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1

Lokeshwor, Y., and W. Vishwanath. "Two new species of stone loaches of the genus Schistura from the Koladyne basin, Mizoram, India (Teleostei: Cobitoidei: Nemacheilidae)." Zootaxa 3731, no. 4 (2013): 561–76. https://doi.org/10.11646/zootaxa.3731.4.8.

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Lokeshwor, Y., Vishwanath, W. (2013): Two new species of stone loaches of the genus Schistura from the Koladyne basin, Mizoram, India (Teleostei: Cobitoidei: Nemacheilidae). Zootaxa 3731 (4): 561-576, DOI: 10.11646/zootaxa.3731.4.8
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2

Прокофьев, А. М. "Рецензия на книгу М. Коттела “Conspectus cobitidum: инвентаризация вьюнов Мира (Teleostei: Cypriniformes: Cobitoidei)”". Вопросы ихтиологии 54, № 2 (2014): 243–48. http://dx.doi.org/10.7868/s0042875214020118.

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3

Liu, Huanzhang, Chyng-Shyan Tzeng, and Hui-Yu Teng. "Sequence variations in the mitochondrial DNA control region and their implications for the phylogeny of the Cypriniformes." Canadian Journal of Zoology 80, no. 3 (2002): 569–81. http://dx.doi.org/10.1139/z02-035.

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The mitochondrial DNA control region of six cobitids and two catostomids was sequenced and compared with sequences of other cypriniforms to study their sequence variations. The extended termination associated sequence (ETAS) domain, central domain, and conserved sequence block (CSB) domain were partitioned and the ETAS sequence, CSB-D, CSB-E, ECSB-F, CSB1, CSB2, and CSB3 were identified. It is suggested that the "hairpin" TACAT-ATGTA is the key sequence of ETAS and GACATA is the symbol of CSB1. Phylogenetic analysis based on the CSB domain showed that all cyprinids evolved as one monophyletic
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4

Bogdanov, B. E. "Variability and systematic status of Siberian spined loach Cobitis melanoleuca (Cypriniformes; Cobitoidei) in the Baikal region." Limnology and Freshwater Biology, no. 1 (2025): 68–83. https://doi.org/10.31951/2658-3518-2025-a-1-68.

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Siberian spined loach Cobitis melanoleuca (sensu lato) is distributed in the Northern Eurasia, and the Baikal region is the center of an areal of this species (or group of species). The present study analyzed the variability of exterior traits in eight populations of Siberian spined loach in the Baikal region (original data) and in the whole range (literature data). The results of the study have shown a high polymorphism of traits traditionally considered species-specific. It was found that populations of spined loaches inhabiting the Cisbaikalia are a mixture of phenotypes typical of Cobitis
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5

Dyldin, Yu V., A. M. Orlov, L. Hanel, V. I. Romanov, R. Fricke, and E. D. Vasil’eva. "Ichthyofauna of the Fresh and Brackish Waters of Russia and Adjacent Areas: Annotated List with Taxonomic Comments. 2. Order Cypriniformes, Suborders Catostomoidei, Cobitoidei and Cyprinoidei." Вопросы ихтиологии 63, no. 4 (2023): 386. http://dx.doi.org/10.31857/s0042875223040069.

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This paper is the second part of the “Ichthyofauna of the Fresh and Brackish Waters of Russia and Adjacent Areas: Annotated List with Taxonomic Comments”, which we started in 2022 based on a critical analysis of scientific publications of the last 200 years, and on the data of the author’s research. The current paper includes all currently known fishes of the order Cypriniformes of Russia and adjacent waters, represented by three suborders Catostomoidei, Cobitoidei and Cyprinoidei, comprising ten families, 64 genera and 177 species. The most specious families were the Leuciscidae (59 species/3
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6

LOKESHWOR, Y., and W. VISHWANATH. "Two new species of stone loaches of the genus Schistura from the Koladyne basin, Mizoram, India (Teleostei: Cobitoidei: Nemacheilidae)." Zootaxa 3731, no. 4 (2013): 561. http://dx.doi.org/10.11646/zootaxa.3731.4.8.

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7

Bohlen, Jörg, Vendula Šlechtová, and Kamphol Udomritthiruj. "Schistura hypsiura, a new species of loach (Cobitoidea: Nemacheilidae) from South-West Myanmar." Raffles Bulletin of Zoology 62 (February 21, 2014): 21–27. https://doi.org/10.5281/zenodo.4504124.

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Bohlen, Jörg, Šlechtová, Vendula, Udomritthiruj, Kamphol (2014): Schistura hypsiura, a new species of loach (Cobitoidea: Nemacheilidae) from South-West Myanmar. Raffles Bulletin of Zoology 62: 21-27, DOI: 10.5281/zenodo.4504124
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8

Tomáš, Scholz, de Chambrier Alain, Shimazu Takeshi, Ermolenko Alexey, and Waeschenbach Andrea. "Proteocephalid tapeworms (Cestoda: Onchoproteocephalidea) of loaches (Cobitoidea): Evidence for monophyly and high endemism of parasites in the Far East." Parasitology International 66 (October 1, 2016): 871–83. https://doi.org/10.1016/j.parint.2016.09.016.

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Tomáš Scholz, Alain de Chambrier, Takeshi Shimazu, Alexey Ermolenko, Andrea Waeschenbach (2017): Proteocephalid tapeworms (Cestoda: Onchoproteocephalidea) of loaches (Cobitoidea): Evidence for monophyly and high endemism of parasites in the Far East. Parasitology International 66: 871-883, DOI: 10.1016/j.parint.2016.09.016
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9

Tomáš, Scholz, de Chambrier Alain, Shimazu Takeshi, Ermolenko Alexey, and Waeschenbach Andrea. "Proteocephalid tapeworms (Cestoda: Onchoproteocephalidea) of loaches (Cobitoidea): Evidence for monophyly and high endemism of parasites in the Far East." Parasitology International 66 (December 31, 2017): 871–83. https://doi.org/10.1016/j.parint.2016.09.016 1383-5769.

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Tomáš Scholz, Alain de Chambrier, Takeshi Shimazu, Alexey Ermolenko, Andrea Waeschenbach (2017): Proteocephalid tapeworms (Cestoda: Onchoproteocephalidea) of loaches (Cobitoidea): Evidence for monophyly and high endemism of parasites in the Far East. Parasitology International 66: 871-883, DOI: http://dx.doi.org/10.1016/j.parint.2016.09.0161383-5769
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10

Nakajima, Jun. "Cobitis takenoi sp. n. (Cypriniformes, Cobitidae): a new spined loach from Honshu Island, Japan." ZooKeys 568 (February 23, 2016): 119–28. https://doi.org/10.3897/zookeys.568.7733.

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A new species of spined loach, Cobitis takenoi sp. n., is described based on the holotype and ten paratypes collected from Tango District, Honshu Island, Japan. The new species is distinguished by a combination of the following character states: 1) the lamina circularis at the base of the pectoral fin in adult male having a simple roundish plate form; 2) a narrowing of the upper segments of the first branched ray of the pectoral fin; 3) a short maxillary barbel whose length equals diameter of the eye; 4) 14 prepelvic myotomes, and 5) L3 and L5 well developed, forming longitudinal obvious strip
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11

Sato, Tatsuro, Jun Nakajima, Liangliang Huang, et al. "Distribution Pattern of Loaches (Teleostei: Cobitoidea) in the River East Tiaoxi, China." Folia Zoologica 60, no. 4 (2011): 328–34. http://dx.doi.org/10.25225/fozo.v60.i4.a10.2011.

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12

Sayyadzadeh, G., H. R. Esmaeili, and K. Abbasi. "Comparative Taxonomically Important Osteological Characters of Loach Fishes from Iran (Cypriniformes: Cobitoidea)." Iranian Journal of Science and Technology, Transactions A: Science 43, no. 5 (2019): 2149–62. http://dx.doi.org/10.1007/s40995-019-00735-z.

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13

Nalbant, Teodor T., and Pier Giorgio Bianco. "The loaches of Iran and adjacent regions with description of six new species (Cobitoidea)." Italian Journal of Zoology 65, sup1 (1998): 109–23. http://dx.doi.org/10.1080/11250009809386803.

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14

Min, Rui, Yahui Zhao, Jingsong Shi, and Junxing Yang. "A new species of Homatula (Teleostei, Cobitoidea, Nemacheilidae) from the Pearl River drainage, Yunnan, China." ZooKeys 1089 (March 17, 2022): 109–24. http://dx.doi.org/10.3897/zookeys.1089.77203.

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Based on morphological and molecular analysis of Homatula species distributed in the Nanpanjiang River in Yunnan, China, we described a new species, Homatula robustasp. nov. It differs from its congeners by a combination of the following characters: naked and robust body with well-developed crests (caudal peduncle depth as a percentage of its length: 70.5–78.5%); lateral line complete; median notch on lower jaw; median gap on lower lip; three pairs of short barbels, with maxillary barbels extending posteriorly to anterior edge of eyes; branched dorsal-fin rays 8½; and vertebrae 37–39. It can f
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15

Lei, Dujuan, Unisa Conteh Kanu, Guang Zhao, et al. "The complete mtDNA genome of Triplophysa dorsalis (Cypriniformes, Balitoridae, Cobitoidea): genome characterization and phylogenetic analysis." Mitochondrial DNA Part A 27, no. 5 (2015): 3745–46. http://dx.doi.org/10.3109/19401736.2015.1079886.

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16

Kanu, Unisa Conteh, Guang Zhao, Peng Xie, et al. "The complete mtDNA genome of Triplophysa strauchii (Cypriniformes, Balitoridae, Cobitoidea): genome charaterization and phylogenetic analysis." Mitochondrial DNA Part A 27, no. 4 (2015): 2637–38. http://dx.doi.org/10.3109/19401736.2015.1041131.

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17

Feng, Chenguang, Weiwei Zhou, Yongtao Tang, et al. "Molecular systematics of the Triplophysa robusta (Cobitoidea) complex: Extensive gene flow in a depauperate lineage." Molecular Phylogenetics and Evolution 132 (March 2019): 275–83. http://dx.doi.org/10.1016/j.ympev.2018.12.009.

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18

Min, Rui, Yahui Zhao, Jingsong Shi, and Junxing Yang. "A new species of Homatula (Teleostei, Cobitoidea, Nemacheilidae) from the Pearl River drainage, Yunnan, China." ZooKeys 1089 (March 17, 2022): 109–24. https://doi.org/10.3897/zookeys.1089.77203.

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Based on morphological and molecular analysis of Homatula species distributed in the Nanpanjiang River in Yunnan, China, we described a new species, Homatula robusta sp. nov. It differs from its congeners by a combination of the following characters: naked and robust body with well-developed crests (caudal peduncle depth as a percentage of its length: 70.5–78.5%); lateral line complete; median notch on lower jaw; median gap on lower lip; three pairs of short barbels, with maxillary barbels extending posteriorly to anterior edge of eyes; branched dorsal-fin rays 8½; and vertebrae 37–39. It can
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19

Chen, W. J., V. Lheknim, and R. L. Mayden. "Molecular phylogeny of the Cobitoidea (Teleostei: Cypriniformes) revisited: position of enigmatic loachEllopostomaresolved with six nuclear genes." Journal of Fish Biology 75, no. 9 (2009): 2197–208. http://dx.doi.org/10.1111/j.1095-8649.2009.02398.x.

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20

Coad, Brian W., and A. G. K. Menon. "The Fauna of India and the Adjacent Countries. Pisces. Vol. IV, Teleostei-Cobitoidea. Part 1, Homalopteridae." Copeia 1989, no. 4 (1989): 1101. http://dx.doi.org/10.2307/1446016.

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21

Sember, Alexandr, Jörg Bohlen, Vendula Šlechtová, Marie Altmanová, Šárka Pelikánová, and Petr Ráb. "Dynamics of tandemly repeated DNA sequences during evolution of diploid and tetraploid botiid loaches (Teleostei: Cobitoidea: Botiidae)." PLOS ONE 13, no. 3 (2018): e0195054. http://dx.doi.org/10.1371/journal.pone.0195054.

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22

Keskar, Ashwini, Rajeev Raghavan, Mandar S. Paingankar, et al. "Molecular phylogeny unveils hidden diversity of hillstream loaches (Cypriniformes: Cobitoidea) in the northern Western Ghats of India." Meta Gene 17 (September 2018): 237–48. http://dx.doi.org/10.1016/j.mgene.2018.07.002.

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23

Liu, Si-qing, Richard L. Mayden, Jia-bo Zhang, et al. "Phylogenetic relationships of the Cobitoidea (Teleostei: Cypriniformes) inferred from mitochondrial and nuclear genes with analyses of gene evolution." Gene 508, no. 1 (2012): 60–72. http://dx.doi.org/10.1016/j.gene.2012.07.040.

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24

He, Y., and D. K. He. "X-ray Microtomography Study of Cranium, Pectoral Fin and Girdle, and Weberian Apparatus Development in Juvenile Triplophysa stewarti (Nemacheilidae)." Вопросы ихтиологии 63, no. 2 (2023): 128. http://dx.doi.org/10.31857/s0042875223020091.

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The stone loach genus Triplophysa is a major component of the Tibetan Plateau ichthyofauna, and its taxonomy and phylogenetic relationships are far from resolved. Skeletogeny patterns contain useful phylogenetic signals, but developmental data for Triplophysa are scarce. Here, we present an X-ray microtomography study of the development of the cranium, pectoral fin and girdle, and Weberian apparatus of juvenile Triplophysa stewarti (12–55 mm standard length). We reveal that the smallest juveniles possess a fontanelle and well-developed Weberian ossicles. Furthermore, the development of the cra
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25

Scholz, Tomáš, Alain de Chambrier, Takeshi Shimazu, Alexey Ermolenko, and Andrea Waeschenbach. "Proteocephalid tapeworms (Cestoda: Onchoproteocephalidea) of loaches (Cobitoidea): Evidence for monophyly and high endemism of parasites in the Far East." Parasitology International 66, no. 1 (2017): 871–83. http://dx.doi.org/10.1016/j.parint.2016.09.016.

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26

Šlechtová, Vendula, Zuzana Musilova, Heok Hui Tan, Maurice Kottelat, and Jörg Bohlen. "One northward, one southward: Contrasting biogeographical history in two benthic freshwater fish genera across Southeast Asia (Teleostei: Cobitoidea: Nemacheilus, Pangio)." Molecular Phylogenetics and Evolution 161 (August 2021): 107139. http://dx.doi.org/10.1016/j.ympev.2021.107139.

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27

Shedko, S. V., I. L. Miroshnichenko, and G. A. Nemkova. "On the systematics and phylogeography of eight-barbel loaches of the genus Lefua (Cobitoidea: Nemacheilidae): mtDNA typing of L. pleskei." Russian Journal of Genetics 44, no. 7 (2008): 817–25. http://dx.doi.org/10.1134/s1022795408070090.

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28

Šlechtová, Vendula, Jörg Bohlen, Jörg Freyhof, and Petr Ráb. "Molecular phylogeny of the Southeast Asian freshwater fish family Botiidae (Teleostei: Cobitoidea) and the origin of polyploidy in their evolution." Molecular Phylogenetics and Evolution 39, no. 2 (2006): 529–41. http://dx.doi.org/10.1016/j.ympev.2005.09.018.

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29

Šlechtová, Vendula, Jörg Bohlen, and Heok Hui Tan. "Families of Cobitoidea (Teleostei; Cypriniformes) as revealed from nuclear genetic data and the position of the mysterious genera Barbucca, Psilorhynchus, Serpenticobitis and Vaillantella." Molecular Phylogenetics and Evolution 44, no. 3 (2007): 1358–65. http://dx.doi.org/10.1016/j.ympev.2007.02.019.

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30

Tang, Qiongying, Huanzhang Liu, Richard Mayden, and Bangxi Xiong. "Comparison of evolutionary rates in the mitochondrial DNA cytochrome b gene and control region and their implications for phylogeny of the Cobitoidea (Teleostei: Cypriniformes)." Molecular Phylogenetics and Evolution 39, no. 2 (2006): 347–57. http://dx.doi.org/10.1016/j.ympev.2005.08.007.

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31

Kottelat, Maurice. "Nomenclatural status of the fish names created by J. C. van Hasselt (1823) and of some cobitoid genera." Japanese Journal of Ichthyology 33, no. 4 (1987): 368–75. http://dx.doi.org/10.1007/bf02904097.

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32

Prokofiev, A. M. "Materials towards the revision of the genus Triplophysa rendahl, 1933 (Cobitoidea: Balitoridae: Nemacheilinae): A revision of nominal taxa of Herzenstein (1888) described within the species “Nemachilus” stoliczkae and “N.” dorsonotatus, with the description of the new species T. scapanognatha sp. nova." Journal of Ichthyology 47, no. 1 (2007): 1–20. http://dx.doi.org/10.1134/s0032945207010018.

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33

Hnátková, Eva, Zuzana Majtánová, Šlechtová Vendula Bohlen, Joerg Bohlen, and Petr Ráb. "Karyotype of Sabanejewia bulgarica (Drensky, 1928) (Teleostei, Cobitidae) from the Danube Delta, Romania." Comparative Cytogenetics 17, no. () (2023): 157–62. https://doi.org/10.3897/compcytogen.17.103152.

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The karyotype of the freshwater fish Sabanejewia bulgarica (Drensky, 1928), from the Danube Delta, was studied by conventional Giemsa staining and the C-banding technique. The diploid chromosome number was 2n = 50. The karyotype contained 2 pairs of metacentric (the first one was much larger than the second one), 6 pairs of submetacentric and 17 pairs of subtelocentric to acrocentric chromosomes. Pericentromeric blocks of heterochromatin were revealed in most of the chromosome pairs. The karyotype phenotype of S. bulgarica was the same as found for S. balcanica from Northern Carpathian Mountai
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34

Goswami, Mukunda, A. Pavan-Kumar, Grishma S. Patil, et al. "Molecular identification of ornamental loaches (Cypriniformes, Cobitoidei) of North East India using mitochondrial genes." Animal Gene, August 2022, 200136. http://dx.doi.org/10.1016/j.angen.2022.200136.

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35

Dyldin, Yu V., A. M. Orlov, L. Hanel, V. I. Romanov, R. Fricke, and E. D. Vasil’eva. "Ichthyofauna of the Fresh and Brackish Waters of Russia and Adjacent Areas: Annotated List with Taxonomic Comments. 2. Order Cypriniformes, Suborders Catostomoidei, Cobitoidei and Cyprinoidei." Journal of Ichthyology, June 19, 2023. http://dx.doi.org/10.1134/s0032945223040045.

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36

Jawad, Laith A., Golnaz Sayyadzadeh, and Hamid Reza Esmaeili. "Identification of loach fishes from Iran (Cypriniformes: Cobitoidea) using some osteological characters." Acta Zoologica, March 11, 2025. https://doi.org/10.1111/azo.12541.

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AbstractThis study examines eight skeletal features to distinguish species within the Cobitidae and Nemacheilidae families, focusing on the vertebral column, fin structure and caudal fin rays. The study analyses the construction and regionalization of the vertebral column, the interdigitation of pterygiophores with neural and haemal spines in the dorsal and anal fins, and the distribution of procurrent rays in the caudal fin. These characteristics are used to identify taxonomically relevant traits and separate species within these two families. The study also proposes a new method for defining
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37

Feng, Chenguang, Yongtao Tang, Sijia Liu, Fei Tian, Cunfang Zhang, and Kai Zhao. "Multiple convergent events created a nominal widespread species: Triplophysa stoliczkae (Steindachner, 1866) (Cobitoidea: Nemacheilidae)." BMC Evolutionary Biology 19, no. 1 (2019). http://dx.doi.org/10.1186/s12862-019-1503-3.

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38

Ráb, Petr, Eva Hnátková, Zuzana Majtánová, Vendula Bohlen Šlechtová, and Joerg Bohlen. "Karyotype Record for the Morphologically Derived, Rarely Collected, Freshwater Fish Ellopostoma mystax (Cypriniformes, Cobitoidea, Ellopostomatidae)." Ichthyology & Herpetology 109, no. 4 (2021). http://dx.doi.org/10.1643/i2020032.

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