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1

Preński, Leszek Bruno. "Studies on hake, Merluccius capensis Castelnau, 1861, from off Namibia. II. Studies on food and feeding." Acta Ichthyologica et Piscatoria 16, no. 2 (December 31, 1986): 3–21. http://dx.doi.org/10.3750/aip1986.16.2.01.

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2

Macpherson, E., and A. Gordoa. "Effect of prey densities on cannibalism in Cape hake (Merluccius capensis) off Namibia." Marine Biology 119, no. 1 (April 1994): 145–49. http://dx.doi.org/10.1007/bf00350116.

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3

Caputo, M., S. Elwen, T. Gridley, SA Kohler, JP Roux, PW Froneman, and JJ Kiszka. "Dietary plasticity of two coastal dolphin species in the Benguela upwelling ecosystem." Marine Ecology Progress Series 669 (July 8, 2021): 227–40. http://dx.doi.org/10.3354/meps13755.

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Defining the trophic relationships of marine predators and their dietary preferences is essential in understanding their role and importance in ecosystems. Here we used stable isotope analysis of skin samples (δ15N values reflecting trophic level and δ13C values reflecting foraging habitat) to investigate resource partitioning and spatial differences of the feeding ecology of dusky dolphins Lagenorhynchus obscurus and Heaviside’s dolphins Cephalorhynchus heavisidii from 2 coastal study sites separated by 400 km along the coast of central (Walvis Bay) and southern (Lüderitz) Namibia in the Benguela upwelling ecosystem. Overall, isotopic niches of both predators were significantly different, indicating partitioning of resources and foraging habitats. Despite their smaller body size, Heaviside’s dolphins fed at a significantly higher trophic level than dusky dolphins. Stable isotope mixing models revealed that both species fed on high trophic level prey (i.e. large Merluccius spp., large Sufflogobius bibarbatus, and Trachurus t. capensis) at Walvis Bay. The diet of both dolphin species included smaller pelagic fish and squid at Lüderitz. Spatial differences highlight that Heaviside’s and dusky dolphins may exhibit dietary plasticity driven by prey availability, and that they likely form distinct population segments. Important prey for both dolphin species, specifically Merluccius spp. and T. t. capensis, are the main target of trawl fisheries in the Benguela upwelling ecosystem, highlighting potential resource overlap between dolphins and fisheries.
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4

Gordoa, Ana, and Joseph E. Hightower. "Changes in Catchability in a Bottom-Trawl Fishery for Cape Hake (Merluccius capensis)." Canadian Journal of Fisheries and Aquatic Sciences 48, no. 10 (October 1, 1991): 1887–95. http://dx.doi.org/10.1139/f91-224.

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Management of the Cape hake (Merluccius capensis) stock off the coasts of Angola and Namibia has been based on surplus production modeling, which incorporates an assumption of constant catchability. Results of least-squares catch-at-age analysis and virtual population analysis were used to estimate catchability from 1968 to 1986. Estimated catchability was either relatively constant or gradually increasing from 1968 to 1978, but fluctuated considerably between about 1979 and 1983. The fluctuations may have been due to an inverse relationship between catchability and abundance or to environmental factors affecting fish distribution. The period of highest estimated catchability corresponded to the period of lowest estimated abundance and below-average water temperatures. If management is based on the assumption of constant catchability but catchability is inversely related to abundance, the risk of overfishing will be greatest when the stock is at the lowest levels. Future assessments should place increased reliance on survey data or incorporate an appropriate model of catchability changes.
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5

Wilhelm, MR, SC Paulus, and S. Kashava. "New age-determination method based on fast growth rate of Cape hake Merluccius capensis in Namibia." African Journal of Marine Science 41, no. 2 (April 3, 2019): 119–24. http://dx.doi.org/10.2989/1814232x.2019.1581659.

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6

Kadila, Hendrina K., Dietlinde N. Nakwaya, Mike Butler, and Johannes A. Iitembu. "Insights into feeding interactions of shallow water cape hake (Merluccius capensis) and cape horse mackerel (Trachurus capensis) from the Northern Benguela (Namibia)." Regional Studies in Marine Science 34 (February 2020): 101071. http://dx.doi.org/10.1016/j.rsma.2020.101071.

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7

REED, CECILE, CHAGANTI KALAVATI, KEN MACKENZIE, CATHERINE COLLINS, and WILLY HEMMINGSEN. "Pseudalataspora vanderlingeni n. sp. (Myxosporea: Bivalvulida) from gall bladders of the Cape hakes Merluccius capensis Castelnau, and M. paradoxus Franca (Teleostei: Merlucciidae)." Zootaxa 4497, no. 3 (October 9, 2018): 422. http://dx.doi.org/10.11646/zootaxa.4497.3.6.

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A new species of myxosporean parasite is described from the gall bladders of the hakes Merluccius capensis Castelnau and M. paradoxus Franca (Pisces: Teleostei) caught off the west and south coasts of South Africa. The new species, Pseudalataspora vanderlingeni, is described morphologically and molecularly and compared with the 15 other species of Pseudalataspora previously described from marine fish. Although a molecular description is available on GenBank for only one of these 15 species, the morphological description supports the status of P. vanderlingeni as a new species. Earlier reports, without detailed descriptions, of Leptotheca sp. and Ceratomyxa sp. from the same hosts caught off Namibia were very likely to have been of P. vanderlingeni. These earlier studies reported high prevalences of infection, similar to those of >60% described in the present study. The effects of fixation and freezing on the dimensions of spores of Pseudalataspora spp. are described, and the status of the genus Pseudalataspora within the family Ceratomyxidae is discussed.
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8

Voges, Elizabeth, Ana Gordoa, Chris Hans Bartholomae, and John G. Field. "Estimating the probability of different levels of recruitment for Cape hakes Merluccius capensis off Namibia, using environmental indices." Fisheries Research 58, no. 3 (November 2002): 333–40. http://dx.doi.org/10.1016/s0165-7836(01)00400-3.

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9

IITEMBU, JOHANNES A., TODD W. MILLER, KOJI OHMORI, ABRAHAM KANIME, and SEAN WELLS. "Comparison of ontogenetic trophic shift in two hake species, Merluccius capensis and Merluccius paradoxus, from the Northern Benguela Current ecosystem (Namibia) using stable isotope analysis." Fisheries Oceanography 21, no. 2-3 (February 13, 2012): 215–25. http://dx.doi.org/10.1111/j.1365-2419.2012.00614.x.

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10

Gordoa, A., and E. Macpherson. "Diurnal variation in the feeding activity and catch rate of cape hake (Merluccius capensis and M. paradoxus) off Namibia." Fisheries Research 12, no. 4 (December 1991): 299–305. http://dx.doi.org/10.1016/0165-7836(91)90014-7.

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11

Thygesen, Uffe Høgsbro, Kasper Kristensen, Teunis Jansen, and Jan E. Beyer. "Intercalibration of survey methods using paired fishing operations and log-Gaussian Cox processes." ICES Journal of Marine Science 76, no. 4 (January 8, 2019): 1189–99. http://dx.doi.org/10.1093/icesjms/fsy191.

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Abstract We present a statistical method for intercalibration of fishery surveys methods, i.e. determining the difference in catchability and size selectivity of two methods, such as trawl gears or vessels, based on data from paired fishing operations. The model estimates the selectivity ratios in each length class by modelling the size distribution of the underlying population at each station and the size-structured clustering of fish at small temporal and spatial scales. The model allows for overdispersion and correlation between catch counts in neighbouring size classes. This is obtained by assuming Poisson-distributed catch numbers conditional on unobserved log-Gaussian variables, i.e. the catch is modelled using log-Gaussian Cox processes. We apply the method to catches of hake (Merluccius paradoxus and M. capensis) in 341 paired trawl hauls performed by two different vessels, viz. the RV Dr Fridtjof Nansen and the FV Blue Sea, operating off the coast of Namibia. The results demonstrate that it is feasible to estimate the selectivity ratio in each size class, and to test statistically the hypothesis that the selectivity is independent of size or species. For the specific case, we find that differences between size classes and species are statistically significant.
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12

Iitembu, Johannes A., and Nicole B. Richoux. "Trophic relationships of hake (Merluccius capensis and M. paradoxus) and sharks (Centrophorus squamosus, Deania calcea and D. profundorum) in the Northern (Namibia) Benguela Current region." African Zoology 50, no. 4 (October 2, 2015): 273–79. http://dx.doi.org/10.1080/15627020.2015.1079142.

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13

Gordoa, Ana, and Carlos M. Duarte. "Size-Dependent Spatial Distribution of Hake (Merluccius capensis and Merluccius paradoxus) in Namibian Waters." Canadian Journal of Fisheries and Aquatic Sciences 48, no. 11 (November 1, 1991): 2095–99. http://dx.doi.org/10.1139/f91-248.

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Data on hake (Merluccius capensis and Merluccius paradoxus) abundance along the Namibian coast showed both species to have strongly aggregated spatial distributions. In addition, average fish size increased with depth, suggesting that the formation of hake aggregation is based on size-dependent shoaling behaviour. Hake spacing was size dependent with areal fish density and biomass being scaled to the −5 and −2 power, respectively, of fish length. The equations describing the size dependence of the areal fish density were log density (fish∙m−2) = 5.06-4.91 log size (cm) for M. capensis and log density (fish∙m−2) = 5.28-4.74 log size (cm) for (M. paradoxus). The maximum areal densities, presumably corresponding to those within the shoals, exceeded those described by the equations above by 20- and 13-fold for M. capensis and M. paradoxus, respectively. The size dependence of hake spacing implies that fishing pressure targeting aggregates of small hake (i.e. shallow shoals) should have a disproportionate effect on fish mortality and recruitment compared with similar fishing pressure targeting aggregates of larger hake (i.e. deep shoals).
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14

Kainge, P., N. Bahamon, CH Bartholomae, JN Kathena, AK van der Plas, and A. Gordoa. "Stock-environment recruitment analysis for Namibian Cape hake Merluccius capensis." African Journal of Marine Science 35, no. 4 (December 2013): 555–64. http://dx.doi.org/10.2989/1814232x.2013.862509.

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15

Wilhelm, Margit R., Jean-Paul Roux, Coleen L. Moloney, and Astrid Jarre. "Data from fur seal scats reveal when Namibian Merluccius capensis are hatched and how fast they grow." ICES Journal of Marine Science 70, no. 7 (July 19, 2013): 1429–38. http://dx.doi.org/10.1093/icesjms/fst101.

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Abstract Wilhelm, M. R., Roux, J-P., Moloney, C. L., and Jarre, A. 2013. Data from fur seal scats reveal when Namibian Merluccius capensis are hatched and how fast they grow. – ICES Journal of Marine Science, 70: . Scat samples were collected regularly at several breeding colonies of Cape fur seals along the Namibian coast. Merluccius capensis otoliths were obtained from these samples, identified, and measured. Cohorts were easily distinguishable using otolith length measurements converted to fish total length. Growth rates of 2- to 21-month-old hake and hatch dates for each of 15 cohorts were estimated from September 1994 to October 2009 (1994–2008 cohorts) using a Schnute growth function and a non-linear mixed-effects model. The function describing growth of these young hake was length Lt (cm) at age t (years) Lt = 3.17 + (25.0 − 3.17) × [1 − e−0.665 × (t − 0.140)]/[1 − e−0.665 × (1.74–0.140)]. Cohort-specific random effects showed a population hatch date estimate of 31 July (austral winter), varying by 94 days among cohorts, from 31 May (1996 cohort) to 1 September (2004 cohort). The mean growth rate from ages 6 to 12 months was 1.26 cm month−1 for the population, ranging between 0.97 cm month−1 (1996 cohort) and 1.38 cm month−1 (2004 cohort). As this rate is almost double the previously estimated value, which is currently used in the stock assessment models, this result may have major implications for the current stock assessment results and the management of the stock. Re-examination of growth rates needs to be extended to older fish.
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16

Johnsen, Espen. "Improving the precision of length frequency distribution estimates from trawl surveys by including spatial covariance—using Namibian Merluccius capensis as an example." Fisheries Research 62, no. 1 (April 2003): 7–20. http://dx.doi.org/10.1016/s0165-7836(02)00275-8.

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17

Kadila, Hendrina K., Dietlinde N. Nakwaya, Mike Butler, and Johannes A. Iitembu. "Corrigendum to “Insights into feeding interactions of shallow water cape hake (Merluccius capensis) and cape horse mackerel (Trachurus capensis) from the Northern Benguela (Namibia)” [Reg. Stud. Mar. Sci. 34 (2020) 101071]." Regional Studies in Marine Science, December 2020, 101592. http://dx.doi.org/10.1016/j.rsma.2020.101592.

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