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1

Mohammed, Nora Mohammed Al-Saqaf, та Othman Saad Saeed Al-Hawshabi. "الصفات المظهرية لـجنسي الـ MELHANIA (DOMBEYOIDEAE) و STERCULIA (STERCULIOIDEAE) من الفصيلة الخبازية بمفهومها الواسع MALVACEAE S.L. في مديرية طورالباحة، محافظة لحج، اليمن". Electronic Journal of University of Aden for Basic and Applied Sciences 3, № 1 (2022): 20–27. http://dx.doi.org/10.47372/ejua-ba.2022.1.140.

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تناولت الدراسة الحالية الصفات المظهرية لجنسي الـ Melhania (Dombeyoideae) و Sterculia (Sterculioideae) ضمن الفصيلة الخبازية بمفهومها الواسع Malvaceae s.l.، في مديرية طورالباحة – محافظة لحج – اليمن، وذلك خلال الفترة من أغسطس 2020 حتى يناير 2022م، إذ جُمْعِت عينات نباتية مثالية لكل نوع نباتي موجود في منطقة الدراسة. تم مناقشة الصفات المظهرية، كما تم سرد مرادفات الأسماء العلمية والأسماء المحلية لكل الأنواع المدروسة وانتشارها في منطقة الدراسة، وتم عمل مفتاح صناعي للتفرقة بين تحت الفصيلتين باستعمال أكثر الصفات المظهرية وضوحاً، كذلك تمت مراجعة التسمية العلمية واستحداثها باستخدام التقسيم الحديث الذي تم فيه إحالة الفصيلة الستروكولية Sterculiaceae إلى تحت الفصيلة Sterculioideae ضمن الفصيلة الخبازية بمفهومها الواسع Malvaceae s.l.، كذلك في هذه الدراسة تم التعامل مع جنس الـ Melhania ضمن تحت الفصيلة Dombeyoideae والذي كان سابقاً ضمن الفصيلة الستروكولية Sterculiaceae. تناولت الدراسة الصفات المظهرية لـنوعين نباتيين تنموان برياً في منطقة الدراسة هما (M. muricata و S. africana)، واثبتت الدراسة أن الصفات المظهرية الخضرية والتكاثرية ذات قيمة عالية في التمييز بين تحت الفصيلتين Dombeyoideae و Sterculioideae)، وبين النوعين (Melhania muricata و Sterculia africana).
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2

Boura, Anaïs, Timothée Le Péchon, and Romain Thomas. "Wood anatomy of the Mascarene Dombeyoideae: Systematic and ecological implications." IAWA Journal 32, no. 4 (2011): 493–519. http://dx.doi.org/10.1163/22941932-90000073.

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The Dombeyoideae (Malvaceae) are one of the most diversified groups of plants in the Mascarene Islands. Species of Dombeya Cav., Ruizia Cav. and Trochetia DC. are distributed in almost all parts of the archipelago and show a wide diversity in their growth forms. This study provides the first wood anatomical descriptions of 17 out of the 22 Mascarene species of Dombeyoideae. Their wood anatomy is similar to that of previously described species: wide vessels, presence of both apotracheal and paratracheal parenchyma, and storied structure. In addition, we also found a second wood anatomical pattern with narrower vessels, high vessel frequency and thick-walled fibres. The two aforementioned wood patterns are considered in a phylogenetic context and used to trace the evolutionary history of several wood anatomical features. For example, the pseudoscalariform pit arrangement supports a sister group relationship between Trochetia granulata Cordem. and T. blackburniana Bojer ex Baker and may be a new synapomorphy of the genus Trochetia. Finally, wood variability is evaluated in relation to geographic, climatic and biological data. Despite the juvenile nature of some of the specimens studied, we discuss how the habit, but also factors related to humidity, influence the variability observed in the Mascarene Dombeyoideae wood structure.
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3

Dorr, Laurence J. "A new combination in the endemic Malagasy genus Helmiopsis (Malvaceae-Dombeyoideae)." Adansonia 23, no. 1 (2001): 151–53. https://doi.org/10.5281/zenodo.4605007.

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4

Barnett, L. C., and L. J. Dorr. "Validation of Nesogordonia holtzii (Malvaceae: Dombeyoideae)." Kew Bulletin 55, no. 4 (2000): 985. http://dx.doi.org/10.2307/4113647.

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5

Jenny, Matthias, Clemens Bayer, and Laurence J. Dorr. "Aethiocarpa reduced to Harmsia (Malvaceae, Dombeyoideae)." TAXON 48, no. 1 (1999): 3–6. http://dx.doi.org/10.2307/1224611.

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6

Dorr, Laurence J., and Kenneth J. Wurdack. "A new disjunct species of Eriolaena (Malvaceae, Dombeyoideae) from Continental Africa." PhytoKeys 111 (November 6, 2018): 11–16. http://dx.doi.org/10.3897/phytokeys.111.29303.

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EriolaenarulkensiiDorr,sp. nov. is described and illustrated. This attractive shrub is endemic to coastal Mozambique. The new species has apically winged seeds, which place it in a group of Malvaceae (Dombeyoideae) that is found in Asia and Madagascar and which had not previously been found in continental Africa.
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7

Dorr, Laurence J., and Kenneth J. Wurdack. "A new disjunct species of Eriolaena (Malvaceae, Dombeyoideae) from Continental Africa." PhytoKeys 111 (November 6, 2018): 11–16. https://doi.org/10.3897/phytokeys.111.29303.

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Eriolaena rulkensii Dorr, sp. nov. is described and illustrated. This attractive shrub is endemic to coastal Mozambique. The new species has apically winged seeds, which place it in a group of Malvaceae (Dombeyoideae) that is found in Asia and Madagascar and which had not previously been found in continental Africa.
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8

Ganesan, S. K., D. J. Middleton, and P. Wilkie. "A REVISION OF PTEROSPERMUM (MALVACEAE: DOMBEYOIDEAE) IN MALESIA." Edinburgh Journal of Botany 77, no. 2 (2019): 161–241. http://dx.doi.org/10.1017/s0960428619000337.

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The species of the tree genus Pterospermum Schreb. (Malvaceae: Dombeyoideae) in Malesia are revised. Twenty-six species of Pterospermum are recognised, six of which are new (P. aureum S.K.Ganesan, P. borneense S.K.Ganesan, P. glabrum S.K.Ganesan, P. havilandii S.K.Ganesan, P. merrillianum S.K.Ganesan and P. zollingerianum S.K.Ganesan) and one (P. grewiifolium Pierre) that is a new distributional record for Malesia. Identification keys are provided. All names are typified, and detailed descriptions of all species recognised are provided. Information on habitat, uses and conservation status is given for all species.
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9

Wilkie, P. "A NEW SPECIES OF PTEROSPERMUM (DOMBEYOIDEAE, MALVACEAE/STERCULIACEAE) FROM CAMBODIA AND VIETNAM." Edinburgh Journal of Botany 64, no. 2 (2007): 179–83. http://dx.doi.org/10.1017/s0960428607000881.

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AbstractPterospermum mengii P.Wilkie (Dombeyoideae, Malvaceae/Sterculiaceae) from Cambodia and Vietnam is described as new to science and illustrated. It is most similar to Pterospermum pecteniforme Kosterm., sharing a distinctive wavy ridge at the margin of each locule of the capsular fruit, but differing by its larger leaves, oblong to obovate leaf shape, deeply cordate leaf base, and stellate indumentum on the undersurface of the leaves and fruit.
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10

FIEVET, LUCILE JOURDAIN, TIMOTHEE LE PECHON, JEAN-YVES DUBUISSON, and WENDY APPLEQUIST. "Dombeya scorpioides (Malvaceae: Dombeyoideae), a new species from southeastern Madagascar." Phytotaxa 704, no. 3 (2025): 232–38. https://doi.org/10.11646/phytotaxa.704.3.2.

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Dombeya scorpioides, a new endemic species of Dombeya (Malvaceae) from southeastern Madagascar, is described. Morphological analysis places this species within Dombeya section Xeropetalum as defined by Arènes. The species is distinguished by its unique biparous cyme, unusually elongated with mostly scorpioid terminal units. It is further differentiated from other species with similar narrow leaves by its hispid ovary with well-developed stellate hairs. Due to its restricted geographic distribution and past and future threats to its habitat, this species should be classified as Endangered according to IUCN Red List Criteria.
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11

Le Péchon, Timothée, Qiang Dai, Li-Bing Zhang, Xin-Fen Gao, and Hervé Sauquet. "Diversification of Dombeyoideae (Malvaceae) in the Mascarenes: Old Taxa on Young Islands?" International Journal of Plant Sciences 176, no. 3 (2015): 211–21. http://dx.doi.org/10.1086/679350.

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12

Péchon, Timothée Le, Nathanaël Cao, Jean-Yves Dubuisson, and Luc D. B. Gigord. "Systematics of Dombeyoideae (Malvaceae) in the Mascarene archipelago (Indian Ocean) inferred from morphology." TAXON 58, no. 2 (2009): 519–31. http://dx.doi.org/10.1002/tax.582016.

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13

Juhari, Muhammad Amirul Aiman Ahmad, Noraini Talip, Che Nurul Aini Che Amri, et al. "Taxonomic implications of pollen of some species of the genus Pterospermum Schreb. (Malvaceae s.l. subfam. Dombeyoideae)." Acta Botanica Brasilica 35, no. 4 (2021): 495–502. http://dx.doi.org/10.1590/0102-33062020abb0442.

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14

Dorr, Laurence J. "The correct author citation of the combination Eriolaena quinquelocularis (Malvaceae: Dombeyoideae) and typification of its basionym." Taxon 63, no. 6 (2014): 1340–41. http://dx.doi.org/10.12705/636.2.

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15

Tran, Van Tien, The Bach Tran, Ha Phuong Vu, Hoang Tuan Cao, and Thu Ha Bui. "Eriolaena bacgiangensis (Malvaceae), a new species from Vietnam." PhytoKeys 256 (May 2, 2025): 13–20. https://doi.org/10.3897/phytokeys.256.152884.

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The new species <i>Eriolaena bacgiangensis</i> from Vietnam is described and illustrated. The identified key was constructed to indicate the differences amongst three allied species of <i>Eriolaena</i>. <i>E. bacgiangensis</i> was compared with similar species, <i>E. candollei</i> and <i>E. wallichii</i>. <i>E. bacgiangensis</i> differs from <i>E. candollei</i> by the length of pedicel (1.4–2.3 cm long vs. 3.0–4.0 cm in <i>E. candollei</i>); presence of epicalyx after flowers at anthesis in <i>E. bacgiangensis</i> (vs. absence of epicalyx after flowers at anthesis in <i>E. candollei</i>); densely fringed epicalyx in <i>E. bacgiangensis</i> (vs. sparsely fringed epicalyx in <i>E. candollei</i>); style significantly exceeds staminal tube length in <i>E. bacgiangensis</i>, while the style is slightly longer than the staminal tube in <i>E. candollei</i>; the fruit apex round in <i>E. bacgiangensis</i> (vs. pointed and beaked in <i>E. candollei</i>). The linear epicalyx lobes are a key diagnostic trait for distinguishing <i>E. bacgiangensis</i> from <i>E. wallichii</i> (linear vs. ovate in <i>E. wallichii</i>). In addition, the other different characteristics of the two species are: leaf blade (not thickly papery in <i>E. bacgiangensis</i> vs. thickly papery in <i>E. wallichii</i>); length of inflorescences (13.0–18.0 cm in <i>E. bacgiangensis</i> vs. ≤ 6.0 cm in <i>E. wallichii</i>); shape of flower bud (lanceolate in <i>E. bacgiangensis</i> vs. globular in <i>E. wallichii</i>); fruit apex (round in <i>E. bacgiangensis</i> vs. apex shortly beaked in <i>E. wallichii</i>).
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16

Al-Hawshabi, Othman S. S., Ahmed K. Osman, Ahmed M. Abbas, Rahmah N. AlQthanin, and Mohamed O. Badry. "Flora and Phytochorology of Lahij Governorate of Yemen: 3-Systematic Revision of Malvaceae s.l. in Toor Al-Baha District." Diversity 17, no. 4 (2025): 242. https://doi.org/10.3390/d17040242.

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This study presents a thorough revision of the Malvaceae s.l. (sensu lato) family within the Flora of the Toor Al-Baha District, Lahij Governorate, South Yemen. The documented taxa were subjected to morphological revision alongside an analysis of life forms and chorological affinities. Thirty taxa are categorized into 12 genera within four subfamilies. Each species is presented with its current classification, accepted names, synonyms (when applicable), vernacular names (when applicable), photographs, and distribution to aid in identifying and recognizing these species. The documented mallow species comprises 80% perennials and 20% annuals. Chamaephytes constituted the predominant life forms, comprising 50% of the total taxa surveyed. A phytogeographical analysis revealed that chorotypes with Sudano-Zambezian affinities are predominant, comprising 83.34% of the recorded mallows. The two subfamilies, Grewioideae and Sterculioideae, are distinguished by the lack of an epicalyx and the presence of an androgynophore, unlike the subfamilies Dombeyoideae and Malvoideae. The highest richness genera included Grewia, with seven taxa, and Hibiscus, Abutilon, and Corchorus, each comprising four species. The research presented nine artificial keys for wild mallows in the Lahij Governorate, comprising three for subfamilies and six for genera.
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17

LE PÉCHON, TIMOTHÉE, SEBASTIEN TRACLET, FABIEN BARTHELAT, et al. "Bridging the Mozambique Channel: first record of Dombeya rosacea (Dombeyoideae, Malvaceae) for Mayotte (Comoros Archipelago, Indian Ocean)." Phytotaxa 460, no. 1 (2020): 75–88. http://dx.doi.org/10.11646/phytotaxa.460.1.4.

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In this study, we used an integrative approach combining morphological and molecular data with ecology, in order to clarify the identity of a Dombeya newly recorded as occurring in Mayotte. Morphological and molecular evidence are in agreement and convincingly show that specimens of this newly recorded species can be confidently assigned to D. rosacea, previously known only from four herbarium sheets from Madagascar. The new populations of this species in the Comoros significantly expand the extant area of occupancy of D. rosacea. Ecological preferences are also discussed, as populations of D. rosacea were found in greatly contrasting sub-humid habitats. This newly reported geographical disjunction between the Comoros and Madagascar strengthens the evidence for a biogeographic connection between the floras of these two regions.
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Le Péchon, Timothée, Jean-Yves Dubuisson, Thomas Haevermans, Corinne Cruaud, Arnaud Couloux, and Luc D. B. Gigord. "Multiple colonizations from Madagascar and converged acquisition of dioecy in the Mascarene Dombeyoideae (Malvaceae) as inferred from chloroplast and nuclear DNA sequence analyses." Annals of Botany 106, no. 2 (2010): 343–57. http://dx.doi.org/10.1093/aob/mcq116.

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19

Kottaimuthu, R. "A new name for Pterospermum gracile P. Wilkie (Malavaceae: Dombeyoideae)." Kew Bulletin 73, no. 3 (2018). http://dx.doi.org/10.1007/s12225-018-9763-4.

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20

Wu, Mingsong, Liu He, Guangyao Ma, Kai Zhang, Haijian Yang, and Xinquan Yang. "The complete chloroplast genome of Diplodiscus trichospermus and phylogenetic position of Brownlowioideae within Malvaceae." BMC Genomics 24, no. 1 (2023). http://dx.doi.org/10.1186/s12864-023-09680-z.

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Abstract Background Malvaceae is an economically important plant family of 4,225 species in nine subfamilies. Phylogenetic relationships among the nine subfamilies have always been controversial, especially for Brownlowioideae, whose phylogenetic position remains largely unknown due to the lack of samples in previous analysis datasets. To greatly clarify the phylogenetic relationship of Malvaceae, we newly sequenced and assembled the plastome of Diplodiscus trichospermus taxonomically located in Brownlowioideae, and downloaded the allied genomes from public database to build a dataset covering all subfamily members of Malvaceae. Results The annotation results showed that the plastome of Diplodiscus trichospermus has a typical quadripartite structure, comprising 112 unique genes, namely 78 protein-coding genes, 30 tRNA genes and 4 rRNA genes. The total length was 158,570 bp with 37.2% GC content. Based on the maximum likelihood method and Bayesian inference, a robust phylogenetic backbone of Malvaceae was reconstructed. The topology showed that Malvaceae was divided distinctly into two major branches which were previously recognized as Byttneriina and Malvadendrina. In the Malvadendrina clade, Malvoideae and Bombacoideae formed, as always, a close sister clade named as Malvatheca. Subfamily Helicteroideae occupied the most basal position and was followed by Sterculioideae which was sister to the alliance of Malvatheca, Brownlowioideae, Dombeyoideae, and Tilioideae. Brownlowioideae together with the clade comprising Dombeyoideae and Tilioideae formed a sister clade to Malvatheca. In addition, one specific conservation SSR and three specific palindrome sequences were observed in Brownlowioideae. Conclusions In this study, the phylogenetic framework of subfamilies in Malvaceae has been resolved clearly based on plastomes, which may contribute to a better understanding of the classification and plastome evolution for Malvaceae.
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Skema, Cynthia, Lucile Jourdain-Fievet, Jean-Yves Dubuisson, and Timothée Le Péchon. "Out of Madagascar, repeatedly: The phylogenetics and biogeography of Dombeyoideae (Malvaceae s.l.)." Molecular Phylogenetics and Evolution, December 2022, 107687. http://dx.doi.org/10.1016/j.ympev.2022.107687.

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22

Cvetković, Tijana, Fabiola Areces-Berazain, Damien D. Hinsinger, et al. "Phylogenomics resolves deep subfamilial relationships in Malvaceae s.l." G3 Genes|Genomes|Genetics 11, no. 7 (2021). http://dx.doi.org/10.1093/g3journal/jkab136.

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Abstract Malvaceae s.l., the most diverse family within Malvales, includes well-known species of great economic importance like cotton, cacao, and durian. Despite numerous phylogenetic analyses employing multiple markers, relationships between several of its nine subfamilies, particularly within the largest lineage /Malvadendrina, remain unclear. In this study, we attempted to resolve the relationships within the major clades of Malvaceae s.l. using plastid genomes of 48 accessions representing all subfamilies. Maximum likelihood and Bayesian analyses recovered a fully resolved and well-supported topology confirming the split of the family into /Byttneriina (/Grewioideae +/Byttnerioideae) and /Malvadendrina. Within /Malvadendrina, /Helicteroideae occupied the earliest branching position, followed by /Sterculioideae, /Brownlowioideae, /Tiliodeae, and /Dombeyoideae formed a clade sister to /Malvatheca (/Malvoideae +/Bombacoideae), a grouping morphologically supported by the lack of androgynophore. Results from dating analyses suggest that all subfamilies originated during hot or warm phases in the Late Cretaceous to Paleocene. This study presents a well-supported phylogenetic framework for Malvaceae s.l. that will aid downstream revisions and evolutionary studies of this economically important plant family.
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23

Le Péchon, Timothée, Alfred Bégué, Stéphane Buord, et al. "Friedmannodendron rodriguesiana , a new segregate of Dombeya Cav. (Dombeyoideae, Malvaceae), endemic to Rodrigues." Botany Letters, July 17, 2024, 1–12. http://dx.doi.org/10.1080/23818107.2024.2375733.

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24

Jourdain-Fievet, Lucile, Jean-Yves Dubuisson, Wendy Applequist, Cynthia Skema, and Timothée Le Pechon. "Updated nomenclature and re-establishment of the genus Dombeya Cav. section Dombeya (dombeyoideae, Malvaceae)." Botany Letters, March 6, 2024, 1–9. http://dx.doi.org/10.1080/23818107.2024.2316807.

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25

Dorr, Laurence J., and Kenneth J. Wurdack. "Indo‐Asian Eriolaena expanded to include two Malagasy genera, and other generic realignments based on molecular phylogenetics of Dombeyoideae (Malvaceae)." TAXON, November 17, 2020. http://dx.doi.org/10.1002/tax.12370.

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