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1

Molina, Lázara Sotolongo, Maira Fernández Zequeira, and Pedro Herrera Oliver. "Pollen morphology of some Cuban Guettarda species (Rubiaceae: Guettardeae)." Grana 41, no. 3 (2002): 142–48. http://dx.doi.org/10.1080/001731302321042605.

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2

V., Nelson Zamora, and Luis J. Poveda. "Una Nueva Especie de Guettarda L. (Rubiaceae, Guettardeae) Para Costa Rica." Annals of the Missouri Botanical Garden 75, no. 3 (1988): 1157. http://dx.doi.org/10.2307/2399388.

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3

Achille, Frédéric, Timothy J. Motley, Porter P. Lowry, and Joël Jérémie. "POLYPHYLY IN GUETTARDA L. (RUBIACEAE, GUETTARDEAE) BASED ON NRDNA ITS SEQUENCE DATA1,2." Annals of the Missouri Botanical Garden 93, no. 1 (2006): 103–21. http://dx.doi.org/10.3417/0026-6493(2006)93[103:piglrg]2.0.co;2.

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4

Chavez, Jayson G., Grecebio Jonathan D. Alejandro, and Ulrich Meve. "Lectotypification ofTimonius pachyphyllusMerr. (Rubiaceae, Guettardeae)." Adansonia 39, no. 2 (2017): 145–48. http://dx.doi.org/10.5252/a2017n2a5.

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5

Muangrom, Wanitcha, Markus Bacher, Andreas Berger, Karin Valant-Vetschera, Srunya Vajrodaya, and Johann Schinnerl. "A novel tryptophan-derived alkaloid and other constituents from Guettarda speciosa (Rubiaceae: Cinchonoideae–Guettardeae)." Biochemical Systematics and Ecology 95 (April 2021): 104239. http://dx.doi.org/10.1016/j.bse.2021.104239.

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6

Borhidi, A., K. Velasco-Gutierrez, and V. Vásquez-Martínez. "Estudios sobre las rubiáceas de México, L Una Guettarda nueva (Guettardeae, Rubiaceae) en Oaxaca." Acta Botanica Hungarica 57, no. 1-2 (2015): 23–27. http://dx.doi.org/10.1556/abot.57.2015.1-2.5.

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7

Taylor, Charlotte M., and David H. Lorence. "Rubiacearum Americanarum Magna Hama Pars II. New and Newly Circumscribed Taxa of Guettarda (Guettardeae)." Novon 11, no. 1 (2001): 127. http://dx.doi.org/10.2307/3393221.

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8

Borhidi, A. "Resinanthus, género nuevo de Rubiaceae (Guettardeae)." Acta Botanica Hungarica 49, no. 3-4 (2007): 311–17. http://dx.doi.org/10.1556/abot.49.2007.3-4.6.

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9

Achille, Frédéric. "Tinadendron, nouveau genre de Rubiaceae, Guettardeae de Mélanésie orientale." Adansonia 28, no. 1 (2006): 167–80. https://doi.org/10.5281/zenodo.5186761.

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10

Chavez, Jayson G., Grecebio Jonathan D. Alejandro, and Ulrich Meve. "Lectotypification of Timonius pachyphyllus Merr. (Rubiaceae, Guettardeae)." Adansonia 39, no. 2 (2017): 145–48. https://doi.org/10.5252/a2017n2a5.

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11

Pérez, David, and Fernando Alzate. "A new species of Tournefortiopsis (Guettardeae, Rubiaceae) from northern Colombia." Phytotaxa 609, no. 2 (2023): 145–50. https://doi.org/10.11646/phytotaxa.609.2.7.

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Pérez, David, Alzate, Fernando (2023): A new species of Tournefortiopsis (Guettardeae, Rubiaceae) from northern Colombia. Phytotaxa 609 (2): 145-150, DOI: 10.11646/phytotaxa.609.2.7, URL: http://dx.doi.org/10.11646/phytotaxa.609.2.7
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12

Chen, Junhao, and Khoon Meng Wong. "Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae)." Phytotaxa 446, no. 2 (2020): 114–20. https://doi.org/10.11646/phytotaxa.446.2.4.

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Chen, Junhao, Wong, Khoon Meng (2020): Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae). Phytotaxa 446 (2): 114-120, DOI: 10.11646/phytotaxa.446.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.446.2.4
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13

Hoo, Pui Kiat, Manickam Sugumaran, and Khoon Meng Wong. "Two new species of Timonius (Rubiaceae: Guettardeae) from the Malay Peninsula." Phytotaxa 599, no. 3 (2023): 171–82. https://doi.org/10.11646/phytotaxa.599.3.4.

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Hoo, Pui Kiat, Sugumaran, Manickam, Wong, Khoon Meng (2023): Two new species of Timonius (Rubiaceae: Guettardeae) from the Malay Peninsula. Phytotaxa 599 (3): 171-182, DOI: 10.11646/phytotaxa.599.3.4, URL: http://dx.doi.org/10.11646/phytotaxa.599.3.4
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14

Torres-Montúfar, Alejandro, Hilda Flores-Olvera, and Helga Ochoterena. "A Taxonomic Treatment of Rogiera (Guettardeae, Rubiaceae)." Systematic Botany 46, no. 1 (2021): 186–210. http://dx.doi.org/10.1600/036364421x16128061189413.

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Abstract—Rogiera (Guettardeae, Rubiaceae) is a Neotropical genus distributed from Mexico to northern South America that includes shrubs, treelets, or trees, which were previously treated in the taxonomically controversial and confused Rondeletia complex. Rogiera can be recognized among other Rubiaceae by the combination of multiflowered inflorescences, heterostylous flowers, quincuncial corolla aestivation, a hairy ring at the corolla mouth, and capsular fruits with loculicidal dehiscence. Despite the recent taxonomic circumscription of Rogiera there is not a comprehensive taxonomic treatment
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15

Darwin, Steven P. "Systematics of Timonius Subgenus Abbottia (Rubiaceae-Guettardeae)." Systematic Botany Monographs 42 (1994): 1. http://dx.doi.org/10.2307/25027838.

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16

Chavez, Jayson G., Ulrich Meve, Nicolai M. Nürk, and Sigrid Liede-Schumann. "Disentangling Antirhea (Rubiaceae): resurrection of Guettardella and description of the new genus Achilleanthus." Botanical Journal of the Linnean Society 197, no. 1 (2021): 85–103. http://dx.doi.org/10.1093/botlinnean/boab013.

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Abstract The Palaeotropical genus Antirhea (Guettardeae, Rubiaceae) currently comprises 37 dioecious species classified into three subgenera, Antirhea, Mesocarpa and Guettardella. It has a complex taxonomic history with varying generic delineation and variable application of species names. Recent phylogenetic studies have indicated the non-monophyly of Antirhea, but the limited sampling does not allow resolution of its circumscription or nomenclatural changes to be proposed. Here, we reconcile the generic boundaries of Antirhea based on phylogenetic analyses of two nuclear (ETS, ITS) and three
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17

Darwin, Steven P. "Six new species of Timonius (Rubiaceae: Guettardeae) from Papuasia." Brittonia 62, no. 2 (2010): 126–36. http://dx.doi.org/10.1007/s12228-009-9102-z.

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18

Torres-Montúfar, Alejandro, Elia Ramírez-Arriaga, Enrique Martínez-Hernández, and Helga Ochoterena. "Systematic relevance of the pollen morphology for the Rondeletia complex (Cinchonoideae, Rubiaceae)." Botanical Journal of the Linnean Society 194, no. 3 (2020): 290–307. http://dx.doi.org/10.1093/botlinnean/boaa047.

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Abstract Seventeen morphologically related genera have been considered as the Rondeletia complex, traditionally in Rondeletieae. Molecular studies agree that this is a polyphyletic group and some genera have been transferred to Guettardeae. Nevertheless, no solid morphological characters support this hypothesis. Our aim is to explore pollen morphology to see if this source of characters has systematic value. Our study uses light and scanning electron microscopy for 30 species of both tribes representing 14 genera. The most important characters are polar outline, endoaperture type and nexine di
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19

Wong, Sin Yeng. "Additions to the ‘Flora of Borneo: The vascular plant genera’ — I: Chewlunia (Rubiaceae) and Zygogynum (Winteraceae)." Webbia 79, no. 1 (2024): 51–52. http://dx.doi.org/10.36253/jopt-15522.

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The newly recognized genus Chewlunia Junhao Chen, P.K.Hoo & K.M.Wong (Rubiaceae: Guettardeae), with eight species (seven from the Philippines and one from Sabah), and the genus Zygogynum Baill. (Winteraceae), recently recorded for the first time for Borneo with a single, novel, species, are incorporated as entries in the Flora of Borneo: The vascular plant genera, taking the Bornean flora to 1594 native genera and 12,592 species.
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20

Ramos Álvarez, C. H., and A. Borhidi. "Una nueva especie del género Chomelia (Rubiaceae, Guettardeae) en México." Acta Botanica Hungarica 57, no. 1-2 (2015): 199–203. http://dx.doi.org/10.1556/abot.57.2015.1-2.15.

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21

Castillo-Campos, Gonzalo, and David H. Lorence. "Antirhea Aromatica (Rubiaceae, Guettardeae), A New Species from Veracruz, Mexico." Annals of the Missouri Botanical Garden 72, no. 2 (1985): 268. http://dx.doi.org/10.2307/2399181.

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22

PÉREZ, DAVID, and FERNANDO ALZATE. "A new species of Tournefortiopsis (Guettardeae, Rubiaceae) from northern Colombia." Phytotaxa 609, no. 2 (2023): 145–50. http://dx.doi.org/10.11646/phytotaxa.609.2.7.

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A new species of Tournefortiopsis from high Andean forests of the Central Colombian Cordillera is described and illustrated. Tournefortiopsis triflora is characterized mainly by its dichasium inflorescence, a feature not previously reported in the genus. It can be distinguished from its related species by its smaller leaves, peduncles, and corolla tube, and by the leaves abaxially glabrous on the areoles. The new species is from an area of biogeographic interest due to the large number of endemic species known to this area. A distribution map, conservation status and taxonomic affinities are p
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23

Darwin, Steven P. "New Species of the Timonius flavescens Alliance (Rubiaceae: Guettardeae) in Papuasia." Systematic Botany 22, no. 1 (1997): 85. http://dx.doi.org/10.2307/2419678.

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24

Borhidi, A. "Revalidation of the genusTournefortiopsisRusby, (Guettardeae, Rubiaceae) and a newGuettardafrom Costa Rica." Acta Botanica Hungarica 50, no. 1-2 (2008): 61–72. http://dx.doi.org/10.1556/abot.50.2008.1-2.3.

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25

Delprete, Piero G. "A new species of Chomelia (Rubiaceae, Guettardeae) from the Brazilian Planalto." Blumea - Biodiversity, Evolution and Biogeography of Plants 53, no. 2 (2008): 393–98. http://dx.doi.org/10.3767/000651908x608034.

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26

Amaya, Alix, and Alex Popovkin. "Malanea revolutifolia (Guettardeae: Rubiaceae), una nueva especie del noreste de Brasil." Rodriguésia 63, no. 4 (2012): 929–32. http://dx.doi.org/10.1590/s2175-78602012000400011.

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Se describe e ilustra Malanea revolutifolia, una nueva especie proveniente del bosque atlántico de Brasil, así mismo se incluye una clave para diferenciar algunas especies de Malanea que crecen en Brasil. M. revolutifolia es similar a M. forsteronioides y M. microphylla de las cuales se distingue principalmente por presentar hojas elípticas fuertemente revolutas 2,8-7,5 × 2-3,5 cm, cáliz profundamente dividido, lóbulos 3-4 mm de largo. Frutos drupas fusiformes a elípticas, pireno 5-7 × 2-2,5 mm.
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27

Salamanes, Julius John D. P., Jayson G. Chavez, Axel H. Arriola, and Grecebio Jonathan D. Alejandro. "A new species ofAntirhea(Guettardeae, Rubiaceae) from Surigao del Norte, Philippines." Acta Botanica Gallica 162, no. 3 (2015): 153–56. http://dx.doi.org/10.1080/12538078.2015.1062800.

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28

HOO, PUI KIAT, MANICKAM SUGUMARAN, and KHOON MENG WONG. "Two new species of Timonius (Rubiaceae: Guettardeae) from the Malay Peninsula." Phytotaxa 599, no. 3 (2023): 171–82. http://dx.doi.org/10.11646/phytotaxa.599.3.4.

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Two new species of Timonius endemic to the Malay Peninsula are described and illustrated: T. megalocalyx and T. denigricans, and their distribution and ecology discussed. Both species have been thought to be conspecific with their most resembling taxa in a previous revision but are here shown to be distinct.
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29

TORRES-MONTÚFAR, ALEJANDRO, and HELGA OCHOTERENA. "Taxonomic treatment of the Mesoamerican genus Rovaeanthus (Rondeletieae, Rubiaceae)." Phytotaxa 403, no. 2 (2019): 122. http://dx.doi.org/10.11646/phytotaxa.403.2.5.

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Rovaeanthus (Rondeletieae, Rubiaceae) is a Mesoamerican endemic genus of two species, characterized by a fleshy, pubescent ring at the corolla mouth. Despite the molecular evidence, which suggested that Rovaeanthus shares a most recent common ancestor with the rest of the tribe Rondeletieae, it was treated as a synonym of Rogiera (Guettardeae), with which it shares the pubescent ring at the corolla mouth. Although the species circumscription of Rovaeanthus is clear, its recognition as a separate genus is not yet widely accepted. An updated identification key for the two species of Rovaeanthus,
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30

Chavez, Jayson G., Grecebio Jonathan D. Alejandro, Danilo N. Tandang, and Ulrich Meve. "Two new species of Timonius (Guettardeae: Rubiaceae) from Samar Island, the Philippines." Botany Letters 167, no. 3 (2020): 363–72. http://dx.doi.org/10.1080/23818107.2020.1759449.

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31

Chen, Junhao, Hugh Tan, and Khoon Meng Wong. "A revision of Timonius (Rubiaceae) in Kinabalu Park, Borneo, with notes on typification and species distinction." Plant Ecology and Evolution 148, no. (3) (2015): 420–30. https://doi.org/10.5091/plecevo.2015.1073.

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<b>Background</b> – <i>Timonius</i> is the largest genus within the tribe Guettardeae of the family Rubiaceae. This genus is poorly studied, particularly in West Malesia. <b>Methods</b> – A revision of the genus in Kinabalu Park was conducted using ordinary practices of herbarium taxonomy. The specimen holdings of the BO, BRUN, K, L, SAN, SING and SNP herbaria were examined. <b>Results</b> – Thirteen species of <i>Timonius</i> are recognised in Kinabalu Park, of which seven are likely to be endemic to the Kinabalu massif including four that appear to occur exclusively on ultramafic substrates.
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CHEN, JUNHAO, and KHOON MENG WONG. "Typification and nomenclatural notes on Bornean Timonius (Rubiaceae: Guettardeae)." Phytotaxa 446, no. 2 (2020): 114–20. http://dx.doi.org/10.11646/phytotaxa.446.2.4.

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As part of a revision of Timonius in Borneo, the types of all relevant names were evaluated. Lectotypes are designated for 14 names in Timonius, including synonyms for accepted names. An erroneous original type citation is corrected and the nomenclatural history of a few names is clarified.
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33

Borhidi, A., and S. Salas-Morales. "Estudios sobre Rubiaceas Mexicanas, XXIX.Arachnothryx sanchezii(Guettardeae) yBouvardia mitlensis(Spermacoceae) nuevas especies de Oaxaca." Acta Botanica Hungarica 52, no. 1-2 (2010): 77–84. http://dx.doi.org/10.1556/abot.52.2010.1-2.8.

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34

Amaya Worm, Alix, Sirli Leython Chacón, María Raymúndez, Pedro Torrecilla, and Bruno Manara †. "Una especie nueva del género neotropical Malanea (Rubiaceae, Guettardeae) de la región Amazónica venezolana." Darwiniana, nueva serie 13, no. 1 (2025): 30–38. https://doi.org/10.14522/darwiniana.2025.131.1261.

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Se describe e ilustra Malanea maigualidensis, una nueva especie de la familia Rubiaceae que crece en la Sierra de Maigualida, el sistema montañoso de roca no sedimentaria más largo y ancho de la Amazonia venezolana. Malanea maigualidensis es similar a M. duckei y M. sarmentosa pero se puede distinguir por su densa pubescencia tomentosa blanco-dorada en la cara abaxial de las hojas y en el lado externo de las estípulas; limbo del cáliz lobulado, lóbulos desiguales de 1-1,5 mm y lóbulos de la corola 2,5-3 mm. En contraste, M. duckei se caracteriza por tener estípulas de 7-8 x 4-5 mm; cáliz subtr
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35

De Toni, K. L. G., and J. E. A. Mariath. "Ovule ontogeny in Rubiaceae (Juss.): Chomelia obtusa (Cinchonoideae–Guettardeae) and Ixora coccinea (Ixoroideae–Ixoreae)." Plant Systematics and Evolution 272, no. 1-4 (2008): 39–48. http://dx.doi.org/10.1007/s00606-007-0635-x.

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36

Taylor, Charlotte M. "Rubiacearum Americanarum Magna Hama Pars XIV. New Species of Malanea (Guettardeae) from Western South America." Novon 13, no. 2 (2003): 223. http://dx.doi.org/10.2307/3393526.

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37

Borhidi, A. L. "Revisión crítica del género Rogiera Planch. (Rubiaceae, Guettardeae) y la validez del género Rovaeanthus Borhidi (Rubiaceae, Rondeletieae)." Acta Botanica Hungarica 60, no. 1-2 (2018): 13–29. http://dx.doi.org/10.1556/034.60.2018.1-2.2.

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38

Chavez, Jayson G., Sigrid Liede-Schumann, and Ulrich Meve. "On the identity of the rubiaceous Timonius quadrasii (Guettardeae: Cinchonoideae): its typification and transfer to Rothmannia (Gardenieae: Ixoroideae)." Webbia 72, no. 2 (2017): 217–20. http://dx.doi.org/10.1080/00837792.2017.1386399.

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39

Taylor, Charlotte M., and Andreas Berger. "Rubiacearum Americanarum Magna Hama Pars XLIX: Overview of the Neotropical Genus Tournefortiopsis (Guettardeae), With Some New Species and Subspecies." Novon, A Journal for Botanical Nomenclature 29 (October 29, 2021): 227–65. http://dx.doi.org/10.3417/2021641.

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Guettarda L. (Rubiaceae) as traditionally circumscribed has been found to be polyphyletic in molecular analyses. Tournefortiopsis Rusby has been separated from it based on details of inflorescence arrangement, pyrene shape, and molecular analyses. We here find additional characters that distinguish this genus: valvate-induplicate corolla lobes and apical horns on the pyrenes. Some features that are usually consistent within species and often genera of Rubiaceae vary within some species of Tournefortiopsis, especially pubescence form and number of corolla lobes and locules. Our taxonomic review
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40

Pessoa, Maria do Céo R., Claes Persson, Alexandre Antonelli, and Maria Regina de V. Barbosa. "First record of Chomelia triflora (J.H. Kirkbr.) Delprete & Achille (Rubiaceae) from Brazil." Check List 13, no. (4) (2017): 159–62. https://doi.org/10.15560/13.4.159.

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This paper presents the first record of <em>Chomelia triflora</em> from Brazil, to date a species only known from French Guiana. After examining herbaria collections and doing fieldwork in the Brazilian Amazon, we found that the species also occurs in and around the Ducke Reserve in the state of Amazonas, Brazil. Our finding increases the data about the Brazilian Amazon and contributes to the better knowledge of <em>Chomelia </em>in Brazil.
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41

Rova, Johan H. E., Piero G. Delprete, and Birgitta Bremer. "The Rondeletia Complex (Rubiaceae): An Attempt to Use ITS,rps16, andtrnL-FSequence Data to Delimit Guettardeae, Rondeletieae, and Sections within Rondeletia1." Annals of the Missouri Botanical Garden 96, no. 1 (2009): 182–93. http://dx.doi.org/10.3417/2006179.

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42

GONZALEZ-ASTORGA, J. "Genetic Variability of the Narrow Endemic Tree Antirhea aromatica Castillo-Campos & Lorence, (Rubiaceae, Guettardeae) in a Tropical Forest of Mexico." Annals of Botany 93, no. 5 (2004): 521–28. http://dx.doi.org/10.1093/aob/mch070.

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43

Delprete, P. G., F. Achille, and A. Mouly. "Four new combinations in Chomelia and Stenostomum (Rubiaceae, Guettardeae) from Central America, the Guianas and the Amazon Basin." Blumea - Biodiversity, Evolution and Biogeography of Plants 55, no. 2 (2010): 164–70. http://dx.doi.org/10.3767/000651910x526906.

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44

Taylor, Charlotte M., and Roy E. Gereau. "Rubiacearum Americanarum Magna Hama Pars XXIII: Overview of the Guettardeae Tribe in Central and South America, with Five New Species and Three New Combinations in Chomelia, Neoblakea, Pittoniotis, and Stenostomum." Novon: A Journal for Botanical Nomenclature 20, no. 3 (2010): 351–62. http://dx.doi.org/10.3417/2009045.

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45

Roxo, Rafaela Sales Pereira, and Maria Regina de Vasconcellos Barbosa. "The genus Malanea Aubl. (Rubiaceae) in the Atlantic Forest of Brazil." Plant Ecology and Evolution 156, no. (3) (2023): 257–75. https://doi.org/10.5091/plecevo.103395.

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Background and aims – Malanea (Rubiaceae) is a Neotropical genus of ca 40 species, 15 of which occur in Brazil, with nine found in the Atlantic Forest domain. This study aims to present a taxonomic treatment of the species occurring in the Atlantic Forest domain. Material and methods – We studied collections deposited in 15 Brazilian herbaria, ALCB, BHCB, BHZB, CEPEC, HUEFS, HUFU, IPA, JPB, MAC, OUPR, PEUFR, RFA, RB, UFP, and UFRN, and collections in NY were studied online. Specimens were identified using protologues and digital images of types available online. Key results – Of the nine speci
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46

Makovicky, E., D. Topa, H. Tajjedin, E. Rastad, and A. Yaghubpur. "THE CRYSTAL STRUCTURE OF GUETTARDITE, PbAsSbS4, AND THE TWINNITE-GUETTARDITE PROBLEM." Canadian Mineralogist 50, no. 2 (2012): 253–65. http://dx.doi.org/10.3749/canmin.50.2.253.

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47

Ferrari, F., I. Messana, B. Botta, and J. F. de Mello. "Constituents of Guettarda platypoda." Journal of Natural Products 49, no. 6 (1986): 1150–51. http://dx.doi.org/10.1021/np50048a041.

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48

Sathasivampillai, Saravanan Vivekanandarajah, Moon Jain, and Pholtan Rajeev Sebastian Rajamanoharan. "BIOACTIVITIES OF GUETTARDA SPECIOSA L. EXTRACTS." Acta Scientifica Malaysia 5, no. 1 (2020): 20–22. http://dx.doi.org/10.26480/asm.01.2021.20.22.

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Guettarda speciosa L. is a tree of the Rubiaceae family. G. specious has been used to treat diarrhea, abscesses, wounds, epilepsy, and malaria in ethnomedicinal systems. So far, there is no systematic review for the bioactivities of G. speciosa. Therefore, the purpose of this minireview is to analyze, summarize and document the reported bioactivities of G. speciosa. Major electronic databases (Web of Science, Scopus, ScienceDirect, PubMed, and Semantic Scholar) were used to identify related studies from 1900 to January 2021. So far, in vivo and in vitro scientific data are available for the bi
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Matos, M. E. O., M. P. Sousa, M. L. L. Machado, and R. Braz Filho. "Quinovic acid glycosides from Guettarda angelica." Phytochemistry 25, no. 6 (1986): 1419–22. http://dx.doi.org/10.1016/s0031-9422(00)81301-1.

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Aquino, R., F. De Simone, F. Senatore, and C. Pizza. "Iridoids and secoiridoids from Guettarda platypoda." Pharmacological Research Communications 20 (December 1988): 105–8. http://dx.doi.org/10.1016/s0031-6989(88)80852-x.

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