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1

Gaudant, Jean, and Milorad Vatsev. "Palaeobatrachids (Amphibia, Anura) in the Oligocene sediments of the “Pirin” Mine (Brezhani Graben, southwestern Bulgaria)." Geologica Balcanica 41, no. 1-3 (2012): 97–103. http://dx.doi.org/10.52321/geolbalc.41.1-3.97.

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Skeletons and tadpoles of Anura belonging to the palaeobatrachids have been collected in the freshwater Early Oligocene sediments of the “Pirin” Mine, near the village of Brezhani, southwestern Bulgaria. Skeletons of cyprinid fishes are also present in the same bituminous argillite. The anatomical study of the palaeobatrachid skeletons has shown that they should be determined as Palaeobatrachus cf. grandipes (Giebel). This new find extends the geographical distribution of this species and of the family Palaeobatrachidae to the southeastern Europe.
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2

Wuttke, Michael, Tomáš Přikryl, Viacheslav Yu Ratnikov, Zdeněk Dvořák, and Zbyněk Roček. "Generic diversity and distributional dynamics of the Palaeobatrachidae (Amphibia: Anura)." Palaeobiodiversity and Palaeoenvironments 92, no. 3 (2012): 367–95. http://dx.doi.org/10.1007/s12549-012-0071-y.

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3

Roček, Zbyněk, Renaud Boistel, Nicolas Lenoir, et al. "Frontoparietal Bone in Extinct Palaeobatrachidae (Anura): Its Variation and Taxonomic Value." Anatomical Record 298, no. 11 (2015): 1848–63. http://dx.doi.org/10.1002/ar.23203.

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4

Vasilyan, Davit, Vladimir S. Zazhigin, and Madelaine Böhme. "Neogene amphibians and reptiles (Caudata, Anura, Gekkota, Lacertilia, and Testudines) from the south of Western Siberia, Russia, and Northeastern Kazakhstan." PeerJ 5 (March 23, 2017): e3025. http://dx.doi.org/10.7717/peerj.3025.

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BackgroundThe present-day amphibian and reptile fauna of Western Siberia are the least diverse of the Palaearctic Realm, as a consequence of the unfavourable climatic conditions that predominate in this region. The origin and emergence of these herpetofaunal groups are poorly understood. Aside from the better-explored European Neogene localities yielding amphibian and reptile fossil remains, the Neogene herpetofauna of Western Asia is understudied. The few available data need critical reviews and new interpretations, taking into account the more recent records of the European herpetofauna. The comparison of this previous data with that of European fossil records would provide data on palaeobiogeographic affiliations of the region as well as on the origin and emergence of the present-day fauna of Western Siberia. An overview of the earliest occurrences of certain amphibian lineages is still needed. In addition, studies that address such knowledge gaps can be useful for molecular biologists in their calibration of molecular clocks.Methods and ResultsIn this study, we considered critically reviewed available data from amphibian and reptile fauna from over 40 Western Siberian, Russian and Northeastern Kazakhstan localities, ranging from the Middle Miocene to Early Pleistocene. Herein, we provided new interpretations that arose from our assessment of the previously published and new data. More than 50 amphibians and reptile taxa were identified belonging to families Hynobiidae, Cryptobranchidae, Salamandridae, Palaeobatrachidae, Bombinatoridae, Pelobatidae, Hylidae, Bufonidae, Ranidae, Gekkonidae, Lacertidae, and Emydidae. Palaeobiogeographic analyses were performed for these groups and palaeoprecipitation values were estimated for 12 localities, using the bioclimatic analysis of herpetofaunal assemblages.ConclusionThe Neogene assemblage of Western Siberia was found to be dominated by groups of European affinities, such as Palaeobatrachidae,Bombina, Hyla,Bufo bufo, and a small part of this assemblage included Eastern Palaearctic taxa (e.g.Salamandrella,Tylototriton,Bufotes viridis). For several taxa (e.g.Mioproteus, Hyla, Bombina,Rana temporaria), the Western Siberian occurrences represented their most eastern Eurasian records. The most diverse collection of fossil remains was found in the Middle Miocene. Less diversity has been registered towards the Early Pleistocene, potentially due to the progressive cooling of the climate in the Northern Hemisphere. The results of our study showed higher-amplitude changes of precipitation development in Western Siberia from the Early Miocene to the Pliocene, than previously assumed.
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Boev, Zlatozar. "Quaternary vertebrate fauna of Bulgaria – composition, chronology and impoverishment." Geologica Balcanica 52, no. 1 (2023): 21–48. http://dx.doi.org/10.52321/geolbalc.52.1.21.

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This study presents for the first time summarized data on 759 species/taxa (628 species at least) of six classes of Quaternary vertebrates of Bulgaria: Chondrichthyes (1); Actinopterygii (34); Amphibia (18); Reptilia (33); Aves (299); and Mammalia (374). The richest fauna has been recorded in the Late Pleistocene (285 species), followed by the Calabrian (255). Bulgaria has lost 32.3% of its former total Quaternary vertebrate fauna. The number of the lost taxa is as follows: species (245), genera (80), families (16), orders (5), of them three mammalian (Perissodactyla, Proboscidea, and Primates), and two avian (Otidiformes and Pteroclidiformes). Extinct families are: one amphibian (Palaeobatrachidae); two reptilian (Varanidae and Elapidae); three avian (Gruidae, Otididae, and Pteroclididae), and ten mammalian (Dipodidae, Eomyidae, Hystricidae, Ochotonidae, Hyaenidae, Phocidae, Equidae, Rhinocerotidae, Elephantidae, and Cercopithecidae). After the small mammals (mainly Cricetidae; 52 taxa), the composition of bovids (27 taxa) and canids (13 taxa) impoverished in a higher extent. The biggest number of recorded vertebrate families is found in the Meghalayan (79), followed by the Greenlandian (63) and the Late Pleistocene (62). At order and family levels, the most varied was the vertebrate fauna in the Meghalayan (39 orders, 79 families). In the Calabrian, the number of genera was a three times greater than in the Northgrippian, which indicates more diversified paleoenvironment. One genus, 25 species, and one subspecies have been described as new to the science from the Quaternary localities in Bulgaria.
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6

Boev, Zlatozar. "Quaternary vertebrate fauna of Bulgaria – composition, chronology and impoverishment." Geologica Balcanica 52, no. 1 (2023): 21–48. https://doi.org/10.52321/GeolBalc.52.1.21.

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This study presents for the first time summarized data on 759 species/taxa (628 species at least) of six classes of Quaternary vertebrates of Bulgaria: Chondrichthyes (1); Actinopterygii (34); Amphibia (18); Reptilia (33); Aves (299); and Mammalia (374). The richest fauna has been recorded in the Late Pleistocene (285 species), followed by the Calabrian (255). Bulgaria has lost 32.3% of its former total Quaternary vertebrate fauna. The number of the lost taxa is as follows: species (245), genera (80), families (16), orders (5), of them three mammalian (Perissodactyla, Proboscidea, and Primates), and two avian (Otidiformes and Pteroclidiformes). Extinct families are: one amphibian (Palaeobatrachidae); two reptilian (Varanidae and Elapidae); three avian (Gruidae, Otididae, and Pteroclididae), and ten mammalian (Dipodidae, Eomyidae, Hystricidae, Ochotonidae, Hyaenidae, Phocidae, Equidae, Rhinocerotidae, Elephantidae, and Cercopithecidae). After the small mammals (mainly Cricetidae; 52 taxa), the composition of bovids (27 taxa) and canids (13 taxa) impoverished in a higher extent. The biggest number of recorded vertebrate families is found in the Meghalayan (79), followed by the Greenlandian (63) and the Late Pleistocene (62). At order and family levels, the most varied was the vertebrate fauna in the Meghalayan (39 orders, 79 families). In the Calabrian, the number of genera was a three times greater than in the Northgrippian, which indicates more diversified paleoenvironment. One genus, 25 species, and one subspecies have been described as new to the science from the Quaternary localities in Bulgaria.
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7

Roček, Zbyněk. ""Lost" And Rediscovered: Holotype Of Palaeobatrachus Diluvianus (Goldfuss, 1831)." Fossil Imprint 72, no. 1-2 (2016): 45–52. https://doi.org/10.14446/FI.2016.45.

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Roček, Zbyněk (2016): "Lost" And Rediscovered: Holotype Of Palaeobatrachus Diluvianus (Goldfuss, 1831). Fossil Imprint 72 (1-2): 45-52, DOI: 10.14446/FI.2016.45, URL: http://dx.doi.org/10.14446/fi.2016.45
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8

Pereda-Suberbiola, Xabier. "Biogeographical affinities of Late Cretaceous continental tetrapods of Europe: a review." Bulletin de la Société Géologique de France 180, no. 1 (2009): 57–71. http://dx.doi.org/10.2113/gssgfbull.180.1.57.

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Abstract The continental tetrapod assemblages from the Santonian-Maastrichtian of Europe consist of dinosaurs (theropods: Abelisauroidea, Alvarezsauridae, Dromaeosauridae, ?Oviraptorosauria, ?Troodontidae, and birds: Enantiornithes, basal Ornithurae; sauropods: Titanosauria; ankylosaurs: Nodosauridae; ornithopods: Hadrosauridae, Rhabdodontidae; and neoceratopsians), pterosaurs (Azhdarchidae), crocodyliforms (eusuchians: Alligatoroidea, Gavialoidea, ?Hylaeochampsidae; sebecosuchian-like ziphosuchians; and, probably, basal neosuchians), choristoderes (?Champsosauridae), squamates (lacertilians: Iguanidae s.l., Paramacellodidae, Polyglyphanodontinae, Varanoidea; snakes: Madtsoiidae; possible amphisbaenians), turtles (cryptodires: Chelydroidea, Kallokibotionidae, Solemydidae; pleurodires: Bothremydidae, Dortokidae), lissamphibians (Albanerpetontidae; anurans: Discoglossidae, Palaeobatrachidae; caudates: Batrachosauroididae, Salamandridae), and mammals (multituberculates: Kogaionidae, ?“Paracimexomys group”; marsupials: Herpetotheriidae; eutherians: “Zhelestidae”). The palaeobiogeographical affinities of the Late Cretaceous continental tetrapods of Europe are complex. The faunas are commonly considered to show a mixed pattern resulting from the addition of “Asiamerican” and Gondwanan forms to European taxa. Albanerpetontids, both paramacellodid and polyglyphonodontine lizards, and hadrosaurid dinosaurs are taxa with Palaeolaurasian (or, in some case, even Neopangean) affinities. Other forms, such as paleobatrachid and batrachosauroidid lissamphibians, solemydid turtles, alligatoroid crocodyliforms, and nodosaurid dinosaurs can be considered as Euramerican taxa. Kallokibotionid and dortokid turtles, rhabdodontid dinosaurs and kogaionid mammals are endemic to Europe. The Gondwanan taxa have been regarded as African immigrants that reached southern Europe via the Mediterranean Tethyan sill. Abelisaurid and titanosaurid dinosaurs, trematochampsid crocodyliforms, podocnemidid and bothremydid turtles, and boid and madtsoiid snakes constitute the basic pattern of the so-called “Eurogondwanan fauna”. However, the validity of some of these taxa is a disputed matter (Titanosauridae, Trematochampsidae), and the presence of other taxa in the Late Cretaceous of Europe is based on controversial data (Boidae, Podocnemididae). Only Abelisauroidea, Madtsoiidae and Bothremydidae (and, yet for confirming, Sebecosuchia) provide evidence of interchanges between Africa and Europe. At least abelisauroids might have reached Europe before the Late Cretaceous. In conclusion, most of the continental tetrapod taxa from the latest Cretaceous of Europe show affinities with those of northern Hemisphere landmasses. Latest Cretaceous trans-Tethyan dispersal events between Africa and Europe remain poorly documented.
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9

Báez, Ana M., and Borja Sanchiz. "A review of Neusibatrachus wilferti, an Early Cretaceous frog from the Montsec Range, northeastern Spain." Acta Palaeontologica Polonica 52, no. 3 (2007): 477–87. https://doi.org/10.5281/zenodo.13741546.

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Báez, Ana M., Sanchiz, Borja (2007): A review of Neusibatrachus wilferti, an Early Cretaceous frog from the Montsec Range, northeastern Spain. Acta Palaeontologica Polonica 52 (3): 477-487, DOI: 10.5281/zenodo.13741546
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10

Roček, Zbyněk. "Larval development in Oligocene palaeobatrachid frogs." Acta Palaeontologica Polonica 48, no. 4 (2003): 595–607. https://doi.org/10.5281/zenodo.13393134.

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11

Syromyatnikova, E. V. "New palaeobatrachid records from Asia." Palaeoworld 27, no. 3 (2018): 410–14. http://dx.doi.org/10.1016/j.palwor.2018.05.002.

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12

Roček, Zbyněk, Ronald Böttcher, and Richard Wassersug. "Gigantism in tadpoles of the Neogene frogPalaeobatrachus." Paleobiology 32, no. 4 (2006): 666–75. http://dx.doi.org/10.1666/05073.1.

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We describe three giant palaeobatrachid fossil tadpoles of the genusPalaeobatrachus(Nieuwkoop-Faber [NF] stages 60–64) from the Miocene of Randecker Maar, Germany. The largest was 150 mm at the beginning of metamorphosis (stage 60), whereas the smallest was 100 mm and approaching the end of metamorphosis (stage 64). In contrast, normal palaeobatrachid tadpoles and their pipid relatives, both extinct and extant, rarely exceed 60 mm in length. We review here both ecological and pathological conditions that are conducive to the development of gigantism in tadpoles. Tadpoles that lack a thyroid gland become exceptionally large and arrest development at early hindlimb stages (NF stages 53–56). However, the advanced metamorphic stages of the giantPalaeobatrachustadpoles indicate that they were able to metamorphose, and thus were not athyroid. Environmental factors—pond size and permanence, predators, duration of the growing season—may all contribute to tadpole gigantism in certain extant anuran species. We identify suites of ecological features that distinguish extant anurans with large tadpoles from high-latitude and high-altitude permanent lakes in temperate regions (e.g., certainRanaandTelmatobius) from tropical species, such asPseudis paradoxa, whose tadpoles normally achieve large size in temporary seasonal ponds. The paleoecology of Randecker Maar suggests thatPalaeobatrachustadpoles lived in a permanent semitropical lake, but one with few predators.
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13

Sanchiz, B., M. Alcobendas, and H. H. Schleich. "Evidence of Seasonal Growth in Extinct Oligocene Palaeobatrachid Anurans." Copeia 1994, no. 1 (1994): 232. http://dx.doi.org/10.2307/1446692.

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14

Villa, Andrea, Loredana Macaluso, and Thomas Mörs. "Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe." Palaeontologia Electronica 27, no. 1 (2024): 1–56. https://doi.org/10.26879/1323.

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Villa, Andrea, Macaluso, Loredana, Mörs, Thomas (2024): Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe. Palaeontologia Electronica (a3) 27 (1): 1-56, DOI: 10.26879/1323, URL: http://dx.doi.org/10.26879/1323
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Venczel, Márton, Vlad Codrea, and Cristina Fărcaş. "A new palaeobatrachid frog from the early Oligocene of Suceag, Romania." Journal of Systematic Palaeontology 11, no. 2 (2012): 179–89. http://dx.doi.org/10.1080/14772019.2012.671790.

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16

Villa, Andrea, Zbyněk Roček, Emanuel Tschopp, Lars W. Van Den Hoek Ostende, and Massimo Delfino. "Palaeobatrachus eurydices, sp. nov. (Amphibia, Anura), the last Western European palaeobatrachid." Journal of Vertebrate Paleontology 36, no. 6 (2016): e1211664. http://dx.doi.org/10.1080/02724634.2016.1211664.

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17

Syromyatnikova, Elena, Alexey Tesakov, Serdar Mayda, Tanju Kaya, and Gerçek Saraç. "Plio-Pleistocene Amphibians And Reptiles From Central Turkey: New Faunas And Faunal Records With Comments On Their Biochronological Position Based On Small Mammals." Fossil Imprint 75, no. 3-4 (2019): 343–58. https://doi.org/10.2478/if-2019-0022.

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Syromyatnikova, Elena, Tesakov, Alexey, Mayda, Serdar, Kaya, Tanju, Saraç, Gerçek (2019): Plio-Pleistocene Amphibians And Reptiles From Central Turkey: New Faunas And Faunal Records With Comments On Their Biochronological Position Based On Small Mammals. Fossil Imprint 75 (3-4): 343-358, DOI: 10.2478/if-2019-0022, URL: http://dx.doi.org/10.2478/if-2019-0022
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18

Codrea, Vlad A., Marian Bordeianu, and Márton Venczel. "Amphibians and squamate reptiles from the late Miocene of Fălciu (Eastern Romania)." Palaeontologia Electronica 25, no. 2 (2021): 1–23. https://doi.org/10.26879/1156.

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Codrea, Vlad A., Bordeianu, Marian, Venczel, Márton (2021): Amphibians and squamate reptiles from the late Miocene of Fălciu (Eastern Romania). Palaeontologia Electronica (a19) 25 (2): 1-23, DOI: 10.26879/1156, URL: http://dx.doi.org/10.26879/1156
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19

Hossini, Saïda, and Jean-Claude Rage. "Palaeobatrachid frogsfrom the earliest Miocene (Agenian) of France, with description of a new species." Geobios 33, no. 2 (2000): 223–31. http://dx.doi.org/10.1016/s0016-6995(00)80019-4.

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20

Venczel, Márton, and Emanoil Stiuc. "Late middle Miocene amphibians and squamate reptiles from Taut,, Romania." Geodiversitas 30, no. 4 (2008): 731–63. https://doi.org/10.5281/zenodo.4665621.

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21

Garcia, Géraldine, Sylvain Duffaud, Monique Feist, et al. "La Neuve, gisement à plantes, invertébrés et vertébrés du Bégudien (Sénonien supérieur continental) du bassin d'Aix-en-Provence." Geodiversitas 22, no. 3 (2000): 325–48. https://doi.org/10.5281/zenodo.5371115.

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Garcia, Géraldine, Duffaud, Sylvain, Feist, Monique, Marandat, Bernard, Tambareau, Yvette, Villatte, Juliette, Sigé, Bernard (2000): La Neuve, gisement à plantes, invertébrés et vertébrés du Bégudien (Sénonien supérieur continental) du bassin d'Aix-en-Provence. Geodiversitas 22 (3): 325-348, DOI: 10.5281/zenodo.5371115
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22

Kvaček, Zlatko, Vasilis Teodoridis, and Miroslav Radoň. "Review of The Late Oligocene Flora of Matrý Near Sebuzín (České Středohoří Mts., The Czech Republic)." Fossil Imprint 74, no. 3-4 (2018): 292–316. http://dx.doi.org/10.2478/if-2018-0018.

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The Oligocene palaeontological locality on Matrý Hill near Sebuzín in the České středohoří Mts., North Bohemia, belongs to the Děčín Formation and is dated to 30.8-24.7 Ma according to the regional stratigraphy. It has yielded, in addition to insects from the Heteroptera group, a fossil bee Apis petrefacta and palaeobatrachid frogs, also numerous plant remains. Their recovery began in 1996. The plant fossil assemblage consists mostly of leaf impressions, occasionally accompanied by casts or impressions of fruits. Noteworthy are records of a fern Woodwardia muensteriana, conifers Pinus cf. rigios, P. cf. hepios, Calocedrus suleticensis, Tetraclinis salicornioides, Torreya bilinica, cf. Cephalotaxus parvifolia and numerous angiosperms, e.g. Liriodendron haueri, Daphnogene cinnamomifolia, Platanus neptuni, Cercidiphyllum crenatum, Sloanea artocarpites, Ulmus pyramidalis, Celtis pirskenbergensis, Carya fragiliformis, C. quadrangula, Betula brongniartii, B. dryadum, Alnus rhenana, A. cf. kefersteinii, Carpinus grandis, Ostrya atlantidis, Acer crenatifolium, A. cf. palaeosaccharinum, A. integrilobum and Craigia bronnii. Several angiosperm foliage specimens of both monocots and dicots have not yet been identified to a particular genus and species. The fossil plant assemblage at Matrý corresponds to two vegetation types, i.e. a zonal riparian forest and zonal mixed mesophytic forest, as corroborated by the Integrated Plant Record vegetation analysis. The vegetation thrived under a humid climate, characterized by average values of MAT (13.4 °C), WMMT (23.8 °C), CMMT (3.6 °C) and MAP (1,117 mm). The Matry fossil flora is similar in composition to the flora of Žichov from the same Oligocene Děčín Formation in the České středohoří Mts.
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23

Syromyatnikova, Elena. "Palaeobatrachid frog from the late Miocene of Northern Caucasus, Russia." Palaeontologia Electronica, 2018. http://dx.doi.org/10.26879/861.

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