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1

Arifullah, Ery, Yahdi Zaim, Aswan, and Djuhaeni. "Tiering style and its interpretation: ichnofabric study in Balikpapan Formation, Kutai Basin, Indonesia." Journal Mathematical and Fundamental Science 51, no. 1 (2019): 90–102. https://doi.org/10.5614/j.math.fund.sci.2019.51.1.8.

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The determination of tiering style is an important task in ichnology. Tiering styles can be modeled by using ichnodiversity, the number of behaviors and number of tiers and their relationships. In this study, this approach was applied in the area of Samarinda, Kutai Basin, Indonesia. The values of ichnodiversity, number of behaviors, and number of tiers were identified. Ichnodiversity was strongly correlated with number of behaviors but weakly correlated with number of tiers. Accordingly, three tiering styles were identified. Typically, tiering styles A, B, and C demonstrated high ichnodiversi
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2

Buatois, Luis A., Maria G. Mángano, Ricardo A. Olea, and Mark A. Wilson. "Decoupled evolution of soft and hard substrate communities during the Cambrian Explosion and Great Ordovician Biodiversification Event." Proceedings of the National Academy of Sciences 113, no. 25 (2016): 6945–48. http://dx.doi.org/10.1073/pnas.1523087113.

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Contrasts between the Cambrian Explosion (CE) and the Great Ordovician Biodiversification Event (GOBE) have long been recognized. Whereas the vast majority of body plans were established as a result of the CE, taxonomic increases during the GOBE were manifested at lower taxonomic levels. Assessing changes of ichnodiversity and ichnodisparity as a result of these two evolutionary events may shed light on the dynamics of both radiations. The early Cambrian (series 1 and 2) displayed a dramatic increase in ichnodiversity and ichnodisparity in softground communities. In contrast to this evolutiona
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3

Buatois, Luis A., and M. Gabriela Mángano. "Ichnodiversity and ichnodisparity: significance and caveats." Lethaia 46, no. 3 (2013): 281–92. http://dx.doi.org/10.1111/let.12018.

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4

Meyer, Neele, Max Wisshak, and André Freiwald. "Bioerosion ichnodiversity in barnacles from the Ross Sea, Antarctica." Polar Biology 44, no. 4 (2021): 667–82. http://dx.doi.org/10.1007/s00300-021-02825-4.

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AbstractBreakdown of skeletal and lithic hard substrates by organisms, a process referred to as bioerosion, is part of the global carbon cycle and receives increased attention, but little is known about bioerosion in polar environments. Here, we study bioerosion traces (addressed by their respective ichnotaxa) recorded in the barnacle Bathylasma corolliforme from the Ross Sea, Antarctica. Traces were visualized via scanning electron microscopy of epoxy casts prepared with the vacuum cast-embedding technique. In 50 samples from shallow 37 m to bathyal 1680 m water depths, 16 different bioerosio
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5

Bellosi, Eduardo, Jorge F. Genise, Mirta González, and Mariano Verde. "Paleogene laterites bearing the highest insect ichnodiversity in paleosols." Geology 44, no. 2 (2016): 119–22. http://dx.doi.org/10.1130/g37250.1.

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6

Luo, Mao, Luis A. Buatois, G. R. Shi, and Zhong-Qiang Chen. "Infaunal response during the end-Permian mass extinction." GSA Bulletin 133, no. 1-2 (2020): 91–99. http://dx.doi.org/10.1130/b35524.1.

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Abstract The end-Permian mass extinction (EPME) profoundly shaped shallow marine ecosystems. Although much has been learned about this event based on the body-fossil record, the global infaunal response to the EPME, as represented by ichnofossils, is much less understood. Here we analyze secular changes in ichnodiversity and ichnodisparity from the late Permian to the Middle Triassic based on a global trace-fossil data set. Results show that, in contrast to the body-fossil record, late Permian global ichnodiversity and ichnodisparity maintained their level until the Griesbachian, followed by a
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7

Knaust, Dirk, Michał Warchoł, and Ian A. Kane. "Ichnodiversity and ichnoabundance: Revealing depositional trends in a confined turbidite system." Sedimentology 61, no. 7 (2014): 2218–67. http://dx.doi.org/10.1111/sed.12134.

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8

WISSHAK, M., A. TRIBOLLET, S. GOLUBIC, J. JAKOBSEN, and A. FREIWALD. "Temperate bioerosion: ichnodiversity and biodiversity from intertidal to bathyal depths (Azores)." Geobiology 9, no. 6 (2011): 492–520. http://dx.doi.org/10.1111/j.1472-4669.2011.00299.x.

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9

SHILLITO, ANTHONY P., and NEIL S. DAVIES. "GRAIN-SIZE CONTROLS ON THE SILURO-DEVONIAN COLONIZATION OF NON-MARINE SUBSTRATES BY INFAUNAL INVERTEBRATES." PALAIOS 37, no. 12 (2022): 731–43. http://dx.doi.org/10.2110/palo.2021.069.

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ABSTRACT Throughout the history of life on Earth, sedimentary environments have placed controls on the trajectory of evolutionary innovations. To survive and thrive in newly colonized sedimentary environments, organisms have needed to develop novel behaviors: often evidenced in the rock record as architectural innovation and diversification in trace fossil morphology. This study focuses on ichnological diversification as a response to challenges presented by different sediment grain sizes during the late Silurian to Early Devonian colonization of the continents by invertebrate life. The ichnod
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10

KIKUCHI, KAZUKI, HAJIME NARUSE, and NOBUHIRO KOTAKE. "EVALUATION OF ICHNODIVERSITY BY IMAGE-RESAMPLING METHOD TO CORRECT OUTCROP EXPOSURE BIAS." PALAIOS 33, no. 5 (2018): 204–17. http://dx.doi.org/10.2110/palo.2017.090.

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11

Bromley, Richard G. "Trace fossils from the Lower and Middle Jurassic marginal marine deposits of the Sorthat Formation, Bornholm, Denmark." Bulletin of the Geological Society of Denmark 50 (December 18, 2003): 185–208. http://dx.doi.org/10.37570/bgsd-2003-50-15.

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The Lower to Middle Jurassic Sorthat and Bagå Formations of the Baltic island of Bornholm, Denmark, are a predominantly fluviatile unit. On the south coast of the island at Korsodde, however, an interval within the Sorthat Formation contains a diverse trace fossil assemblage indicating a marine incursion. Study of this interval revealed 15 ichnotaxa, among which one is new: Bornichnus tortuosus nov. igen. et isp. Several of the trace fossils present are generally considered characteristic of the lower shoreface to offshore environments (e.g. Teichichnus and Asterosoma). However, the low degree
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12

Buatois, Luis A., M. Gabriela Mángano, Nicholas J. Minter, et al. "Quantifying ecospace utilization and ecosystem engineering during the early Phanerozoic—The role of bioturbation and bioerosion." Science Advances 6, no. 33 (2020): eabb0618. http://dx.doi.org/10.1126/sciadv.abb0618.

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The Cambrian explosion (CE) and the great Ordovician biodiversification event (GOBE) are the two most important radiations in Paleozoic oceans. We quantify the role of bioturbation and bioerosion in ecospace utilization and ecosystem engineering using information from 1367 stratigraphic units. An increase in all diversity metrics is demonstrated for the Ediacaran-Cambrian transition, followed by a decrease in most values during the middle to late Cambrian, and by a more modest increase during the Ordovician. A marked increase in ichnodiversity and ichnodisparity of bioturbation is shown during
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13

Arifullah, Ery, Yahdi Zaim, Aswan Aswan, and Djuhaeni Djuhaeni. "Basin-scale Paleoecology: Using Semi-quantitative Analysis of the Ichnofabric within Kutai Basin (Indonesia)." Journal of Mathematical and Fundamental Sciences 53, no. 2 (2021): 285–304. http://dx.doi.org/10.5614/j.math.fund.sci.2021.53.2.8.

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Ichnofossils are still not used in paleoecological studies, even though they are a valuable proxy for paleoecology. This study focused on a semi-quantitative approach to a number of ichnofabric variables, i.e. ichnofossil association, bioturbation index (BI), ichnodiversity (ID), number of behaviors (NB), penetration depth (PD), and burrow diameter (DM). It was proved that the scores of those variables were low to medium because of the paleoecological fluvial-marine transition depositional processes in the Serravallian-Tortonian interval in Kutai Basin, Indonesia. This paper contributes an ich
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14

Minter, Nicholas J., Spencer G. Lucas, Allan J. Lerner, and Simon J. Braddy. "Augerinoichnus Helicoidalis, a new helical trace fossil from the nonmarine Permian of New Mexico." Journal of Paleontology 82, no. 6 (2008): 1201–6. http://dx.doi.org/10.1017/s0022336000055402.

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New Mexico contains a significant record of trace fossil assemblages, in terms of both abundance and ichnodiversity, from Lower Permian nonmarine depositional settings. Most notable amongst these are the trace fossil assemblages in the Robledo Mountains Formation of the Robledo Mountains in Doña Ana County, southern New Mexico, as recognized by the recent proposal to designate this area as a national monument. These trace fossil assemblages formed on a tidal flat under largely non-marine conditions (Mack and James, 1986; Hunt et al., 1993; Lucas et al., 1995a, 1998) and are dominated by the tr
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15

Arifullah, Ery, Yahdi Zaim, Aswan Aswan, and Djuhaeni Djuhaeni. "Basin-scale Paleoecology: Using Semi-quantitative Analysis of the Ichnofabric within Kutai Basin (Indonesia)." Journal of Mathematical and Fundamental Sciences 53, no. 2 (2021): 286–305. https://doi.org/10.5614/j.math.fund.sci.2021.53.2.8.

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Ichnofossils are still not used in paleoecological studies, even though they are a valuable proxy for paleoecology. This study focused on a semi-quantitative approach to a number of ichnofabric variables, i.e. ichnofossil association, bioturbation index (BI), ichnodiversity (ID), number of behaviors (NB), penetration depth (PD), and burrow diameter (DM). It was proved that the scores of those variables were low to medium because of the paleoecological fluvial-marine transition depositional processes in the Serravallian-Tortonian interval in Kutai Basin, Indonesia. This paper contributes an ich
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16

Naimi, Mohammed Nadir, and Amine Cherif. "Sedimentology and ichnology of the mid-Cretaceous succession of Ouled Nail Mounts (Eastern Saharan Atlas, Algeria)." Geologia Croatica 74, no. 3 (2021): 209–23. http://dx.doi.org/10.4154/gc.2021.15.

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Shallow marine deposits characterize the upper Albian – lower Cenomanian deposits of Northern Algeria. In Djebel Azzeddine (Ouled Nail Mounts), the corresponding sediments have been subdivided into three distinctive units A to C. The first discovered ammonite fauna from the Bou Saada area allowed the attribution of a part of the mid-Cretaceous post-Continental Intercalaire deposits to the upper Albian. The ammonite-bearing level indicates a maximum flooding surface and could be correlated with similar levels from Northern Algeria. The studied succession is characterized by a low ichnodiversity
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17

Bayet-Goll, Aram, Carlos Neto De Carvalho, Mohammad Hossein Mahmudy-Gharaei та Reza Nadaf. "Ichnology and sedimentology of a shallow marine Upper Cretaceous depositional system (Neyzar Formation, Kopet-Dagh, Iran): Palaeoceanographic influence on ichnodiversity". Cretaceous Research 56 (вересень 2015): 628–46. http://dx.doi.org/10.1016/j.cretres.2015.07.008.

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18

Tchoumatchenco, Platon, and Alfred Uchman. "Lower and Middle Jurassic flysch trace fossils from the eastern Stara Planina Mountains, Bulgaria: A contribution to the evolution of Mesozoic ichnodiversity." Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 213, no. 2 (1999): 169–99. http://dx.doi.org/10.1127/njgpa/213/1999/169.

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19

Miguez-Salas, Olmo, and Francisco J. RodrÍguez-Tovar. "Ichnofabric analysis as a tool for characterization and differentiation between calcareous contourites and calciturbidites." Journal of Sedimentary Research 91, no. 11 (2021): 1151–65. http://dx.doi.org/10.2110/jsr.2020.147.

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ABSTRACT The Eocene–Miocene Cyprus paleoslope system records complex deep-marine sedimentation comprising background vertical settling of autochthonous pelagic–hemipelagic particles (chalks) which were punctuated by calcareous bottom currents (contourites) and gravity flows (calciturbidites). The Eocene Lefkara Formation at the Petra Tou Romiou beach section (Cyprus) shows the incidence of deep-marine bottom currents and distal turbiditic episodes in a context of pelagic–hemipelagic sedimentation. Trace-fossil analysis of this section, using an ichnofabric approach (i.e., ichnodiversity, Biotu
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20

Mángano, M. Gabriela, Christopher David Hawkes, and Jean-Bernard Caron. "Trace fossils associated with Burgess Shale non-biomineralized carapaces: bringing taphonomic and ecological controls into focus." Royal Society Open Science 6, no. 1 (2019): 172074. http://dx.doi.org/10.1098/rsos.172074.

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The association of trace fossils and non-biomineralized carapaces has been reported from Cambrian Lagerstätten worldwide, but the abundance, ichnodiversity, taphonomy and ecological significance of such associations have yet to be fully investigated. Two main end-member hypotheses are explored based on the study of a relatively wide variety of trace fossils preserved associated to Tuzoia carapaces from the middle Cambrian Burgess Shale in British Columbia. In the ecological Tuzoia garden hypothesis, the bacterially enriched surface of carapaces provides opportunities for intricate ecologic int
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21

Vinn, Olev, Mehdi Hadi, Dangpeng Xi, Majid Mirzaie Ataabadi, and Francisco J. Vega. "Paleoecology, bioerosion, and paleoenvironments in the Late Miocene of Dar Pahn Unit, Makran, SE Iran." Revista Brasileira de Paleontologia 27, no. 3 (2024): e20240469. http://dx.doi.org/10.4072/rbp.2024.3.0469.

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This study focuses on sedimentation patterns and paleoecological conditions during the Upper Miocene in Southeast Iran, specifically analyzing the Rudig section. The sedimentation was predominantly terrigenous and occurred in marine conditions, forming multiple shallowing upwards sequences comprising silty marl, siltstone, and sandstone. The analysis of molluscan shells from this period indicates minimal bioerosion, with encrusters represented solely by Balanus sp. This low level of encrustation suggests rapid sedimentation and brief exposure times. Bioerosion was limited, with Caulostrepsis t
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Davies, Neil S., Anthony P. Shillito, and Cameron R. Penn-Clarke. "Cold feet: Trackways and burrows in ice-marginal strata of the end-Ordovician glaciation (Table Mountain Group, South Africa)." Geology 48, no. 12 (2020): 1159–63. http://dx.doi.org/10.1130/g47808.1.

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Abstract New observations from an outcrop of Upper Ordovician Table Mountain Group strata (Matjiesgoedkloof, Western Cape Province, South Africa) have revealed an unexpected ichnofauna that is hosted within diamictites and sandstones that were deposited by a retreating low-latitude (∼30°S) ice sheet during the Hirnantian glaciation. The locality provides a rare window onto animal-sediment interactions in an early Paleozoic ice-marginal shallow-marine environment and contains a trace fossil community with a surprising ichnodiversity and ichnodisparity of burrows, trackways, and trails (Archaeon
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23

Shahkarami, Setareh, M. Gabriela Mángano, and Luis A. Buatois. "Ichnostratigraphy of the Ediacaran-Cambrian boundary: new insights on lower Cambrian biozonations from the Soltanieh Formation of northern Iran." Journal of Paleontology 91, no. 6 (2017): 1178–98. http://dx.doi.org/10.1017/jpa.2017.72.

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AbstractStrata in the Central Alborz Mountains, northern Iran, are interpreted to show continuous sedimentation from Ediacaran through Cambrian times. The Soltanieh Formation consists of five members: Lower Dolomite, Lower Shale, Middle Dolomite, Upper Shale and Upper Dolomite members. The clastic units (Lower and Upper Shale members) represent sedimentation in distal marine settings, ranging from the shelf to offshore, and contain abundant trace fossils of biostratigraphic utility. Four ichnozones have been recognized. Ichnozone 1, containingHelminthoidichnites tenuis,Helminthopsis tenuis, an
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Dalman, Sebastian G., and Robert E. Weems. "A New Look at Morphological Variation in the IchnogenusAnomoepus, with Special Reference to Material from the Lower Jurassic Newark Supergroup: Implications for Ichnotaxonomy and Ichnodiversity." Bulletin of the Peabody Museum of Natural History 54, no. 1 (2013): 67–124. http://dx.doi.org/10.3374/014.054.0104.

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25

Schatz, Elizabeth R., María Gabriela Mángano, Luis A. Buatois, and Carlos Oscar Limarino. "Life in the Late Paleozoic ice age: trace fossils from glacially influenced deposits in a Late Carboniferous fjord of western Argentina." Journal of Paleontology 85, no. 3 (2011): 502–18. http://dx.doi.org/10.1666/10-046.1.

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The early Late Carboniferous rocks of the Guandacol Formation in western Argentina preserve the glacial to postglacial transition. In the study area, this unit has been divided in three intervals: 1) a lower diamictitic interval; 2) a middle interval chiefly composed of mudstone, and 3) an upper sandstone-dominated interval. The lower interval records infill of a fjord incised into the underlying Ordovician limestone. The middle and upper intervals reflect postglacial sedimentation. Four ichnotaxa, occurring as both discrete and compound trace fossils, are documented from the lower and middle
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Martins, Taynara Cristina Matos, Joelson Lima Soares, and Hudson Pereira Santos. "Icnofósseis no Arenito do Guamá (Pará, Brasil): potenciais registros silurianos." Revista Brasileira de Paleontologia 25, no. 2 (2022): 103–19. http://dx.doi.org/10.4072/rbp.2022.2.01.

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Ichnofossils of the Guamá Sandstone (Pará, Brazil): potential Silurian records. The Guamá Sandstone is a unit restricted to the Bragantina Platform in the northeast Pará State. It is characterized by thick layers of well-rounded, well-sorted, medium-grained quartz-sandstone with textural and compositional maturity. In general, the layers exhibit few preserved primary sedimentary structures. The facies association indicates a sandy coastal paleoenvironment comprising the foreshore and shoreface zones. We found in the Guamá Sandstone the ichnospecies Psammichnites isp., cf. Schaubcylindrichnus c
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Mángano, M. Gabriela, and Luis A. Buatois. "The rise and early evolution of animals: where do we stand from a trace-fossil perspective?" Interface Focus 10, no. 4 (2020): 20190103. http://dx.doi.org/10.1098/rsfs.2019.0103.

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The trace-fossil record provides a wealth of information to track the rise and early evolution of animals. It comprises the activity of both hard- and soft-bodied organisms, is continuous through the Ediacaran (635–539 Ma)– Cambrian (539–485 Ma) transition, yields insights into animal behaviour and their role as ecosystem engineers, and allows for a more refined characterization of palaeoenvironmental context. In order to unravel macroevolutionary signals from the trace-fossil record, a variety of approaches is available, including not only estimation of degree of bioturbation, but also analys
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Mangano, Gabriela M., Luis A. Buatois, Ronald R. West, and Christopher G. Maples. "Ichnology of a Pennsylvanian Equatorial Tidal Flat -- The Stull Shale Member at Waverly, Eastern Kansas." Bulletin (Kansas Geological Survey), no. 245 (January 1, 2002): 1–139. https://doi.org/10.17161/kgsbulletin.no.i245.22279.

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Integrated stratigraphic, sedimentologic, and ichnologic analyses indicate that the Stull Shale Member (Kanwaka Shale, Shawnee Group) at Waverly (Coffey County, eastern Kansas) was deposited along a microtidal shoreline directly connected with the open sea and outside of a northeast-south west-oriented embayment. Six major subenvironments have been identified in the lower interval of the Waverly section: sand flat, mixed flat, mud flat, supratidal paleosols, intertidal runoff channels, and fluvial channel. Paleogeographic information indicates that the position of the Waverly tidal flat was eq
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Veenma, Yorick P., Neil S. Davies, and Anthony P. Shillito. "The ichnological optimum: Enhanced trace fossil diversity in the Cambrian offshore driven by ecology, sedimentation, and outcrop." Journal of the Geological Society, April 8, 2025. https://doi.org/10.1144/jgs2024-220.

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The trace fossil record provides a valuable archive of the Cambrian Explosion and its variable expressions at the local level. To characterize factors controlling such variability, we present a framework that deconstructs local ichnodiversity into 1) ‘ original ichnodiversity’, reflecting ecology and behavioural diversity; 2) ‘ registered ichnodiversity’, reflecting colonization windows and substrate preservation; and 3) ‘ observable ichnodiversity’, reflecting present-day outcrop quality. An environment with favourable controls on all three levels results in an ichnological optimum , exemplif
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Zhang, Li‐Jun, Luis A. Buatois, M. Gabriela Mángano, et al. "Trace fossils from the Meishucun section of South China: revisiting ichnotaxonomy, behavioural diversification and ecosystem engineering from a key Ediacaran–Cambrian succession." Papers in Palaeontology 11, no. 2 (2025). https://doi.org/10.1002/spp2.70009.

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AbstractThe Ediacaran–Cambrian transition was a time of profound geobiological revolution and sedimentary change in Earth's history, including arguably the most dramatic evolutionary radiation and a remarkable biotic replacement. The complex feedback between diversification of metazoans and their ecological interactions, especially after the extinction of the latest Ediacaran biota, is still debated. Here, we systematically studied the trace fossil content of the Ediacaran–Cambrian succession from the Meishucun section of South China. A total of 20 ichnogenera comprising 31 ichnospecies have b
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Meyer, Neele, Max Wisshak, and André Freiwald. "Ichnodiversity and bathymetric range of microbioerosion traces in polar barnacles of Svalbard." Polar Research 39 (September 14, 2020). http://dx.doi.org/10.33265/polar.v39.3766.

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Valencia, Fernando L., M. Gabriela Mángano, Luis A. Buatois, and Juan Carlos Laya. "Animal–substrate interactions preserved in ancient lagoonal chalk." Scientific Reports 12, no. 1 (2022). http://dx.doi.org/10.1038/s41598-022-18713-8.

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AbstractTrace-fossil assemblages reflect the response of the benthos to sets of paleoenvironmental conditions during and immediately after sedimentation. Trace fossils have been widely studied in pelagic shelf and deep-sea chalk deposits from around the globe but never documented from ancient lagoonal chalk successions. Here we report the first detailed ichnologic analysis of a lagoonal chalk unit, using as an example the Upper Cretaceous Buda Formation from the Texas Gulf Coast Basin. In this unit, variable interconnection with the open ocean, accompanied by marked fluctuations in physicochem
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Gougeon, Romain, M. Gabriela Mángano, Luis A. Buatois, Guy M. Narbonne, Brittany A. Laing, and Maximiliano Paz. "The Ediacaran–Cambrian Chapel Island Formation of Newfoundland, Canada: evaluating the impact of outcrop quality on trace-fossil data sets at the Cambrian GSSP and less-explored sections." Canadian Journal of Earth Sciences, December 23, 2022. http://dx.doi.org/10.1139/cjes-2022-0060.

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In 1992, the Chapel Island Formation at Fortune Head was selected as the Cambrian GSSP, which was placed at the first appearance of the ichnotaxon Treptichnus pedum. Although the transition from the Ediacaran to the Cambrian was well studied in Fortune Head and Grand Bank Head, it is also exposed at Lewin’s Cove and Point May. Here, we report new ichnologic and sedimentologic data sets from the four localities. Ichnologic data sets consist of bioturbation intensities, burrow width and depth, ichnotaxonomic composition, and stratigraphic appearances. Sedimentologic data sets include sediment gr
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Meyer, Neele, Max Wisshak, Evan N. Edinger, Kumiko Azetsu-Scott, and André Freiwald. "Ichnodiversity in the eastern Canadian Arctic in the context of polar microbioerosion patterns." Polar Research 41 (April 29, 2022). http://dx.doi.org/10.33265/polar.v41.8083.

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Studies of marine microbioerosion in polar environments are scarce. They include our recent investigations of bioerosion traces preserved in sessile balanid skeletons from the Arctic Svalbard archipelago and the Antarctic Ross Sea. Here, we present results from a third study site, Frobisher Bay, in the eastern Canadian Arctic, together with a synthesis of our current knowledge of polar bioerosion in both hemispheres. Barnacles from 62 to 94 m water depth in Frobisher Bay were prepared using the cast-embedding technique to enable visualization of microboring traces by scanning electron microsco
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Wisshak, Max. "Taming an ichnotaxonomical Pandora's box: revision of dendritic and rosetted microborings (ichnofamily: Dendrinidae)." European Journal of Taxonomy, no. 390 (December 29, 2017). http://dx.doi.org/10.5852/ejt.2017.390.

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Dendritic and/or rosetted microborings in calcareous and osteic skeletal substrates have a diverse trace fossil record, spanning most of the Phanerozoic, whereas the ichnodiversity of comparable bioerosion traces produced in modern seas is rather limited. The most prominent occurrences are known from Devonian brachiopods and from Upper Cretaceous belemnite rostra. Ichnotaxonomically, they are comprised within one of the few ichnofamilies established to date, the Dendrinidae Bromley et al., 2007. As an outcome of the present revision of this ichnofamily, the plethora of 84 ichnospecies establis
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Goers, Alexa R., Stephen T. Hasiotis, and Eugene C. Rankey. "Controls on bioturbation and sediment distribution in carbonate shoreface deposits: insights from heterogeneity in Pleistocene and recent strata." Journal of Sedimentary Research, August 11, 2023. http://dx.doi.org/10.2110/jsr.2022.024.

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Burrowing organisms alter sedimentary textures, influence cement distribution, and affect petrophysical characteristics of carbonate strata. Although many descriptions of carbonate successions reference bioturbation, quantitative data on spatial variability of trace fossils is rare, and fewer studies address trace-fossil influence on postdepositional modification of sedimentary deposits, which can affect petrophysical properties. To address these unknowns and determine the controls on ichnology in carbonate shoreface successions, this study evaluates the along- and across-strike distribution o
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Tindal, Benjamin H., Anthony P. Shillito, and Neil S. Davies. "First report of fish trace fossils (Undichna) from the Middle Devonian Achanarras Limestone, Caithness Flagstone Group." Scottish Journal of Geology, August 10, 2021, sjg2020–023. http://dx.doi.org/10.1144/sjg2020-023.

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Two newly-discovered specimens of the fish locomotion trace Undichna (U. britannica and Undichna isp.), are described from the Middle Devonian Achanarras Limestone Member (Caithness Flagstone Group, NE Scotland). Fish trace fossils have not previously been reported from the Achanarras Limestone Member, despite decades of study of the unit as a key locality for fish body fossils. The traces comprise discontinuous sinusoidal grooves; one showing multiple parallel incisions, created by the fins of an acanthodian fish swimming close to the substrate. The apparent absence of trace fossils attributa
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38

Castanera, Diego, Elisabete Malafaia, Bruno C. Silva, Vanda F. Santos, and Matteo Belvedere. "New dinosaur, crocodylomorph and swim tracks from the Late Jurassic of the Lusitanian Basin: implications for ichnodiversity." Lethaia, November 4, 2020. http://dx.doi.org/10.1111/let.12402.

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39

Meek, Dean M., Luis A. Buatois, M. Gabriela Mángano, and Bruce M. Eglington. "Increased habitat segregation at the dawn of the Phanerozoic revealed by correspondence analysis of bioturbation." Scientific Reports 13, no. 1 (2023). http://dx.doi.org/10.1038/s41598-023-49716-8.

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AbstractThe Agronomic Revolution of the early Cambrian refers to the most significant re-structuration of the benthic marine ecosystem in life history. Using a global compilation of trace-fossil records across the Ediacaran-Cambrian transition, this paper investigates the relationship between the benthos and depositional environments prior to, during, and after the Agronomic Revolution to shed light on habitat segregation via correspondence analysis. The results of this analysis characterize Ediacaran mobile benthic bilaterians as facies-crossing and opportunistic, with low levels of habitat s
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40

Zouheir, Tariq, Abdelkbir Hminna, Hafid Saber, et al. "Ichnodiversity and facies of Triassic red beds in the Irohalene area (Argana Basin, Western High Atlas, Morocco): implications for palaeoenvironment." Historical Biology, June 19, 2022, 1–34. http://dx.doi.org/10.1080/08912963.2022.2069018.

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41

Lockley, Martin G., Charles W. Helm, Andrew M. W. Lawfield, and Kevin J. Sharman. "New evidence for avian and small non-avian theropod ichnotaxa from the Lower Cretaceous of Canada: implications for theropod ichnodiversity." Cretaceous Research, June 2022, 105292. http://dx.doi.org/10.1016/j.cretres.2022.105292.

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42

Bayet-Goll, Aram, Mehdi Daraei, Gerd Geyer, Carlos Neto de Carvalho, and Nasrin Bahrami. "Exotic facies episodes of a carbonate platform: implications for middle and late Cambrian ecosystems and impact of bioturbation in the Alborz Basin, Iran." Geological Society, London, Special Publications, November 23, 2021, SP522–2020–269. http://dx.doi.org/10.1144/sp522-2020-269.

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AbstractThe sedimentological and ichnological data of the Mila Group in the Alborz Basin, northern Iran indicate that the appearance of exotic facies and resurgence of microbialites resulted from the development of extraordinary palaeoceanographic conditions with widespread environmental stress and enhanced precipitation of CaCO3. Inhospitable environmental conditions in the Mila Group platform led to a considerable reduction in ichnodiversity, bioturbation intensity and depth and size of burrows, which led to periods of dominating calcimicrobe ecology. The development of exotic or anachronist
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43

Veenma, Yorick P., and Neil S. Davies. "Short-stasis signatures in Cambrian and Ordovician shallow-marine sandstones: Implications for the ichnological record and time preserved at outcrop." Geological Society, London, Special Publications 556, no. 1 (2024). http://dx.doi.org/10.1144/sp556-2024-96.

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Shallow-marine sandstones are an archetypal facies in the stratigraphic record of the Cambrian Explosion and the Ordovician Radiation but have been considered uninformative due to erosional signatures, time-incompleteness, and unfossiliferous outcrops. New analyses of the Carnedd-y-Filliast Grits (Furongian, Wales) and littoral sandstone successions from Laurentia and Baltica reveal that bedding planes in such outcrops commonly register sedimentary stasis. Most stasis intervals were brief, reflecting 1) a high sedimentation frequency; 2) post-depositional substrate resculpting by wave action d
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Enriquez, Nathan J., Nicolás E. Campione, Corwin Sullivan, et al. "Probable deinonychosaur tracks from the Upper Cretaceous Wapiti Formation (upper Campanian) of Alberta, Canada." Geological Magazine, December 23, 2020, 1–14. http://dx.doi.org/10.1017/s0016756820001247.

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Abstract Late Cretaceous tracks attributable to deinonychosaurs in North America are rare, with only one occurrence of Menglongipus from Alaska and two possible, but indeterminate, occurrences reported from Mexico. Here we describe the first probable deinonychosaur tracks from Canada: a possible trackway and one isolated track on a single horizon from the Upper Cretaceous Wapiti Formation (upper Campanian) near Grande Prairie in Alberta. The presence of a relatively short digit IV differentiates these from argued dromaeosaurid tracks, suggesting the trackmaker was more likely a troodontid. Oth
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Miguez-Salas, Olmo, Francisco J. Rodríguez-Tovar, and Wouter de Weger. "The Late Miocene Rifian corridor as a natural laboratory to explore a case of ichnofacies distribution in ancient gateways." Scientific Reports 11, no. 1 (2021). http://dx.doi.org/10.1038/s41598-021-83820-x.

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AbstractOceanic gateways have modulated ocean circulation and have influenced climatic variations throughout the Earth´s history. During the late Miocene (7.8–7.35 Ma), the Atlantic Ocean and the Mediterranean Sea were connected through the Rifian Corridor (Morocco). This gateway is one of the few examples of deep ancient seaways with a semi-continuous sedimentary record. Deposits comprise turbidites intercalated between deep-sea mudstone (i.e., hemipelagites and drift deposits), channelized sandstone contourite facies, and shallow marine sandstone. Herein an ichnological analysis was conducte
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Oligmueller, Andrew R., and Stephen T. Hasiotis. "An ichnotaxonomic assessment of the Cretaceous Dakota Group, Front Range, Colorado, USA, and its comparison to other Western interior seaway deposits." Paleontological Contributions, no. 23 (July 23, 2024). http://dx.doi.org/10.17161/pc.vi23.22542.

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The Aptian-Cenomanian Dakota Group along the Colorado Front Range is known for its dinosaur tracks; however, it also contains extensive invertebrate marine and continental trace fossils. The Dakota Group in Colorado is subdivided into the Lytle, Plainview, Glencairn (Cañon City), Skull Creek Shale (Denver, Fort Collins), and Muddy formations. Thirty-two invertebrate ichnogenera and 34 ichnospecies were identified: Archaeonassa, Arenicolites, Asterosoma, Asthenopodichnium, Aulichnites, Chondrites, Cochlichnus, Conichnus, Cruziana, Cylindrichnus, Diplocraterion, Gyrolithes, Lockeia, Macaronichnu
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Gesualdi, Vincenzo, Matteo Belvedere, Marko Yurac, et al. "Diverse dinosaur tracks from the Upper Jurassic – Lower Cretaceous Chacarilla Formation of Quebrada de Arcas, northeast Chile: Evidence of high ichnodiversity in an arid palaeoenviroment." Palaeogeography, Palaeoclimatology, Palaeoecology, June 2025, 113088. https://doi.org/10.1016/j.palaeo.2025.113088.

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48

Bayet-Goll, Aram. "Ordovician matground and mixground ecosystems in shoreface–offshore and barrier-island environments from Central Iran, northern Gondwana." Geological Magazine, April 11, 2022, 1–29. http://dx.doi.org/10.1017/s0016756822000097.

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Abstract The shift of matground ecosystems to bioturbator-free settings was investigated in the Ordovician wave-dominated marine strata of the Shirgesht Formation in Central Iran. Ten ichnofabrics are recognized in shoreface–offshore and barrier-island sedimentary facies, representing a proximal-to-distal depositional trend along the studied profile. In the offshore settings, intensive burrowing on several tiers and bioirrigation, referred to the Thalassinoides (Th), Cruziana–Skolithos–Rosselia (CrSkRo), crowded Trichophycus (CT) and Cruziana–Helminthopsis (CrHe) ichnofabrics, prevented the de
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49

Wang, Zekun, Imran A. Rahman, and Li-Jun Zhang. "Quantifying the rise of animals during the Ediacaran–Cambrian using ichnodissimilarity." Paleobiology, December 10, 2024, 1–16. https://doi.org/10.1017/pab.2024.40.

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Abstract The trace fossil record provides important insights into the evolution of early animals during the Ediacaran/Cambrian transition, with changes in ichnodiversity through time and between environments informing on the diversification of major body plans, behaviors, and niches. To quantify variation in the diversity of trace fossils across this critical interval, we propose a measure of trace fossil dissimilarity (ichnodissimilarity) based on vector calculation. Furthermore, by comparing discrepancies between the angular bisector and mean vector of two sets of vectorized fossil data, we
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Choudhary, Anjali, Suruchi Chauhan, Bhaskar Paul, Dhruv Patel, and Bhawanisingh Desai. "Ichnoguild analysis of the Bajocian-Bathonian Yellow Flagstone Member (Goradongar Formation) from the Kachchh Basin, Western India." Journal of the Palaeontological Society of India, October 22, 2024. http://dx.doi.org/10.1177/05529360241286278.

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The Yellow Flagstone is the oldest member of the Goradongar Formation exposed in the Pachchham Island, Kachchh Basin. The members exposed in four sections are mapped for trace fossils, and 17 ichnofossils have been documented. The concept of ichnoguild (IG) is applied to reveal the infaunal tiering structure and the strategies organisms adapt to exploit the resources. Ten IG of similar characteristics and ecological functions have been identified. Five are shallow-tier IGs ( Asterosoma IG, Gyrochorte IG, Nereites IG, Planolites IG and Bolonia IG), three middle-tier IGs ( Rhizocorallium IG, Are
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