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Journal articles on the topic 'Copris'

Author: Grafiati

Published: 5 June 2025

Last updated: 24 June 2025

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1

Darling, James D. G., and François Génier. "Revision of the taxonomy and distribution of the Neotropical Copris incertus species complex (Coleoptera: Scarabaeidae: Scarabaeinae)." Canadian Entomologist 150, no. 5 (2018): 539–77. http://dx.doi.org/10.4039/tce.2018.32.

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AbstractCopris incertus Say, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae: Coprini) has been described as a New World coprophagous scarab distributed from Mexico to Ecuador with large discontinuities in its range between the Yucatán province and Costa Rica. The C. incertus species complex of the Copris minutus (Drury, 1773) species group consists of C. incertus, Copris laeviceps Harold, 1869, and Copris lugubris Boheman, 1858. Based on external morphology and male genitalia, we discovered that multiple species have been classified as C. incertus. Of these species, five are new: Copris amazonicusnew species, Copris brevicornisnew species, Copris davidinew species, Copris moroninew species, and Copris susanaenew species. Herein, we revise the organisation of the C. incertus species complex and propose a new species complex, the C. laeviceps species complex, which includes: C. davidi, Copris igualensis Warner, 1990, and C. laeviceps, formerly included in the C. incertus species complex. We provide an identification key along with species distribution maps, images of habitus, and diagnostic characters.

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2

KOHLMANN, BERT, ENIO CANO, and LEONARDO DELGADO. "New species and records of Copris (Coleoptera: Scarabaeidae; Scarabaeinae) from Central America." Zootaxa 167, no. 1 (2003): 1. http://dx.doi.org/10.11646/zootaxa.167.1.1.

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Two new species of Copris Geoffroy from Guatemala and Honduras are described and illustrated: Copris caliginosus sp. nov. and Copris nubilosus sp. nov. Both species are related to C. sallei Harold, which is redescribed and illustrated for the first time. A key for the remotus complex is also included. New distributional records of Copris are provided for five species and subspecies from Guatemala and El Salvador.

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3

BUI, VAN BAC, THOMAS ZIEGLER, and MICHAEL BONKOWSKI. "Checklist of beetles in the subgenus Copris (Paracopris) Balthasar from Asia with description of a new species, and redescription of Copris (Paracopris) punctulatus Wiedemann (Coleoptera: Scarabaeidae: Scarabaeinae)." Zootaxa 4712, no. 1 (2019): 51–64. http://dx.doi.org/10.11646/zootaxa.4712.1.3.

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We present the first checklist of Asian beetles of the subgenus Copris (Paracopris) Balthasar, 1939 including new data on the distribution. In addition, Copris (Paracopris) javanensis Bui, Ziegler & Bonkowski, new species from Java (Indonesia) is described and Copris (Paracopris) punctulatus Wiedemann, 1823 is redescribed based on specimens deposited in the Muséum National d’Histoire Naturelle (Paris, France), including detailed illustrations of both sexes and male genitalia. At the same time, a neotype is designated for C. (P.) punctulatus as the type specimen was lost.

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4

DESCHODT, CHRISTIAN M., ADRIAN L. V. DAVIS, and CLARKE H. SCHOLTZ. "A new synonymy in the fidius group of Copris Müller 1764 (Coleoptera: Scarabaeidae: Scarabaeinae) and a new species from the highland grasslands of South Africa." Zootaxa 3949, no. 3 (2015): 431. http://dx.doi.org/10.11646/zootaxa.3949.3.8.

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The fidius group constitutes the basal clade in a phylogeny of the Afrotropical members of the genus, Copris Müller, 1764. In this paper we describe Copris crassus Deschodt and Davis as a new species in this group. We also demonstrate that the distributional function between two body dimensions (lateral horn length and mid-line elytron length) differs between Copris fidius (Olivier 1789) and C. crassus. Copris bihamatus Balthasar, 1965 is also a member of the fidius group. Nguyen-Phung (1988) suspected that it was a synonym of C. fidius (Olivier) so we compare specimens of similar body size and use the distributional function between body dimensions to demonstrate that C. bihamatus is indeed a synonym of C. fidius. We provide a key for all known valid species in the fidius group, except C. serius Nguyen-Phung, 1987.

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5

Bui, Van Bac, Thomas Ziegler, and Michael Bonkowski. "Checklist of beetles in the subgenus Copris (Paracopris) Balthasar from Asia with description of a new species, and redescription of Copris (Paracopris) punctulatus Wiedemann (Coleoptera: Scarabaeidae: Scarabaeinae)." Zootaxa 4712, no. 1 (2019): 51–64. https://doi.org/10.11646/zootaxa.4712.1.3.

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Bui, Van Bac, Ziegler, Thomas, Bonkowski, Michael (2019): Checklist of beetles in the subgenus Copris (Paracopris) Balthasar from Asia with description of a new species, and redescription of Copris (Paracopris) punctulatus Wiedemann (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 4712 (1): 51-64, DOI: 10.11646/zootaxa.4712.1.3

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6

López-Guerrero, Irma, Mario Zunino, and Gonzalo Halffter. "Taxonomic Use of Genitalic Characters in Mexican Copris (Coleoptera: Scarabaeidae, Scarabaeinae): The Case of Copris klugi sierrensis Matthews and the C. armatus Species Complex." Coleopterists Bulletin 63, no. 2 (2009): 203–12. https://doi.org/10.1649/1145.1.

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López-Guerrero, Irma, Zunino, Mario, Halffter, Gonzalo (2009): Taxonomic Use of Genitalic Characters in Mexican Copris (Coleoptera: Scarabaeidae, Scarabaeinae): The Case of Copris klugi sierrensis Matthews and the C. armatus Species Complex. The Coleopterists Bulletin 63 (2): 203-212, DOI: 10.1649/1145.1, URL: http://dx.doi.org/10.1649/1145.1

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7

MCCLEVE, SCOTT, and BERT KOHLMANN. "New species and new records of Copris (Coleoptera: Scarabaeidae: Scarabaeinae) from Mexico and the United States." Zootaxa 1096, no. 1 (2005): 17. http://dx.doi.org/10.11646/zootaxa.1096.1.2.

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One new species of Copris Geoffroy from Sonora and Chihuahua is described and illustrated: Copris warneri sp. nov. The new species is closely related to C. arizonensis Schaeffer. Drawings of the male and female and a distribution map are included. New distributional records for C. arizonensis Schaeffer and C. martinezi Matthews and Halffter are provided.

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8

Veenakumari, K., and G. K. Veeresh. "Subsociality in dung beetles Copris repertus Walker and Copris indicus Gill. (Coleoptera: Scarabaeidae)." Journal of the Bombay Natural History Society 94 (June 6, 1997): 530–35. https://doi.org/10.5281/zenodo.13656052.

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9

MALDANER, MARIA E., MARIO CUPELLO, DANIELA C. FERREIRA, and FERNANDO Z. VAZ-DE-MELLO. "Type specimens and names assigned to Coprophanaeus (Megaphanaeus) d’Olsoufieff, 1924, the largest New World dung beetles (Coleoptera: Scarabaeidae: Phanaeini)." Zootaxa 4272, no. 1 (2017): 83. http://dx.doi.org/10.11646/zootaxa.4272.1.4.

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Currently 13 species-group names assigned to the subgenus Megaphanaeus of Coprophanaeus are deemed available (or 'potentially available'), four of which denote valid species. In the present work we review the nomenclatural history of those names and conclude that two of them, Scarabaeus satelles Lichtenstein, 1796 and Copris ajax Sturm, 1826, are unavailable. For the other 11 names, we found type specimens of five, Scarabaeus bellicosus Olivier, 1789, Copris ensifer Germar, 1821, Phanaeus septentrionalis Pêssoa, 1934, P. vicinus Martínez, 1944, and P. vicinus var. argentinus Martínez, 1944. Two lectotypes are designated, one for Copris ensifer and other for Phanaeus septentrionalis. We were unable to locate the type series of the four names described by Castelnau (1840), i.e. Phanaeus ducalis, P. sylvanus, P. heros and P. miles, and of P. bonariensis Gory, 1844 and Scarabaeus lancifer Linné, 1767. For the latter, based on some iconotypes, we found that its type series was composite, including specimens of two different Megaphanaeus species, and, in addition, it was mixed with Copris ensifer type series. In order to fix the name to a sole species, a neotype is designated for Scarabaeus lancifer. The history of the names are presented in detail, and the application of these names to species is briefly re-discussed.

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10

HERNANDEZ, Blas, Jean Michel MAES, Gretel VARGAS, Natya TORUÑO, and Luis ZEAS. "Copris maesi RATCLIFFE (Scarabaeidae) en Selva Negra, Matagalpa, Nicaragua." Revista Nicaraguense de Entomologia 273 (June 30, 2022): 1–15. https://doi.org/10.5281/zenodo.7314578.

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11

Kohlmann, Bert, and Enio Cano And Leonardo Delgado. "New species and records of Copris (Coleoptera: Scarabaeidae; Scarabaeinae) from Central America." Zootaxa 167 (December 31, 2003): 1–16. https://doi.org/10.5281/zenodo.156784.

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12

Mccleve, Scott, and Bert Kohlmann. "New species and new records of Copris (Coleoptera: Scarabaeidae: Scarabaeinae) from Mexico and the United States." Zootaxa 1096 (December 31, 2005): 17–28. https://doi.org/10.5281/zenodo.170530.

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Mccleve, Scott, Kohlmann, Bert (2005): New species and new records of Copris (Coleoptera: Scarabaeidae: Scarabaeinae) from Mexico and the United States. Zootaxa 1096: 17-28, DOI: 10.5281/zenodo.170530

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13

Hill, Robert A., and Andrew Sutherland. "Hot off the press." Natural Product Reports 32, no. 6 (2015): 760–64. http://dx.doi.org/10.1039/c5np90021b.

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A personal selection of 32 recent papers is presented covering various aspects of current developments in bioorganic chemistry and novel natural products such as coprisamide A from a bacterium isolated from Copris tripartitus.

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14

Pizzo, A., V. Citeroni, F. Mazzone, M. Dellacasa, and C. Palestrini. "Are Horn Morphological Patterns Able to Differentiate the Two Closely Related Species Copris klugi Harold and Copris sierrensis Matthews?" Neotropical Entomology 44, no. 2 (2015): 153–59. http://dx.doi.org/10.1007/s13744-015-0277-6.

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15

Ziani, Stefano. "Morphological revision of the western Palaearctic species of the genus Copris Geoffroy, 1762 with three foretibial external teeth (Coleoptera: Scarabaeoidea: Scarabaeidae)." Insecta Mundi 2017, no. 528 (2017): 1–26. https://doi.org/10.5281/zenodo.4645819.

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Ziani, Stefano (2017): Morphological revision of the western Palaearctic species of the genus Copris Geoffroy, 1762 with three foretibial external teeth (Coleoptera: Scarabaeoidea: Scarabaeidae). Insecta Mundi 2017 (528): 1-26, DOI: 10.5281/zenodo.4645819

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16

Ziani, Stefano. "Subsequent spelling cavolinii for Scarabaeus cavolini Petagna, 1792 currently Copris hispanus cavolinii) to be maintained as prevailing usage (Coleoptera: Scarabaeoidea: Scarabaeidae: Coprini)." Insecta Mundi 2020, no. 777 (2020): 1–5. https://doi.org/10.5281/zenodo.5755752.

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Ziani, Stefano (2020): Subsequent spelling cavolinii for Scarabaeus cavolini Petagna, 1792 currently Copris hispanus cavolinii) to be maintained as prevailing usage (Coleoptera: Scarabaeoidea: Scarabaeidae: Coprini). Insecta Mundi 2020 (777): 1-5, DOI: 10.5281/zenodo.5755752

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17

Huerta, Carmen, and Hea Son Bang. "Fecundity And Offspring Survival Of Copris Tripartitus Waterhouse (Coleoptera, Scarabaeidae: Scarabaeinae) Under Laboratory Rearing Conditions." Coleopterists Bulletin 58, no. 4 (2004): 501–7. https://doi.org/10.1649/638.

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Huerta, Carmen, Bang, Hea Son (2004): Fecundity And Offspring Survival Of Copris Tripartitus Waterhouse (Coleoptera, Scarabaeidae: Scarabaeinae) Under Laboratory Rearing Conditions. The Coleopterists Bulletin 58 (4): 501-507, DOI: 10.1649/638, URL: http://dx.doi.org/10.1649/638

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18

Vaz-de-Mello, José Mondaca Fernando Z. "Homocopris Burmeister, 1846 (Coleoptera: Scarabaeidae): lectotype designation for Copris torulosus Eschscholtz, 1822, and correction on the geographic distribution of the two Chilean species." Insecta Mundi 2024, no. 68 (2024): 1–5. https://doi.org/10.5281/zenodo.14662048.

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Vaz-de-Mello, José Mondaca Fernando Z. (2024): Homocopris Burmeister, 1846 (Coleoptera: Scarabaeidae): lectotype designation for Copris torulosus Eschscholtz, 1822, and correction on the geographic distribution of the two Chilean species. Insecta Mundi 2024 (68): 1-5, DOI: 10.5281/zenodo.14662048

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19

Drăghici, Andreea-Cătălina. "Copris hispanus cavolinii (Coleoptera: Scarabaeidae) in the Collection of “Grigore Antipa” National Museum of Natural History (Bucharest)." Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 60, no. 1 (2017): 373–75. http://dx.doi.org/10.1515/travmu-2017-0005.

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Abstract Copris hispanus cavolinii (Petagna, 1792) is mentioned for the first time in the Coleoptera Collection of “Grigore Antipa” National Museum of Natural History (Bucharest). Three new records are added in the present short note for C. hispanus cavolinii based on the studied material.

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20

Deschodt, Christian M., Adrian L. V. Davis, and Clarke H. Scholtz. "A new synonymy in the fidius group of Copris Müller 1764 (Coleoptera: Scarabaeidae: Scarabaeinae) and a new species from the highland grasslands of South Africa." Zootaxa 3949, no. 3 (2015): 431–38. https://doi.org/10.11646/zootaxa.3949.3.8.

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Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2015): A new synonymy in the fidius group of Copris Müller 1764 (Coleoptera: Scarabaeidae: Scarabaeinae) and a new species from the highland grasslands of South Africa. Zootaxa 3949 (3): 431-438, DOI: 10.11646/zootaxa.3949.3.8

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21

Stanbrook, Roisin, Claire Raisin, and Kevina Vulinec. "Observations on the Tunneling Behavior and Seed Dispersal Efficacy of Copris nubilosus Kohlmann, Cano, and Delgado (Coleoptera: Scarabaeinae: Coprini)." Coleopterists Bulletin 71, no. 4 (2017): 777–80. https://doi.org/10.1649/0010-065X-71.4.777.

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Stanbrook, Roisin, Raisin, Claire, Vulinec, Kevina (2017): Observations on the Tunneling Behavior and Seed Dispersal Efficacy of Copris nubilosus Kohlmann, Cano, and Delgado (Coleoptera: Scarabaeinae: Coprini). The Coleopterists Bulletin 71 (4): 777-780, DOI: 10.1649/0010-065X-71.4.777, URL: http://dx.doi.org/10.1649/0010-065x-71.4.777

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22

Zidek, Jiri. "Catalogue of species-group names assigned to Copris Geoffroy, Coptodactyla Burmeister, Litocopris Waterhouse, Microcopris Balthasar, Paracopris Balthasar, Pseudocopris Ferreira, Pseudopedaria Felsche, Sinocopris Ochi, Kon and Bai, and Thyregis Blackburn (Coleoptera: Scarabaeidae: Scarabaeinae: Coprini)." Insecta Mundi, no. 756 (March 27, 2020): 1–24. https://doi.org/10.5281/zenodo.3746274.

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Zidek, Jiri (2020): Catalogue of species-group names assigned to Copris Geoffroy, Coptodactyla Burmeister, Litocopris Waterhouse, Microcopris Balthasar, Paracopris Balthasar, Pseudocopris Ferreira, Pseudopedaria Felsche, Sinocopris Ochi, Kon and Bai, and Thyregis Blackburn (Coleoptera: Scarabaeidae: Scarabaeinae: Coprini). Insecta Mundi 756: 1-24, DOI: 10.5281/zenodo.3746274

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23

Bang, H. S., O. S. Kwon, S. J. Hwang, Y. I. Mah, and Keith Grant Watdhaugh. "Developmental Biology and Phenology of a Korean Native Dung Beetle, Copris ochus (Motschulsky) (Coleoptera: Scarabaeidae)." Coleopterists Bulletin 58, no. 4 (2004): 522–33. https://doi.org/10.1649/662.

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Bang, H. S., Kwon, O. S., Hwang, S. J., Mah, Y. I., Watdhaugh, Keith Grant (2004): Developmental Biology and Phenology of a Korean Native Dung Beetle, Copris ochus (Motschulsky) (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 58 (4): 522-533, DOI: 10.1649/662, URL: http://dx.doi.org/10.1649/662

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24

Zidek, Jiri. "Addendum and update to Zidek (2020): Catalogue of species-group names assigned to Copris Geoffroy, Coptodactyla Burmeister, Litocopris Waterhouse, Microcopris Balthasar, Paracopris Balthasar, Pseudocopris Ferreira, Pseudopedaria Felsche, Sinocopris Ochi, Kon and Bai, and Thyregis Blackburn." Insecta Mundi 2020, no. 821 (2020): 1–3. https://doi.org/10.5281/zenodo.5353673.

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Zidek, Jiri (2020): Addendum and update to Zidek (2020): Catalogue of species-group names assigned to Copris Geoffroy, Coptodactyla Burmeister, Litocopris Waterhouse, Microcopris Balthasar, Paracopris Balthasar, Pseudocopris Ferreira, Pseudopedaria Felsche, Sinocopris Ochi, Kon and Bai, and Thyregis Blackburn. Insecta Mundi 2020 (821): 1-3, DOI: 10.5281/zenodo.5353673

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25

Bunalski, Marek, Najmeh Samin, Hassan Ghahari, and Trevor J. Hawkeswood. "Contributions to the knowledge the scarab beetles of Golestan province, Northern Iran with checklist of Iranian Scarabaeoidea (Coleoptera)." Polish Journal of Entomology 83, no. 2 (2014): 141–70. http://dx.doi.org/10.2478/pjen-2014-0011.

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AbstractThis paper deals with the fauna of Scarabaeoidea (Coleoptera) in Golestan province, Northern Iran. In total 19 species from 16 genera (Adoretus, Aphodius, Amphimallon, Cetonischema, Copris, Eulasia, Euonthophagus, Glaphyrus, Netocia, Onthophagus, Oxythyrea, Pentodon, Valgus, Polyphylla, Stalagmosoma, Trichius) were collected and recorded from this region. A checklist of Iranian Scarabaeoidea is provided.

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26

SOLÍS, ÁNGEL, and BERT KOHLMANN. "New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama." Zootaxa 139, no. 1 (2003): 1. http://dx.doi.org/10.11646/zootaxa.139.1.1.

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The following new species of dung beetles (Coleoptera: Scarabaeidae) from Costa Rica and Panama are described: Copris tridentatus sp. nov., Onthophagus notiodes sp. nov., Onthophagus xiphias sp. nov. and Trichillum arcus sp. nov. Pedaridium bradyporum (Boucomont) is reported from the Pacific slope of Costa Rica for the first time. Drawings of the dorsal habitus of all species are provided.

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27

Poloni, Riccardo, and Alessio Morelli. "New or interesting beetles from the Abruzzi region (Central Italy) (Insecta Coleoptera)." Bollettino della Società Entomologica Italiana 153, no. 1 (2021): 39–41. http://dx.doi.org/10.4081/bollettinosei.2021.39.

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New data on the beetle fauna of the Abruzzi region (Central Italy) are presented. New data on ecology and distribution are provided for three species of Cerambycidae previously recorded for the region and three species (one Cerambycidae and two Scarabaeoidea) are recorder for the first time for the region: Agapanthia (Epoptes) cynarae (Germar, 1817), Copris hispanus cavolinii (Petagna, 1792), Triodontella nitidula (Rossi, 1790).

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28

RÖSSNER, ECKEHARD, JOACHIM SCHÖNFELD, and DIRK AHRENS. "Onthophagus (Palaeonthophagus) medius (Kugelann, 1792)—a good western palaearctic species in the Onthophagus vacca complex (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini)." Zootaxa 2629, no. 1 (2010): 1. http://dx.doi.org/10.11646/zootaxa.2629.1.1.

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Based on a morphological taxonomic revision of available species, and supported by a separate molecular phylogenetic study, we identified Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) as a good western palaearctic species and a sibling species of O. vacca (Linnaeus, 1767). In this paper we diagnose and illustrate the key morphological features and distribution of both species. A neotype is designated for Copris medius Kugelann, 1792.

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29

Martínez Morales, Imelda, Carmen Huerta, and Magdalena Cruz. "Comportamiento reproductor en hembras de Copris incertus Say (Coleoptera, Scarabaeidae)." Bulletin de la Société entomologique de France 101, no. 2 (1996): 121–30. http://dx.doi.org/10.3406/bsef.1996.17225.

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30

Sugiura, Shinji, Yuichi Yamaura, and Hiroshi Makihara. "Sexual and Male Horn Dimorphism in Copris ochus (Coleoptera: Scarabaeidae)." Zoological Science 24, no. 11 (2007): 1082–85. http://dx.doi.org/10.2108/zsj.24.1082.

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31

Huerta, Carmen. "Il genere Copris Müller: Tassonomia, filogenesi e note di zoogeografía." Dugesiana 20, no. 1 (2013): 45. https://doi.org/10.32870/dugesiana.v20i1.4078.

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32

Bang, Hea-Son, Young-Eun Na, Myung-Hyun Kim, Kee-An Roh, and Jung-Taek Lee. "Effect of Temperature on the Development of Copris tripartitus Waterhouse (Coleoptera: Scarabaeidae)." Korean journal of applied entomology 46, no. 3 (2007): 357–62. http://dx.doi.org/10.5656/ksae.2007.46.3.357.

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33

López-Guerrero, Irma, Mario Zunino, and Gonzalo Halffter. "Taxonomic Use of Genitalic Characters in Mexican Copris (Coleoptera: Scarabaeidae, Scarabaeinae): The Case of Copris klugi sierrensis Matthews and the C. armatus Species Complex." Coleopterists Bulletin 63, no. 2 (2009): 203–12. http://dx.doi.org/10.1649/1145.1.

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34

Bezborodov, V. G. "The genus Copris (Coleoptera, Scarabaeidae) in the Far East of Russia." Entomological Review 96, no. 4 (2016): 432–38. http://dx.doi.org/10.1134/s0013873816040059.

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35

BANG, Hea Son, K. G. WARDHAUGH, Seok Jo HWANG, and Oh Seok KWON. "Development of Copris tripartitus (Coleoptera: Scarabaeidae) in Two Different Rearing Media." Entomological Research 33, no. 3 (2003): 201–4. http://dx.doi.org/10.1111/j.1748-5967.2003.tb00070.x.

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36

MASAN, Peter, and Bruce HALLIDAY. "Mesostigmatid mites associated with the dung beetle Copris lunaris (Coleoptera: Scarabaeidae)." European Journal of Entomology 106, no. 4 (2009): 545–50. http://dx.doi.org/10.14411/eje.2009.068.

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37

Indarjani, Raden, and Mochamad Miko. "DISTRIBUSI VERTIKAL KOMUNITAS KUMBANG KOTORAN Scarabaeidae DI HABITAT TAMAN NASIONAL GUNUNG SALAK." Konservasi Hayati 16, no. 2 (2020): 77–84. http://dx.doi.org/10.33369/hayati.v16i2.12670.

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Scarabaeidae dung beettles is a bioindicator through its function as decomposer, Scrabaeidae maintains the balance of environmental conditions that represented by the variation community structures. A study of Scarabaeidae was conducted to observe the variation of community structures related to different heights (900, 1400 dan 2000 masl) in Salak Halimun Mountains National Park (TNGHS), West Java. Pit Fall Dung Trap method was applied in a belt transect paralele with tracking line. In every height , there were 5 traps created with 10 meters distance among each other. In addition, the study also measured main abiotic paramters, such as type of susbtrate, soil acidity, soil temperature that presumed to have strong influence in establishment of the communities. The result showed that there was a variation in community structures related to different heights. In 900 masl, diversity index (H’) was 0.80 which caterogised as low, in 1400 masl, H’= 1,29 (middle) and H’=0,84 (low) was found in the 2000 masl. The community structures of Scarabaeidae in TNGHS were made up by genus of Onthophagus dan Copris that conssits of 4 (four) spesies, they were O. taurus, O. semicupreus, O. babirussoides dan Copris punctulatus. Onthophagus taurus considered as dominant species that filled 60 % or 339 individual of the communities. Meanwhile, Onthopagus babirussoides can be considered as rare species which only 8% been caught during study.

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Khaustov, Alexander A., and Andrey V. Frolov. "Five new species of Pavania (Acari: Heterostigmata: Dolichocybidae) phoretic on scarab beetles (Coleoptera: Scarabaeidae) from French Guiana and Malaysia." Systematic and Applied Acarology 25, no. 4 (2020): 707–27. http://dx.doi.org/10.11158/saa.25.4.10.

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Five new species of Pavania Lombardini (Acari: Heterostigmata: Dolichocybidae) associated with scarabaeine beetles are described: P. brevicaudata sp. nov., on Canthon vulcanoae in French Guiana; P. semireducta sp. nov., on Canthon quadriguttatus in French Guiana; P. pusilla sp. nov., on Canthon bicolor in French Guiana; P. hansreiaphila sp. nov., on Hansreia affinis in French Guiana; and P. copridis sp. nov., on Copris agnus in Borneo, Malaysia. A key to world species of Pavania is provided. The family Dolichocybidae is reported from French Guiana and Malaysia for the first time.

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Young, Orrey P. "SURVIVAL OF COPRIS MINUTUS (COLEOPTERA: SCARABAEIDAE) DURING CARRION AND DUNG FEEDING REGIMES." Journal of the New York Entomological Society 113, no. 3 & 4 (2005): 218–21. http://dx.doi.org/10.1664/0028-7199(2005)113[0218:socmcs]2.0.co;2.

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Kim, Hyeon-Jeong, Dong-Hee Kim, Jin-Young Lee, et al. "Study of Anti-inflammatory Effect of CopA3 Peptide Derived from Copris tripartitus." Journal of Life Science 23, no. 1 (2013): 38–43. http://dx.doi.org/10.5352/jls.2013.23.1.38.

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TAKAKU, Gen. "A Description of the Male of Pachylaelaps copris Ishikawa, 1984(Acari: Pachylaelapidae)." Journal of the Acarological Society of Japan 5, no. 1 (1996): 1–6. http://dx.doi.org/10.2300/acari.5.1.

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Tasbulatova, A. T., and A. Umirzakova. "Biodiversity of invertebrates in the Iirsu gorge of Sairam-Ugam SNNP." Kazakhstan zoological bulletin 3, no. 1 (2022): 21–26. http://dx.doi.org/10.54944/kzbww723hg70.

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In clause results, annual research insects living on territory of Sayram-Ygams national park are stated. The most common invertebrates are described. When collecting the material, standard entomological methods were used - collection with an entomological net, at night on an artificial light source, manual collection. As a result of research, about 70 specimens of Lepidoptera and 40 specimens of Coleoptera were collected. Rare Lepidoptera species: Hemaris fucifirmis (Linnaeus, 1758), Hemaris tityus (Linnaeus, 1758), Papilio alexanor voldemar Kreuzberg, 1989, Parnassius apollonius (Eversmann, 1847). Common species from the Coleoptera order are: Lytta flavovittata Ballion, 1878, Cetonia aurata (Linnaeus, 1760), Copris lunaris (Linnaeus, 1758).

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Natta, Gianluca, Alex Laini, Angela Roggero, Fabrizio Fabbriciani, Antonio Rolando, and Claudia Palestrini. "Behavioural Repeatability and Behavioural Syndrome in the Dung Beetle Copris umbilicatus (Coleoptera, Scarabaeidae)." Insects 14, no. 6 (2023): 529. http://dx.doi.org/10.3390/insects14060529.

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Although personality studies have primarily focused on vertebrates, the evidence showing invertebrates to be capable of displaying personalities has been steadily growing in recent years. In this study, we investigated the behavioural repeatability (repetition of a behaviour over time) and behavioural syndromes (a set of correlated behaviours) in Copris umbilicatus, which is a dung beetle species showing complex sub-social behaviour. We analysed three behaviours (activity, thanatosis and distress call emission) by measuring seven distinct behavioural traits (i.e., three activity-, one thanatosis- and three distress call-related traits). We found moderate to high levels of individual repeatability in all behavioural traits considered. The duration of thanatosis was inversely correlated with two activity traits, hinting a behavioural syndrome for thanatosis and activity, with bolder individuals exhibiting shorter thanatosis and higher locomotor activity in contrast with fearful individuals, which display longer thanatosis and poor locomotor activity. No relationships were found between the behavioural traits and body size or sex. Results of the principal component analysis (PCA) suggested personality differences among individuals. Dung beetles provide an impressive variety of ecosystem services. Since the provision of these services may depend on the personalities represented in local populations and communities, studies on the ecology of personality in dung beetles should be encouraged in future research.

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Cruz R, Magdalena, and Carmen Huerta. "COMPORTAMIENTO Y ACTIVIDAD REPRODUCTORA DE LOS MACHOS DE COPRIS INCERTUS SAY (COLEOPTERA SCARABAEIDAE: SCARABAEINAE)." ACTA ZOOLÓGICA MEXICANA (N.S.), no. 74 (August 14, 1998): 163–73. http://dx.doi.org/10.21829/azm.1998.74741723.

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Copris incertus es un escarabajo coprófago cavador cuya hembra elabora nidos que cuida hasta que la progenie emerge. En este trabajo se describen las modificaciones que presenta el aparato reproductor del macho desde la emergencia y a lo largo del período de nidificación. Se consideran principalmente las variaciones a nivel del testículo y glándulas anexas (mesadenias). Los individuos jóvenes presentan un volumen del folículo testicular y del reservorio glandular mínimos, las actividades espermatogénica y glandular son reducidas. Al madurar el individuo, tanto el volumen como la actividad testicular y glandular se incrementan notablemente. En individuos viejos se observa un proceso de degeneración testicular, pero la actividad secretora de las glándulas anexas se mantiene por la acumulación continua de sus secreciones en el reservorio.

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Kim, Sung Hun, Goeun Park, Jin-Soo Park, and Hak Cheol Kwon. "Antifungal Streptomyces spp., Plausible Partners for Brood-Caring of the Dung Beetle Copris tripartitus." Microorganisms 9, no. 9 (2021): 1980. http://dx.doi.org/10.3390/microorganisms9091980.

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The dung beetle Copris tripartitus Waterhouse (Coleoptera: Scarabaeidae) is a coprophagous insect that lives in and feeds primarily on the feces of mammalian herbivores and is known to protect their offspring from the pathogen-rich environment by performing parental care for brood balls. Brood balls under continuous management by dung beetle are rarely contaminated by entomopathogenic fungi compared to abandoned brood balls. On the supposition that dung beetles may benefit from mutualistic bacteria that protect their offspring against fungal pathogens, we evaluated the antifungal activities of bacteria isolated from the dung beetle and brood ball. As a result, bacterial isolates, mainly streptomycetes, manifested potent and broad-spectrum antifungal activity against various fungi, including entomopathogens. Of the isolates, Streptomyces sp. AT67 exhibited pronounced antifungal activities. Culture-dependent and independent approaches show that this strain has occurred continuously in dung beetles that were collected over three years. Moreover, metabolic profiling and chemical investigation demonstrated that the strain produced an antifungal polyene macrocyclic lactam, sceliphrolactam, as a major product. Our findings imply that specific symbiotic bacteria of C. tripartitus are likely to contribute brood ball hygiene by inhibiting fungal parasites in the environment.

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Akamine, Mayumi. "Size-dependent seasonal activity for males of the dung beetle Copris acutidens (Coleoptera: Scarabaeidae)." Canadian Entomologist 151, no. 6 (2019): 757–67. http://dx.doi.org/10.4039/tce.2019.55.

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AbstractThis study aimed to determine differences in activities between two male morphs of the dung beetle Copris acutidens Motschulsky (Coleoptera: Scarabaeidae) during the reproductive period and to examine the size distribution of reproductive males that stayed in nests. The activities of two male morphs distinguished by a threshold value of body size were compared with those by horn length. Regardless of body size or horn length, earlier activity of minor males was observed during the reproductive period. The sex ratio showed the greatest female bias when minor males were the more abundant than major males, indicating that minor males were the most active when competition was the weakest and these could avoid direct combat with major males. In morphs distinguished by horn length, more major males than minor males stayed in nests with females although the major males became the most active from the middle of the reproductive period. Thus, longer horns may directly confer a competitive advantage to males, enabling them to stay in nests with females, whereas early activity of minor males does not always indicate the effect of horn length directly. Therefore, this behaviour may occur regardless of whether the morphs differ in body size or horn length.

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Nguyen-Phung, Thérèse. "Révision des Copris afro-tropicaux. V. Une nouvelle espèce du « groupe fidius Olivier » [Col. Scarabaeidae]." Bulletin de la Société entomologique de France 91, no. 9 (1986): 271–73. http://dx.doi.org/10.3406/bsef.1986.18229.

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Huerta, Carmen, and Hea Son Bang. "Fecundity and Offspring Survival of Copris tripartitus Waterhouse (Coleoptera, Scarabaeidae: Scarabaeinae) Under Laboratory Rearing Conditions." Coleopterists Bulletin 58, no. 4 (2004): 501–7. http://dx.doi.org/10.1649/638.

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Bang, H. S., O. S. Kwon, S. J. Hwang, Y. I. Mah, and Keith Grant Watdhaugh. "Developmental Biology and Phenology of a Korean Native Dung Beetle, Copris ochus (Motschulsky) (Coleoptera: Scarabaeidae)." Coleopterists Bulletin 58, no. 4 (2004): 522–33. http://dx.doi.org/10.1649/662.

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Hwang, Jae-Sam, Juneyoung Lee, Yeon-Ju Kim, et al. "Isolation and Characterization of a Defensin-Like Peptide (Coprisin) from the Dung Beetle, Copris tripartitus." International Journal of Peptides 2009 (October 22, 2009): 1–5. http://dx.doi.org/10.1155/2009/136284.

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The antibacterial activity of immune-related peptides, identified by a differential gene expression analysis, was investigated to suggest novel antibacterial peptides. A cDNA encoding a defensin-like peptide, Coprisin, was isolated from bacteria-immunized dung beetle, Copris tripartitus, by using differential dot blot hybridization. Northern blot analysis showed that Coprisin mRNA was up-regulated from 4 hours after bacteria injection and its expression level was reached a peak at 16 hours. The deduced amino acid sequence of Coprisin was composed of 80 amino acids with a predicted molecular weight of 8.6 kDa and a pI of 8.7. The amino acid sequence of mature Coprisin was found to be 79.1% and 67.4% identical to those of defensin-like peptides of Anomala cuprea and Allomyrina dichotoma, respectively. We also investigated active sequences of Coprisin by using amino acid modification. The result showed that the 9-mer peptide, LLCIALRKK-NH2, exhibited potent antibacterial activities against Escherichia coli and Staphylococcus aureus.

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