparar as comunidades de planárias terrestres do PE-Turvo com as de outros tipos de vegetação do estado, utilizou-se o método de ordenação NMDS. Essa análise indicou que as comunidades de planárias terrestres do PE-Turvo e de outras áreas de Floresta Decidual, Semidecidual e Floresta Ombrófila Densa diferenciam-se das comunidades de áreas de Floresta Ombrófila Mista. A alta riqueza de espécies observada no PE-Turvo reforça sua importante contribuição para a conservação da diversidade regional.
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16

ZABORSKI, EDMOND R. "Observations on Feeding Behavior by the Terrestrial Flatworm Bipalium adventitium (Platyhelminthes: Tricladida: Terricola) from Illinois." American Midland Naturalist 148, no. 2 (2002): 401. http://dx.doi.org/10.1674/0003-0031(2002)148[0401:oofbbt]2.0.co;2.

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17

Fernandes, M. C., E. P. Alvares, P. Gama, and M. Silveira. "Serotonin in the nervous system of the head region of the land planarian Bipalium kewense." Tissue and Cell 35, no. 6 (2003): 479–86. http://dx.doi.org/10.1016/s0040-8166(03)00074-0.

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18

Morffe, Jans, Nayla García, Byron Adams, and Koichi Hasegawa. "First record of the land planarian Bipalium kewense Moseley, 1878 (Tricladida: Geoplanidae: Bipaliinae) from Cuba." BioInvasions Records 5, no. 3 (2016): 127–32. http://dx.doi.org/10.3391/bir.2016.5.3.01.

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19

Ducey, Peter K., Michael Messere, Kellie Lapoint, and Stacey Noce. "Lumbricid Prey and Potential Herpetofaunal Predators of the Invading Terrestrial Flatworm Bipalium adventitium (Turbellaria: Tricladida: Terricola)." American Midland Naturalist 141, no. 2 (1999): 305–14. http://dx.doi.org/10.1674/0003-0031(1999)141[0305:lpaphp]2.0.co;2.

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20

BARAHONA-SEGOVIA, RODRIGO M., JUAN FRANCISCO ARAYA, and LAURA PAÑINAO-MONSÁLVEZ. "New records of the giant planarian Polycladus gayi Blanchard, 1845 (Platyhelminthes: Geoplanidae) with notes on its conservation biology." Zootaxa 4822, no. 4 (2020): 595–600. http://dx.doi.org/10.11646/zootaxa.4822.4.9.

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Land planarians (Tricladida:Geoplanidae) comprise about 910 species distributed in four subfamilies and can be found on all continents except Antarctica (Sluys & Riutort 2018; Sluys 2019). The Neotropical region possesses nearly 31% of all the described terrestrial planarian species, most of them belonging to the subfamily Geoplaninae (Sluys 1999; Grau & Carbayo 2010). Land planarians are mostly habitat-specialists, living in the humid soils of native forest, and predating on invertebrates like earthworms, isopods, mollusks and harvestmen, among others (Ogren 1995; Carbayo & Leal-Zanchet 2003; Boll & Leal-Zanchet 2016). Although most planarian species seem to be physiologically sensitive, for example to environmental moisture, a few land planarian genera like Bipalium Stimpson and Obama Carbayo et al., have successfully invaded many habitats, even in highly perturbed areas (Kawaguti 1932; Sluys 2019). Therefore, some of these invertebrate species appear to be good candidates as habitat quality bioindicators according to some authors (Sluys 1998; Gerlach et al., 2013; Negrete et al., 2014).
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21

Justine, Jean-Lou, Leigh Winsor, Delphine Gey, Pierre Gros, and Jessica Thévenot. "Giant worms chez moi! Hammerhead flatworms (Platyhelminthes, Geoplanidae, Bipalium spp., Diversibipalium spp.) in metropolitan France and overseas French territories." PeerJ 6 (May 22, 2018): e4672. http://dx.doi.org/10.7717/peerj.4672.

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Background Species of the genera Bipalium and Diversibipalium, or bipaliines, are giants among land planarians (family Geoplanidae), reaching length of 1 m; they are also easily distinguished from other land flatworms by the characteristic hammer shape of their head. Bipaliines, which have their origin in warm parts of Asia, are invasive species, now widespread worldwide. However, the scientific literature is very scarce about the widespread repartition of these species, and their invasion in European countries has not been studied. Methods In this paper, on the basis of a four year survey based on citizen science, which yielded observations from 1999 to 2017 and a total of 111 records, we provide information about the five species present in Metropolitan France and French overseas territories. We also investigated the molecular variability of cytochrome-oxidase 1 (COI) sequences of specimens. Results Three species are reported from Metropolitan France: Bipalium kewense, Diversibipalium multilineatum, and an unnamed Diversibipalium ‘black’ species. We also report the presence of B. kewense from overseas territories, such as French Polynesia (Oceania), French Guiana (South America), the Caribbean French islands of Martinique, Guadeloupe, Saint Martin and Saint Barthélemy, and Montserrat (Central America), and La Réunion island (off South-East Africa). For B. vagum, observations include French Guiana, Guadeloupe, Martinique, Saint Barthélemy, Saint Martin, Montserrat, La Réunion, and Florida (USA). A probable new species, Diversibipalium sp. ‘blue,’ is reported from Mayotte Island (off South–East Africa). B. kewense, B. vagum and D. multilineatum each showed 0% variability in their COI sequences, whatever their origin, suggesting that the specimens are clonal, and that sexual reproduction is probably absent. COI barcoding was efficient in identifying species, with differences over 10% between species; this suggests that barcoding can be used in the future for identifying these invasive species. In Metropolitan south–west France, a small area located in the Department of Pyrénées-Atlantiques was found to be a hot-spot of bipaliine biodiversity and abundance for more than 20 years, probably because of the local mild weather. Discussion The present findings strongly suggest that the species present in Metropolitan France and overseas territories should be considered invasive alien species. Our numerous records in the open in Metropolitan France raise questions: as scientists, we were amazed that these long and brightly coloured worms could escape the attention of scientists and authorities in a European developed country for such a long time; improved awareness about land planarians is certainly necessary.
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22

Ogren, Robert E. "Description of a New Three-Lined Land Planarian of the Genus Bipalium (Turbellaria: Tricladida) from Pennsylvania, U.S.A." Transactions of the American Microscopical Society 106, no. 1 (1987): 21. http://dx.doi.org/10.2307/3226281.

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23

Ducey, Peter K., Matthew McCormick, and Elizabeth Davidson. "Natural History Observations on Bipalium cf. vagum Jones and Sterrer (Platyhelminthes: Tricladida), a Terrestrial Broadhead Planarian New to North America." Southeastern Naturalist 6, no. 3 (2007): 449–60. http://dx.doi.org/10.1656/1528-7092(2007)6[449:nhoobc]2.0.co;2.

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24

Kamimura, K., and T. Yoshieda. "13.A case of nasal parasitism due to the land planaria, Bipalium fuscolineatum(Proceedings of the 62nd Annual Meeting of Western Region)." Medical Entomology and Zoology 59, no. 2 (2008): 118. http://dx.doi.org/10.7601/mez.59.118_3.

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25

Almeida, Ana Laura, Tiago Maurício Francoy, Marta Álvarez‐Presas, and Fernando Carbayo. "Convergent evolution: A new subfamily for bipaliin‐like Chilean land planarians (platyhelminthes)." Zoologica Scripta 50, no. 4 (2021): 500–508. http://dx.doi.org/10.1111/zsc.12479.

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26

"Pharyngeal regeneration in the land planarian Bipalium kewense." Hydrobiologia 227, no. 1 (1991): 57. http://dx.doi.org/10.1007/bf00027581.

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27

Getlekha, Nuntaporn. "Karyological Studies on Hammerhead Flatworm, Bipalium kewense (Tricladida, Terricola) from Thailand." Chiang Mai University Journal of Natural Sciences 19, no. 3 (2020). http://dx.doi.org/10.12982/cmujns.2020.0029.

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28

AA, Abdel-Haleem. "Histological and Scanning Electron Microscopic Studies on the Sensory Organs of the Terrestrial Flatworm Bipalium kewense." Journal of Cytology & Histology 05, no. 03 (2014). http://dx.doi.org/10.4172/2157-7099.1000233.

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29

Agnolin, Federico, Agustín Agnolin, and Elián Guerrero. "Invertebrados exóticos nuevos o poco conocidos en la ciudad de Buenos Aires." Acta Zoológica Lilloana, December 2, 2019, 48–67. http://dx.doi.org/10.30550/j.azl/2019.63.2/4.

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Las especies invasoras pueden causar diversos problemas a nivel socioeconómico y para la conservación de la biodiversidad. La detección temprana de las especies naturalizadas puede ser de vital importancia para el monitoreo y la planificación de estrategias de contención del proceso de invasión. La fauna de planarias terrestres (Tricladida, Geoplanoidea), babosas terrestres (Mollusca, Limacidae), diplópodos (Myriapoda, Diplopoda) y sínfilos (Myriapoda, Symphylla) de la Ciudad Autónoma de Buenos Aires se encuentra poco estudiada, y la adición de especies exóticas transportadas por las actividades humanas complejiza la situación de poco conocimiento. En este trabajo se dan a conocer nuevos registros de especies de estos grupos que se han naturalizado en el ámbito urbano de la ciudad y sus alrededores (Bipalium kewense, Austroplana sanguinea, Ambigolimax valentianus, Brachyiulus pusillus, Oxidus gracilis y Scutigerella immaculata). Se evaluó el potencial como invasor de cada especie en base a lo registrado en otros países. Hasta el día de la fecha no existen registros publicados de planarias terrestres, sínfilos, o diplópodos para esta área, por lo cual el presente trabajo constituye la primera contribución sobre estos grupos. Las especies mencionadas perjudican a las actividades agropecuarias y horticulturales de diversas formas. Una tarea a futuro deberá ser la cuantificación de los daños que causan para establecer su importancia y el costo/beneficio de su control o manejo.
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