Dissertations / Theses on the topic 'Ducks Waterfowl'

Strategies animals use to find and consume food in the face of conflicting forces such as competition and predation are central questions in the fields of ecological theory and management. Whereas theoretical models abound, proper empirical tests of these theories are less abundant. In studying the relationship between food abundance, predation risk, and competition there exists an array of confounding factors, which need to be accounted for by manipulating some aspect of the system. I used a guild of spring migratory ducks as a model system and manipulated food abundance in areas differing in presumed risk to assess the relative effects of food abundance, predation risk, competition, and life history characteristics on foraging strategies used by ducks. Using a randomized block design, I established a pair of 0.4 ha plots (block) in emergent, open water, and forested wetlands in the Wabash River Floodplain in eastern Illinois. I randomly selected one plot within each block to supplement with 2000 kg/ha of corn (Zea mays), creating an area of very high duck food abundance next to a control area with no added food. I conducted instantaneous focal animal samples and used video recorders to estimate the proportion of time mallard (Anas platyrhynchos), blue-winged teal (Anas discors), wood duck (Aix sponsa), ring-necked duck (Aythya collaris), and lesser scaup (Aythya affinis) devoted to feeding, identify the specific behaviors used, and estimate feeding stint lengths and frequencies. I used these metrics as an index of risk that ducks were willing to engage in for a known food reward. I predicted that species with a faster life history strategy, factors that increase perceived predation risk (cover, water depth, group size), and increase energetic demand (due to nesting or temperature), would elicit more risk-taking behaviors in ducks. This would be realized by an increase in the proportion of time spent feeding, longer feeding durations, and deeper feeding behaviors in treatment plots compared with control plots. Consistent with my life history prediction, species with a faster life history strategy were willing to engage in more risky behavior (feeding more) for a greater food reward (food treatment). Mallards, lesser scaup, and wood ducks exhibited risk-taking behavior consistent with perceived predation risk. Mallards devoted more time to feeding and used longer feeding stints when in areas with less cover. Alternatively, lesser scaup devoted more time to feeding when in areas of more cover. Wood ducks devoted more time to feeding in treatment plots, when in shallow areas, and larger flocks. When blue-winged teal fed on the surface (eyes above water), they devoted more overall time to feeding indicating that surface feeding is less risky than deeper feeding. Wood ducks and lesser scaup exhibited behavior consistent with an increase in energetic demand as observed by an increase in the proportion of time devoted to feeding later in the spring. I also examined how food abundance influenced aspects of ducks' foraging niches. I found that dabbling ducks used a greater variety of behaviors (behavior niche breadth) when in treatment plots compared to control plots and also shifted to slightly deeper feeding behaviors in treatment plots. This greater breadth when food was more abundant was due to individuals of the same species diverging from one another, rather than each individual using a broader array of behaviors. Overall, I found substantial variability among and within species in how they manage risk while foraging, although this was partially explained by life history theory, and what types of conditions they perceive as risky. I document the importance of taking the state of the forager (life history, perceived risk, energetic demand) into account when examining patterns of risk-taking.

This study is an examination of the use of coastal estuaries and nearby farmland as habitat by dabbling ducks (mallard and American wigeon) during migrating and wintering periods. Its aim was to identify aspects of British Columbia coastal habitat of importance to dabblers through an analysis of the ducks' movements among habitat types and through a description of where and on what ducks feed. Data were collected in 1979 and 1980 along a 30 km stretch of coastline on southeastern Vancouver Island. Results are presented in 3 parts. Part 1 examines the relative use of farm and coastal habitat through a series of censuses conducted weekly at 8 farm and 8 coastal sites. The strong negative correlation between counts at farm and coastal sites indicates that dabblers treat them as alternative habitats. The numbers of ducks on farms was positively correlated with the area of standing water on the fields. Farm habitat, apparently preferred during warm, wet weather, was not used when fields were dry or frozen. Part 2 is a description of feeding location on fields, at estuaries and at a shallow, nonestuarine bay. It is based on observations at selected sites at high and low tide levels. Each duck in each observation period was classified as to location and activity. Both species fed primarily in shallow water, their feeding location shifting with the tides. Both marsh and marine sections of estuaries were used extensively for feeding. The shallow bay was used especially by American wigeon at low tide in fall and early winter. The high marsh areas at estuaries were particularly attractive to mallards when flooded by exceptionally high tides. Feeding intensities were consistently high at farm sites for both species. In Part 3, 23 mallards and 40 American wigeon were shot while feeding in estuarine locations commonly used for feeding. Analysis of gullet contents revealed that both species ate a wide variety of items. Main foods were, for mallards, seeds, invertebrates and green algae and, for American wigeon, green algae, roots, seeds and green vegetation. Algae and marine Invertebrates are not usually considered to be important dabbler foods in estuaries. Major conclusions and recommendations are: 1) Both farm and coastal sites are important to dabblers, with fields being favoured as feeding locations under good flood conditions and coastal habitat being vital during dry or freezing periods. As dabblers move among sites, assessment and management of wintering dabbler habitat should be by wetland complexes rather than by individual estuaries. 2) Dabblers feed in or near shallow water. Fields that do not flood are not worth maintaining as dabbler habitat. Assessment of estuarine marshes should consider the availability of food at all points of the tidal cycle. 3) As both species feed on a wide variety of items, factors affecting shallow water flooding and thus food availability are more important than plant species composition. 4) Marine deltas and beaches are important as refuges from disturbance and as feeding grounds. They form an integral part of dabbler coastal habitat.
Science, Faculty of
Zoology, Department of
Graduate

The habitat use and energetics of American black ducks (Anas rubripes) wintering at Chincoteague National Wildlife Refuge, Virginia, were investigated. Twenty-two female black ducks were systematically radiotracked on the 25,600 ha study area between 15 December 1985 and 28 February 1986. Diurnal time and energy budgets were constructed by distributing 1,471 scans (collected in 1985-86 and 1986-87) over a time-tide matrix within refuge, saltmarsh, and tidal water habitats. Sixty-four ducks were collected during early, mid, and late winter in 1985-86 to determine changes in carcass composition. The Habitat Suitability Index (HSI) model for wintering black ducks was evaluated. Age affected range and core areas but did not affect habitat selection. Tide, ice, and time of day affected habitat use. Refuge pools were used during the day and saltmarsh was used at night. Subtidal water was used during periods of icing. Black ducks fed least and rested most when in refuge pools but fed most and rested least when in tidal waters. Black ducks curtailed feeding and increased time spent in alert and locomotion behaviors in response to disturbance. Whole carcass analysis indicated that black ducks were at least as fat and heavy in the spring as they were in the fall. Comparisons with similar work in Maine suggested that black ducks wintering in Maine and Virginia expend the same energy at a given temperature. However, because of lower temperatures, black ducks collected at Chincoteague were in relatively better condition than ducks wintering in Maine.
Master of Science

Muscovy ducks are native only to Central and South America, Mexico, and parts of southern Texas and are considered invasive in some areas outside of their native range. Although they have been introduced worldwide, they remain largely unstudied. The primary focus of this study was to relate Muscovy duck abundance to habitat characteristics of wetlands in Tampa, Florida. Muscovy abundance was measured using point count methods at 21 wetland sites that occur within an eight km radius of the University of South Florida’s main campus. Habitat features at these sites were assessed using field methods and Geographic Information Systems (GIS) (Arc 10.1v). Mann-Whitney U tests and Chi-squared tests were performed to identify significant differences between quantitative data groups. A Chi-squared test determined that there was not a positive correlation between Muscovy abundance and fountains or water regime, yet identified a significant relationship between Muscovy abundance and fencing, in which Muscovies did not frequently occupy ponds with fencing. Mann-Whitney U tests did not identify significances between Muscovy abundance and other habitat groups. Since Muscovy ducks are listed as an invasive species, identifying habitat preferences and deterrents will assist land managers and property owners with habitat modifications in preventing or controlling nuisance Muscovy populations.

Several species of raptors are found in prairie landscapes managed and enhanced for waterfowl. Red-tailed Hawks (Buteo jamaicensis ) and Great Horned Owls (Bubo virginianus) may benefit from such management in a manner that is counter to its goals and objectives; that is, waterfowl may comprise a significant proportion of their diet, resulting in a decline in waterfowl numbers. The overall aims of this three-year study were to determine whether the feeding habits of the two raptor species are selective and to determine if waterfowl is a preferred prey group. The diet was determined through pellet analysis, prey remains and direct nest observations during the nestling growth period. Availability of most prey species was assessed through small mammal trapping and by conducting waterfowl censuses. It was determined that both raptors select for duck species. The average waterfowl biomass consumed per nestling represented 21.5% of the total biomass consumed for Great Horned Owls and 23.5% for Red-tailed Hawks. (Abstract shortened by UMI.)

Selecting a nest site is an important decision for waterfowl. Because most nest failure is due to depredation, the primary selective pressure in choosing a nest site should be to reduce depredation risk. This task is difficult because predators use differing tactics to locate nests, such as olfactory or visual cues. I investigated several components of waterfowl nest-site selection and success on sites with shelterbelts (planted tree-rows) in North Dakota, during the 2006 and 2007 nesting seasons. I found that meteorological conditions impacted nest depredation; artificial nests were more likely to be depredated when either temperature or dew point was high. These meteorological conditions should improve foraging efficiency for olfactory predators by increasing odor concentration. Waterfowl selected nesting sites with greater visual concealment than random locations (lateral concealment). However, the only difference found between successful and depredated nests was lateral dispersion, an olfactory concealment characteristic. Nest density was higher in areas without shelterbelts than in areas near shelterbelts. Nest success for waterfowl decreased as shelterbelt height increased. Other shelterbelt characteristics, like porosity and orientation, did not affect nest success or nest density. Given that nest predators differ in foraging habitat, temporal patterns of activity, and searching modalities, nest site characteristics that conceal the nest from 1 predator species may increase its vulnerability to another predator. For instance, risk due to olfactory predators should be reduced near shelterbelts because locating nests would be more difficult as turbulence is generated by the shelterbelts. Concomitantly, shelterbelts could also increase the presence of visual predators, by providing nesting sites and vantage points. In my study, any benefits shelterbelts provide in reducing nest depredation by olfactory predators may have been offset by increasing nest depredation from visual predators. Hence nesting near shelterbelts was neither a liability nor a benefit to ducks.

Doctor of Philosophy
Department of Biology
David A. Haukos
Mottled ducks (Anas fulvigula) are dabbling waterfowl species native to coastal wetlands of the Gulf of Mexico of the United States and Mexico. Although closely related to common waterfowl species such as the mallard (A. platyrhynchos) and American black duck (A. rubripes), the mottled duck exhibits unique behavior, mainly in its life history as a non-migratory species. As such, because of population declines caused by predation, habitat destruction, and environmental contaminants, this species requires specialized conservation concerns and species-specific management to protect population numbers. The goal of this study was to assess ongoing effect of observed lead (Pb) contamination and exposure issues in mottled ducks and their habitats, which I achieved by conducting assessments that will provide managers habitat and organism level metrics to detect and mitigate lead in mottled ducks and their environments. My field study was conducted at the Texas Chenier Plain National Wildlife Refuge Complex (TCPC), which was the area of greatest mottled duck density on the Texas Coast. I first created a body condition index to provide managers a tool to monitor population health, and a proxy for lead exposure and avian health without destructively sampling individuals. I then used presence-only maximum entropy (MaxENT) and multivariate statistical modeling procedures in conjunction with mottled duck movement data to elucidate sets of habitat conditions that were conducive to predicting the occurrence of mottled ducks and environmental lead “hot spots”. MaxENT analyses suggested that lead in the top portion of the soil column is similarly related to all environmental variables considered, may be increasingly available after large-scale environmental disturbances. Lack of variation in coarse-scale habitat use between breeding and non-breeding seasons may further point to a food-based exposure pathway for lead as mottled ducks switch from an invertebrate to plant diet, either as a result of changing age classes or normal adult phenology, during the period of increased lead exposure. Using stable isotope ratio analysis, I then tested environmental samples of soil and vegetation as well as mottled duck blood to determine isotopic signatures that were consistent with particular sources of lead deposition (e.g., lead shot pellets, leaded fossil fuel combustion, industrial effluents). Comparisons suggested a great deal of similarity to lead shot reference values in vegetation and blood samples, especially in blood samples with higher concentrations of lead present. Last, I conducted a formal Ecological Risk Assessment (ERA) procedure to quantify the risk to mottled ducks from lead exposure in their current habitat and direct managers towards effective mitigation and habitat management strategies to reduce exposure in the future. One scenario suggested that mottled ducks were at greatest risk from eating an invertebrate-based diet, but lead content values at the TCPC suggest that a plant-based diet may provide a higher lead exposure risk for mottled ducks, depending on true levels of bioavailability in environmental media. Overall, I determined that mottled ducks experience greatest lead exposure risk from lead shot pellets on the TCPC or in nearby habitat, while potentially also experiencing low levels of exposure from several other sources. Additionally, management efforts that focus on plants that do not provide food resources for mottled ducks as a potential environmental sink for lead contamination, such as phytoremediation, may prove effective in reducing the overall lead load from historical activities that likely deposited much of the lead in this ecosystem.

The Laysan teal, an endangered species, is restricted to a single population on Laysan Island, a remote atoll of the Hawaiian archipelago. Little is known of the Laysan teal's ecology, therefore, I examined food habits, habitat use, and population dynamics. These aspects of its ecology are fundamental to the species management and conservation. I described diel and nocturnal habitat use, home range, and foraging with radio telemetry in 1998-2000. Most individuals showed strong site fidelity during the tracking period, but habitat selection varied between individuals. Mean home range size was 9.78 ha (SE 2.6) using the fixed kernel estimator (95% kernel; 15 birds with >25 locations). Foraging was strongly influenced by time of day: birds spent only 4% of their time foraging in the day, but spent 45% of their time foraging at night. Time activity budgets from the island's four habitat zones indicated that the coastal zone was rarely used for foraging. The birds foraged 42% of the time they spent in the terrestrial zone at night, but foraged only 4-6% of the time they spent there during other times. Fecal analysis and behavioral observations revealed that the Laysan teal is not a 100% macro-insectivore as previously reported, but consumed seeds, succulent leaves, and algae, in addition to adult diptera, diptera larvae and pupae, ants, seeds, lepidoptera, coleoptera, and Artemia. I concluded that this species exhibits high plasiticity in foraging behavior. Laysan teal appear to opportunistically select abundant, high energy prey for the breeding season, due to constrained resources on Laysan Island. I also studied the parameters influencing the Laysan teal's population dynamics. Adult survival is high, but duckling survival on Laysan is low, and is a primary demographic parameter limiting population growth. Estimates indicate the population density was high (between 546-827) from 1991 until August 1993, prior to a population crash that occurred between September and December 1993. The most current population estimate (Sept-Nov 2001) is 444 (SE 181) adults. Additional populations (translocation), along with control of non-native mammalian predators, are needed to reduce extinction risks to the Laysan teal.
Ph. D.

Clearing of old growth forests resulted in a substantial loss of nesting habitat for cavity-nesting waterfowl during the late 19th and early 20th centuries. Since then, many hardwood forests have matured into size classes capable of producing cavities suitable for nesting ducks. To quantify changes in cavity availability in U. S. Fish and Wildlife Service Region 3, which contains most of the midwestern U.S., I examined current cavity availability at 4 sites where cavity availability had been estimated in the past; Mingo National Wildlife Refuge (NWR) in Missouri, Shiawassee NWR in Michigan, Mead Wildlife Area (WA) in Wisconsin, and Muscatatuck NWR in Indiana. I found densities of 1.8 ± 0.4, 1.4 ± 0.3, 0.9 ± 0.4, and 1.8 ± 0.4 suitable cavities per hectare at each of these sites, respectively. Suitable cavities per hectare increased at Mingo NWR (433%) since 1966 and Shiawassee NWR (1400%) since 1974, but remained similar at Mead WA and Muscatatuck NWR since the mid-1980's, after accounting for differences in past, study-specific criteria for cavity suitability. Differences among sites were likely due to variation in species composition, stage of forest maturation, timber management, and time elapsed since the previous studies. Comparison of size-class distributions for all trees and for trees with cavities indicated that cavities occur in the largest trees and that forests have yet to mature into the most prolific cavity-producing size classes. This conclusion was corroborated by forest growth modeling results from Forest Vegetation Simulator, a forest growth-modeling program from the USDA Forest Service. I used Forest Inventory and Analysis data to model growth from 2008 to 2058 at 10-year intervals for Region 3. Cavity per tree estimates from the 4 study sites were applied to modeling outputs, and cavity availability was projected to almost double over the entire region by 2058. Thus, the observed and further expected increases in cavity density in the region justify re-examination of nest box programs and possibly a reduction of artificial nest boxes in some areas of the region where sufficient natural cavities exist. Because current and future estimates indicate sufficient nest sites for cavity-nesting waterfowl, efforts should be spent protecting and restoring brood-rearing wetlands, which are known to be declining in many areas of Region 3.

Military jet over-flight activities pose a potential threat to staging waterfowl and pilots. The migration period is important for nutrient acquisition and courtship as these waterfowl enter the breeding season. Using a focal animal (continuous) technique for five-minute intervals, diurnal Time/Activity (TA) budgets for Canada Geese (Branta canadensis canadensis) (n=751), American Black Duck (Anas rubripes) (n=474) and Common Goldeneye (Bucephala clangula) (n=1274) were compiled during 216 hours of behavioural observation. The study was conducted from 26 April to 27 May, 2002 at the outlet of Lac Fourmont, Labrador (52° 03' 30"N, 60° 31' 01" W), in an ashkui or area of open water in an otherwise frozen landscape, which is known historically as a place to hunt waterfowl. These staging grounds are within the 130 000 km2 Low-level Training Area (LLTA) of the Quebec-Labrador Peninsula. Generalized Linear Modeling (GLM) of ranked variables was used to analyze behaviour by: observer, sex, pair status, time of day, date, and each weather parameter. Male Common Goldeneye spent more time in courtship behaviours (2.7%) than females (1.1%) and they spent most of the daylight hours feeding (males 53.3% and females 54.5%), with little time resting (4.8% and 5.2%, respectively). In contrast, Black Ducks and Canada Geese spent relatively little time feeding (12.4% and 5% respectively) and most of their time sleeping (35% and 38% respectively) and locomotion (37.8% and 11% respectively). Ashkui are important to Common Goldeneyes for foraging, and to Canada Geese and Black Ducks for resting prior to the breeding season.
Ninety-one low-level jet over-flights occurred and sound levels (n=336 h) at the study site were measured. Effects of low-level jet over-flights were analyzed using GLM of ranked variables in order to analyze a number of variables simultaneously. All behavioural observations that occurred in the quarter hour periods up to 165 minutes after an over-flight were analyzed. Alert and courtship behaviours of Canada Geese increased after over-flights. Other behaviours were negatively affected to a lesser degree. Locomotor activities by Black Ducks increased significantly immediately following over-flights with a stronger movement response with increased noise. Increases in agonistic and comfort behaviours of Common Goldeneye were detected following over-flights with few other significant affects on their behaviour.
Key words. Anas rubripes, Branta canadensis, Bucephala clangula, disturbance, jet aircraft, Labrador, low-level flying, military activity, noise, spring staging, time-activity budgets, waterfowl

We collected 6,823 cloacal swabs over four years (2005–2006: 1,460; 2006– 2007: 2,171; 2007–2008: 2,424; and 2008–2009: 768) from 30 potential avian host species. Most samples (88.3 percent) were from dabbling ducks (genus Anas), while diving ducks (genus Aythya) accounted for 5.0 percent, and geese (genera Anser, Chen, and Branta) 3.0 percent of the samples tested. Waterfowl (Anatidae) comprised 98.7 percent of samples, with 1.8 percent from non-migratory dabbling ducks (genus Anas). All samples were screened for avian influenza virus (AIV) by AIV-matrix real-time RT-PCR (rRT-PCR); all rRT-PCR positive samples (541) were processed for virus isolation as well as 4,473 rRT-PCR negative samples. Differences were observed in apparent prevalence estimates over the four years between virus isolation (0.5, 1.3, 3.9, and 0.7 percent) and rRT-PCR (5.9, 6.5, 11.2, and 5.5 percent). We isolated 138 AIVs, of which two were obtained from rRT-PCR negative samples. Unlike previous reports of seasonal variation in AIV prevalence, we documented differences in prevalence estimates among months using rRT-PCR only during 2008–2009 and by virus isolation only during 2006–2007 and 2007–2008. Several of the AIV subtypes we identified are common in North America (e.g., H3, H4, and H6); H3N8 and H4N6 were the most common subtype combinations isolated. Similar to most surveillance studies, we found no significant difference in AIV infection based on host sex, but did find that juveniles were more likely to be positive for AIV than adults. We also documented that dabbling ducks were more likely to be positive for AIV than diving ducks, although not all dabbling ducks are equally likely to be positive. Molecular sequence analysis revealed no insertions of multiple basic amino acids at the cleavage site, which supported the identification of low pathogenic AIV. Phylogenetic anlyses performed on H5, H6, H7, N1, N2, N3, and N4 subtypes sequenced indicated similarity to other North American isolates with the exception of seven H6 which were more similar in amino acid translation to an isolate from Japan. In sum, this is the first multiyear study of avian influenza viruses on waterfowl wintering grounds of the Central Flyway, a historically understudied area of North America.

Several factors have contributed to impeding the progress of migratory waterfowl spatial modeling, such as (1) waterfowl’s reliance on wetlands, (2) lack of understanding about shifts in distributions through time, and (3) large-scale seasonal migration. This doctoral dissertation provides an array of tools to address each of these concerns in order to better understand and conserve this group of species. The second chapter of this dissertation addresses issues of modeling species dependent on wetlands, a dynamic and often ephemeral habitat type. Correlation models of the relationships between climatic variables and species occurrence will not capture the full habitat constraints of waterfowl. This study introduces a novel data source that explicitly models the depth to water table, which is a simulated long-term measure of the point where climate and geological/topographic water fluxes balance. The inclusion of the depth to water table data contributes significantly to the ability to predict species probability of occurrence. Furthermore, this data source provides advantages over traditional proxies for wetland habitat, because it is not a static measure of wetland location, and is not biased by sampling method. Utilizing the long-term banding bird data again, the third chapter examines the behavior of waterfowl niche selection through time. By using the methods developed in chapter two, probability of occurrence models for the 1950s and the 1990s were developed. It was then possible to detect movements in geographic and environmental space, and how movements in these two spaces are related. This type of analysis provides insight into how different bird species might respond to environment changes and potentially improve climate change forecasts. The final chapter presents a new method for predicting the migratory movement of waterfowl. The method incorporates not only the environmental constraints of stopover habitat, but also includes likely distance and bearing traveled from a source point. This approach uses the USGS’ banding bird database; more specifically, it relies on banding locations, which have multiple recoveries within short time periods. Models made from these banding locations create a framework of migration movement, and allow for predictions to be made from locations where no banding/recovery data are available.
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碩士
國立臺灣大學
獸醫學研究所
93
In the previous study goose parvovirus (GPV) was regarded to infect goose and Muscovy duck, and the Muscovy duck parvovirus (MDPV) was considered to infect goose only but not Muscovy duck. In our lab, a MDPV strain was isolated from goose. To understand the host range of the MDPV and its distribution in the host, white Roman goose and Muscovy duck were artificial infected with MDPV in this study. After the inoculation of MDPV in white Roman goose and Muscovy duck, the MDPV DNA was mainly detected in liver, small intestinal and kidney. The MDPV DNA was detected in most organs at 14th day post-inoculation (DPI). The MDPV DNA continued to be detected till the end of the study at the 16th and 18th DPI. MDPV DNA could be detected in the Muscovy duck, Pekin duck, tsaiya duck , and mule duck after experimental inoculation. The result showed MDPV could infect Muscovy duck, tsaiya duck, mule duck and Pekn duck and resulting in high mortality. To understand the occurrence of MDPV and GPV in the field, cohort study was performed in the duck farms in Yun-Lin. The results show that the vertical transmission of GPV or MDPV occurred in the field, and the occurrence of GPV might be more common than MDPV. The nucleotide sequence of the structure protein, VP3, of the field strains of MDPV and GPV were analyzed. The results that the similarity of GPV Taiwan strain and the four isolated strains was between 97.1-99.8％. The similarity of MDPV Hungary strain and the four isolated strains was between 88.5-99.0％. The variability was higher in MDPV group than in GPV group.