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Journal articles on the topic 'Phrynopus'

Author: Grafiati

Published: 4 June 2021

Last updated: 28 July 2025

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1

Lehr, Edgar, Jiri Moravec, and Juan Carlos Cusi. "Two new species of Phrynopus (Anura, Strabomantidae) from high elevations in the Yanachaga-Chemillén National park in Peru (Departamento de Pasco)." ZooKeys 235 (October 31, 2012): 51–71. https://doi.org/10.3897/zookeys.235.3885.

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We describe two new species of <i>Phrynopus</i> from a cloud forest of the Cordillera Yanachaga, Yanachaga-Chemillén National Park in Peru and describe and document the first clutch and case of parental care for a species of <i>Phrynopus</i>. One of the new species of <i>Phrynopus </i>is described based on two females (SVL 19.1–21.0 mm) which were found in leaf litter and moss layer in a cloud forest at 2900 m elevation. This new species is most similar to <i>Phrynopus bracki</i>. The second new species of <i>Phrynopus </i>was found in the transitional formation between cloud forest and wet pu

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2

LEHR, EDGAR, and ANAHÍ ORÓZ. "Two new species of Phrynopus (Anura: Strabomantidae) from the Cordillera de Carpish in central Peru (Departamento de Huánuco)." Zootaxa 3512, no. 1 (2012): 53. http://dx.doi.org/10.11646/zootaxa.3512.1.3.

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We describe two new species of Phrynopus from cloud forests of the Cordillera de Carpish, Departamento de Huánuco incentral Peru, increasing the number of currently known Phrynopus to 23. Phrynopus interstinctus n. sp. is described basedon one female (SVL 23.8 mm) and two males (SVL 14.9–16.5 mm). Specimens were found between 3100 and 3180 melevation. This species has weakly developed dorsolateral folds, minute dentigerous processes of vomers, no tympanum,and coloration in life consisting of a reddish-brown, dark grayish-brown or olive-brown dorsum, a black venter with whiteblotches and flecks

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3

CHAPARRO, JUAN C., JOSE M. PADIAL, and IGNACIO DE LA RIVA. "Two sympatric new species of Phrynopus (Anura: Strabomantidae) from Yanachaga Chemillen National Park (central Peruvian Andes)." Zootaxa 1761, no. 1 (2008): 49. http://dx.doi.org/10.11646/zootaxa.1761.1.5.

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Two sympatric new species of Phrynopus (Anura: Strabomantidae) are described from elfin forests and puna grasslands (3363–3589 m) of the eastern slopes of the Andes of Departamento Pasco in central Peru. Phrynopus miroslawae sp. nov. is a medium-sized species characterized by lacking vomerine teeth and tympanic membrane, and by having dorsolateral folds, areolate ventral skin, dorsum gray with black spots, venter cream with small scattered black blotches, and iris bronze. Phrynopus nicoleae sp. nov. is a small species characterized by lacking tympanic membrane and by having vomerine teeth, an

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4

RODRIGUEZ, LILY O., and ALESSANDRO CATENAZZI. "Four new species of terrestrial-breeding frogs of the genus Phrynopus (Anura: Terrarana: Craugastoridae) from Río Abiseo National Park, Peru." Zootaxa 4273, no. 3 (2017): 381. http://dx.doi.org/10.11646/zootaxa.4273.3.4.

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We describe four new species of terrestrial-breeding frogs belonging to the genus Phrynopus from specimens collected on the eastern slopes of the Cordillera Oriental (2800–3850 m) near and within Río Abiseo National Park, Provincia Mariscal Cáceres, Departments of San Martín and La Libertad, northeastern Peru. All four species lack a visible tympanum and inhabit the upper ridges and slopes within or adjacent to the Park. Phrynopus anancites sp. nov. and P. capitalis sp. nov. inhabit the wet montane grasslands on the upper ridges and valleys from 3600 to 3850 m. Phrynopus anancites (SVL = 25.3

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5

Lehr, Edgar, May Rudolf von, Jiri Moravec, and Juan Carlos Cusi. "A new species of Phrynopus (Amphibia, Anura, Craugastoridae) from upper montane forests and high Andean grasslands of the Pui Pui Protected Forest in central Peru." ZooKeys 713 (November 2, 2017): 131–57. https://doi.org/10.3897/zookeys.713.20776.

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We describe a new species of Phrynopus from the upper montane forests and high Andean grasslands (puna) of the Pui Pui Protected Forest and its close surroundings (Región Junín, central Peru) and compare it morphologically and genetically with other species of Phrynopus. Phrynopus inti sp. n. is known from four localities outside and two localities inside the Pui Pui Protected Forest between 3350 and 3890 m a.s.l. Studied specimens of the new species are characterized by a snout-vent length of 27.2–35.2 mm in males (n = 6), and 40.4 mm in a single female, by having the skin on dorsum and flank

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6

VENEGAS, PABLO J., ANDY C. BARBOZA, IGNACIO DE LA RIVA, and JOSÉ M. PADIAL. "A new species of Phrynopus from the northeastern Andes of Peru, its phylogenetic position, and notes on the relationships of Holoadeninae (Anura: Craugastoridae)." Zootaxa 4446, no. 4 (2018): 501. http://dx.doi.org/10.11646/zootaxa.4446.4.5.

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We report the discovery of a geographically disjunct and morphologically distinctive species of direct-developing frog of the genus Phrynopus (Phrynopus mariellaleo sp. nov.) that changes considerably our understanding of the distribution of species in this Andean genus. The type locality lies on a subcordillera (Cerro de Campanario area) of the extreme northeastern portion of the Cordillera Central of Peru, on the headwaters of the Mayo River, Amazonas department, at 2575 m asl (6°6’42.9’’S, 77°26’24’’W). This area is situated 170 km to the NE from the northernmost record of Phrynopus known s

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7

Chávez, Germán, Luis Alberto García Ayachi, and Alessandro Catenazzi. "A new species of frog (Terrarana, Strabomantidae, Phrynopus) from the Peruvian Andean grasslands." PeerJ 8 (June 24, 2020): e9433. http://dx.doi.org/10.7717/peerj.9433.

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We describe a new, medium-sized species of terrestrial frog of the genus Phrynopus from a single locality in the central Andes of Peru (Departamento de Huánuco) at 3,730 meters of elevation. Phylogenetic analyses supported Phrynopus remotum sp. nov. as an independent lineage, sister to most of its congeners. The new species is morphologically distinguishable by the presence of small tubercles on upper eyelids and heels, an areolate venter, and the absence of dorsolateral folds or ridges. This species inhabits the highlands adjacent to the Marañón Dry valley. The only sympatric amphibian specie

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8

Lehr, Edgar. "Rediscovery of Phrynopus peruanus Peters 1874 (Amphibia, Anura, Leptodactylidae)." Zootaxa 1485 (December 31, 2007): 51–57. https://doi.org/10.5281/zenodo.176955.

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9

Lehr, Edgar, and César Aguilar. "The taxonomic status of Phrynopus pereger Lynch 1975 (Amphibia, Anura, Leptodactylidae)." Zootaxa 1284 (December 31, 2006): 53–60. https://doi.org/10.5281/zenodo.173462.

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10

Mamani, Luis, and Sergio Malqui. "A new species of Phrynopus (Anura: Craugastoridae) from the central Peruvian Andes." Zootaxa 3838, no. 2 (2014): 207–14. https://doi.org/10.11646/zootaxa.3838.2.5.

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11

Duellman, William E., and Blair Hedges. "Two new minute species of Phrynopus (Lissamphibia: Anura) from the Cordillera oriental in Peru." Zootaxa 1675 (December 31, 2008): 59–66. https://doi.org/10.5281/zenodo.274057.

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12

Lehr, Edgar, and Anahí Oróz. "Two new species of Phrynopus (Anura: Strabomantidae) from the Cordillera de Carpish in central Peru (Departamento de Huánuco)." Zootaxa 3512 (December 31, 2012): 53–63. https://doi.org/10.5281/zenodo.210408.

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Lehr, Edgar, Oróz, Anahí (2012): Two new species of Phrynopus (Anura: Strabomantidae) from the Cordillera de Carpish in central Peru (Departamento de Huánuco). Zootaxa 3512: 53-63, DOI: 10.5281/zenodo.210408

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13

May, Rudolf Von. "Rediscovery of the Endangered frog Phrynopus montium (Shreve, 1938) (Amphibia, Anura, Craugastoridae) in the cloud forest of central Peru." Zootaxa 4338, no. 1 (2017): 195–200. https://doi.org/10.11646/zootaxa.4338.1.13.

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May, Rudolf Von (2017): Rediscovery of the Endangered frog Phrynopus montium (Shreve, 1938) (Amphibia, Anura, Craugastoridae) in the cloud forest of central Peru. Zootaxa 4338 (1): 195-200, DOI: 10.11646/zootaxa.4338.1.13

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14

Chaparro, Juan C., José M. Padial, and Ignacio De La Riva. "Two sympatric new species of Phrynopus (Anura: Strabomantidae) from Yanachaga Chemillén National Park (central Peruvian Andes)." Zootaxa 1761 (December 31, 2008): 49–58. https://doi.org/10.5281/zenodo.181938.

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Chaparro, Juan C., Padial, José M., Riva, Ignacio De La (2008): Two sympatric new species of Phrynopus (Anura: Strabomantidae) from Yanachaga Chemillén National Park (central Peruvian Andes). Zootaxa 1761: 49-58, DOI: 10.5281/zenodo.181938

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15

Rodriguez, Lily O., and Alessandro Catenazzi. "Four new species of terrestrial-breeding frogs of the genus Phrynopus (Anura: Terrarana: Craugastoridae) from Río Abiseo National Park, Peru." Zootaxa 4273, no. 3 (2017): 381–406. https://doi.org/10.11646/zootaxa.4273.3.4.

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Rodriguez, Lily O., Catenazzi, Alessandro (2017): Four new species of terrestrial-breeding frogs of the genus Phrynopus (Anura: Terrarana: Craugastoridae) from Río Abiseo National Park, Peru. Zootaxa 4273 (3): 381-406, DOI: 10.11646/zootaxa.4273.3.4

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16

Chaparro, Juan C., Ignacio De La Riva, José M. Padial, José A. Ochoa, and Edgar Lehr. "A new species of Phrynopus from Departamento Cusco, southern Peru (Anura: Brachycephalidae)." Zootaxa 1618 (December 31, 2007): 61–68. https://doi.org/10.5281/zenodo.179135.

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Chaparro, Juan C., Riva, Ignacio De La, Padial, José M., Ochoa, José A., Lehr, Edgar (2007): A new species of Phrynopus from Departamento Cusco, southern Peru (Anura: Brachycephalidae). Zootaxa 1618: 61-68, DOI: 10.5281/zenodo.179135

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17

Lehr, Edgar, and Daniel Rodríguez. "Two new species of Andes Frogs (Craugastoridae: Phrynopus) from the Cordillera de Carpish in central Peru." Salamandra 53, no. 3 (2017): 327–38. https://doi.org/10.5281/zenodo.2585687.

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We describe two new species of Phrynopus from the Unchog Elfin Forest of the Cordillera de Carpish in the eastern Andes of central Peru, Región Huánuco. Specimens were obtained from the Puna at elevations between 3276 and 3582 m above sea level. One of the new species is described based on a single male which has a pale gray coloration with large tubercles on dorsum and flanks. It is most similar to P. bufoides and P. thompsoni. The second new species is described based on a male and a female. This new species has a grayish-brown coloration with reddish-brown groin, and discontin

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18

Díaz, Vladimir, Luis Mamani, and Alessandro Catenazzi. "A new species of Andean frog of the genus Phrynopus (Anura: Strabomantidae) from southeastern Peru." Zootaxa 5293, no. 2 (2023): 333–48. https://doi.org/10.11646/zootaxa.5293.2.7.

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Díaz, Vladimir, Mamani, Luis, Catenazzi, Alessandro (2023): A new species of Andean frog of the genus Phrynopus (Anura: Strabomantidae) from southeastern Peru. Zootaxa 5293 (2): 333-348, DOI: 10.11646/zootaxa.5293.2.7, URL: http://dx.doi.org/10.11646/zootaxa.5293.2.7

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19

Riva, Ignacio De La, Juan C. Chaparro, and José M. Padial. "The taxonomic status of Phyllonastes Heyer and Phrynopus peruvianus (Noble) (Lissamphibia, Anura): resurrection of Noblella Barbour." Zootaxa 1685, no. 1 (2008): 67–68. https://doi.org/10.11646/zootaxa.1685.1.5.

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Riva, Ignacio De La, Chaparro, Juan C., Padial, José M. (2008): The taxonomic status of Phyllonastes Heyer and Phrynopus peruvianus (Noble) (Lissamphibia, Anura): resurrection of Noblella Barbour. Zootaxa 1685 (1): 67-68, DOI: 10.11646/zootaxa.1685.1.5, URL: http://dx.doi.org/10.11646/zootaxa.1685.1.5

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20

Chávez, Germán, Roy Santa-Cruz, Daniel Rodriguez, Edgar Lehr Abstract.-We, Peru. One, and La Libertad. "Two new species of frogs of the genus Phrynopus (Anura: Terrarana: Craugastoridae) from the Peruvian Andes." Amphibian & Reptile Conservation 9, no. 1 (2015): 15–25. https://doi.org/10.5281/zenodo.13270172.

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Chávez, Germán, Santa-Cruz, Roy, Rodriguez, Daniel, Abstract.-We, Edgar Lehr, One, Peru., Libertad, La (2015): Two new species of frogs of the genus Phrynopus (Anura: Terrarana: Craugastoridae) from the Peruvian Andes. Amphibian & Reptile Conservation (e105) 9 (1): 15-25, DOI: 10.5281/zenodo.13270172

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21

Venegas, Pablo J., Andy C. Barboza, Ignacio De La Riva, and José M. Padial. "A new species of Phrynopus from the northeastern Andes of Peru, its phylogenetic position, and notes on the relationships of Holoadeninae (Anura: Craugastoridae)." Zootaxa 4446, no. 4 (2018): 501–24. https://doi.org/10.11646/zootaxa.4446.4.5.

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Venegas, Pablo J., Barboza, Andy C., Riva, Ignacio De La, Padial, José M. (2018): A new species of Phrynopus from the northeastern Andes of Peru, its phylogenetic position, and notes on the relationships of Holoadeninae (Anura: Craugastoridae). Zootaxa 4446 (4): 501-524, DOI: 10.11646/zootaxa.4446.4.5

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22

DÍAZ, VLADIMIR, LUIS MAMANI, and ALESSANDRO CATENAZZI. "A new species of Andean frog of the genus Phrynopus (Anura: Strabomantidae) from southeastern Peru." Zootaxa 5293, no. 2 (2023): 333–48. http://dx.doi.org/10.11646/zootaxa.5293.2.7.

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The Cordillera de los Andes is one of the most important regions for biodiversity. Among amphibians, many endemic species of terrestrial-breeding frogs have recently been discovered. Herein we describe Phrynopus sancristobali from the Andes of southeastern Peru based on molecular and morphological data. The new species is known from the ecotone between humid puna and montane forest at 3910 m a.s.l. on the left side of the Apurímac Valley in the Department of Ayacucho. The new species differs from congeners by having dorsum bearing pustules and light brown with dark brown reticulations surround

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23

LEHR, EDGAR. "NEW ELEUTHERODACTYLINE FROGS (LEPTODACTYLIDAE: PRISTIMANTIS, PHRYNOPUS) FROM PERU." Bulletin of the Museum of Comparative Zoology 159, no. 2 (2007): 145–78. http://dx.doi.org/10.3099/0027-4100(2007)159[145:neflpp]2.0.co;2.

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24

NA. "New eleutherodactyline frogs (Leptodactylidae: Pristimantis, Phrynopus) from Peru." Bulletin of the Museum of Comparative Zoology at Harvard College 159 (June 12, 2007): 145–78. https://doi.org/10.5281/zenodo.13492681.

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25

NA. "New eleutherodactyline frogs (Leptodactylidae: Pristimantis, Phrynopus) from Peru." Bulletin of the Museum of Comparative Zoology at Harvard College 159 (June 7, 2007): 145–78. https://doi.org/10.5281/zenodo.13492681.

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26

NA. "New eleutherodactyline frogs (Leptodactylidae: Pristimantis, Phrynopus) from Peru." Bulletin of the Museum of Comparative Zoology at Harvard College 159 (July 3, 2007): 145–78. https://doi.org/10.5281/zenodo.13492681.

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27

NA. "New eleutherodactyline frogs (Leptodactylidae: Pristimantis, Phrynopus) from Peru." Bulletin of the Museum of Comparative Zoology at Harvard College 159 (July 10, 2007): 145–78. https://doi.org/10.5281/zenodo.13492681.

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28

DUELLMAN, WILLIAM E., and S. BLAIR HEDGES. "Two new minute species of Phrynopus (Lissamphibia: Anura) from the Cordillera oriental in Peru." Zootaxa 1675, no. 1 (2008): 59. http://dx.doi.org/10.11646/zootaxa.1675.1.4.

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Two small new species of terrestrial frogs of the genus Phrynopus are described from the Andes in the Departament of Pasco in central Peru. The presence of a tympanum in one species is shared with P. peruanus. The second species shares characters with the larger P. dagmarae from the Departament of Huánuco. The new species are the smallest known members of the genus.

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29

Hedges, S. Blair. "A New Species of Phrynopus (Anura: Leptodactylidae) from Perú." Copeia 1990, no. 1 (1990): 108. http://dx.doi.org/10.2307/1445826.

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30

LEHR, EDGAR. "Rediscovery of Phrynopus peruanus Peters 1874 (Amphibia, Anura, Leptodactylidae)." Zootaxa 1485, no. 1 (2007): 51–57. http://dx.doi.org/10.11646/zootaxa.1485.1.5.

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Phrynopus peruanus, the type species of the genus, was described by Peters in 1874 based on three specimens from “Maraynioc in Peru”. In 1975, Lynch redescribed the species based on two preserved females, but skin texture, male characters, coloration in life, and precise information on the type locality and natural history remained unknown or uncertain. In December 2005, four specimens of P. peruanus were found at the Puna of Maraynioc at 3825 m a.s.l. in the Vitoc Valley (Departamento de Junín). Males were calling from inside the bunched Peruvian Feather Grass (Stipa icchu). Coloration in lif

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31

DE LA RIVA, IGNACIO, JUAN C. CHAPARRO, and JOSÉ M. PADIAL. "A new, long-standing misidentified species of Psychrophrynella Hedges, Duellman & Heinicke from Departamento Cusco, Peru (Anura: Strabomantidae)." Zootaxa 1823, no. 1 (2008): 42. http://dx.doi.org/10.11646/zootaxa.1823.1.3.

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We present the description of a new species of the recently described genus Psychrophrynella. Hitherto, frogs of this new species were traditionally misidentified as Phrynopus peruvianus. The new species is known only from elevations of 3270–3450 m in the vicinity of the type locality in the Cordillera de Paucartambo, Departamento Cusco, Peru. It is unique within Psychrophrynella in having a marked, oblique inner tarsal tubercle. The geographically closest species Psychrophrynella bagrecito Lynch, also possesses a tarsal tubercle.

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32

Lehr, Edgar, and May Rudolf von. "A new species of terrestrial-breeding frog (Amphibia, Craugastoridae, Pristimantis) from high elevations of the Pui Pui Protected Forest in central Peru." ZooKeys 660 (March 7, 2017): 17–42. https://doi.org/10.3897/zookeys.660.11394.

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We describe a new species of Pristimantis from upper montane forests and high Andean grasslands of the Pui Pui Protected Forest and its close surroundings, Región Junín, central Peru. The description of the new species is based on 34 specimens found at elevations between 3400 and 3936 m a.s.l. Pristimantis attenboroughi sp. n. is characterized by a snout–vent length of 14.6–19.2 mm in adult males (n = 21), 19.2–23.0 mm in adult females (n = 10), and is compared morphologically and genetically with other taxonomically and biogeographically relevant species of Pristimantis. The new species is ch

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33

CHAPARRO, JUAN C., IGNACIO DE LA RIVA, JOSÉ M. PADIAL, JOSÉ A. OCHOA, and EDGAR LEHR. "A new species of Phrynopus from Departamento Cusco, southern Peru(Anura: Brachycephalidae)." Zootaxa 1618, no. 1 (2007): 61–68. http://dx.doi.org/10.11646/zootaxa.1618.1.3.

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We describe a new species of Phrynopus (Anura: Brachycephalidae) from two close localities at the upper limits of cloud forest in the southern Peruvian Departamento Cusco, between 3555–3950 m a.s.l. The new species is characterized by having medium size (maximum snout-vent length 23.4 mm), dentigerous processes of vomers absent, tympanic membrane inconspicuous, dorsal skin coarsely shagreen in life, dorsolateral folds, ventral skin areolate, dorsum tan, venter bold black with conspicuous bluish-gray spots, and a bluish-white iris.

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34

Romero de Pérez, Gloria, and Pedro M. Ruiz C. "HISTOLOGIA, HISTOQUÍMICA Y ESTRUCTURA FINA DE LA GLANDULA MENTONIANA DE DOS ESPECIES DE HYLA (GRUPO BOGOTENSIS) Y DEL ANTEBRAZO DE PHRYNOPUS ADENOBRACHIUS." Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales 20, no. 78 (2024): 575–84. http://dx.doi.org/10.18257/raccefyn.20(78).1996.3053.

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Se estudió la histoquímica y la estructura por microscopía óptica (MO) y microscopía electrónica de transmisión (TEM) de la glándula mentoniana de machos adultos de dos especies de Hylabogotensis grupo similares en forma y función con las glándulas mentonianas (hedónicas) de especies de Plethodontidae. Un gran ensanchamiento del antebrazo de los machos adultos de Phrynopus adenobrachius, estudiada con los mismos métodos (MO, TEM) muestra que un sistema “lagunar” en el estrato esponjoso de la dermis contribuye al afianzamiento de la pareja durante el amplexo.

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35

Chávez, Germán, Luis A. García-Ayachi, and Alessandro Catenazzi. "A new species of frog (Terrarana, Strabomantidae, Phrynopus) from the Peruvian Andean Grasslands." Evolutionary Systematics 7, no. 1 (2023): 105–16. http://dx.doi.org/10.3897/evolsyst.7.96258.

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We describe a new terrestrial frog from the puna grasslands adjacent to the Mantaro dry valley in southern Peru. Phrynopus apumantarumsp. nov. is similar in appearence to P. bufoides but is differentiable by lacking discoidal fold and enlarged warts on dorsum, lacking a prominent post ocular fold and having canthal and post ocular stripe. Lastly, we propose to place the new species under the Vulnerable (VU) category of the IUCN Red List, due its small distribution and habitat loss recorded at the type locality.

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36

Chávez, Germán, Luis A. García-Ayachi, and Alessandro Catenazzi. "A new species of frog (Terrarana, Strabomantidae, Phrynopus) from the Peruvian Andean Grasslands." Evolutionary Systematics 7 (March 16, 2023): 105–16. https://doi.org/10.3897/evolsyst.7.96258.

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We describe a new terrestrial frog from the puna grasslands adjacent to the Mantaro dry valley in southern Peru. Phrynopus apumantarum sp. nov. is similar in appearence to P. bufoides but is differentiable by lacking discoidal fold and enlarged warts on dorsum, lacking a prominent post ocular fold and having canthal and post ocular stripe. Lastly, we propose to place the new species under the Vulnerable (VU) category of the IUCN Red List, due its small distribution and habitat loss recorded at the type locality.

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37

Lehr, Edgar, Mikael Lundberg, and César Aguilar. "Three New Species of Phrynopus from Central Peru (Amphibia: Anura: Leptodactylidae)." Copeia 2005, no. 3 (2005): 479–91. http://dx.doi.org/10.1643/ch-04-278r2.

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38

MAY, RUDOLF VON. "Rediscovery of the Endangered frog Phrynopus montium (Shreve, 1938) Amphibia, Anura, Craugastoridae) in the cloud forest of central Peru." Zootaxa 4338, no. 1 (2017): 195. http://dx.doi.org/10.11646/zootaxa.4338.1.13.

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Phrynopus montium (Shreve, 1938) was described from Cascas, Junín Department, Peru, at an undetermined elevation between 3000 and 4000 m a.s.l. in the Cordillera Central. With the exception of the type specimens used in the original description, no additional sightings of P. montium had been reported for 76 years. Additionally, its coloration in life, microhabitat use, and precise elevational distribution remained unknown. During a field expedition conducted in 2014, two individuals of P. montium were found at an upper cloud forest site in central Peru, 8.7 km south from Hacienda Cascas. Here,

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DE LA RIVA, IGNACIO, JUAN C. CHAPARRO, and JOSÉ M. PADIAL. "The taxonomic status of Phyllonastes Heyer and Phrynopus peruvianus (Noble) (Lissamphibia, Anura): resurrection of Noblella Barbour." Zootaxa 1685, no. 1 (2008): 67. http://dx.doi.org/10.11646/zootaxa.1685.1.5.

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Between December 1899 and August 1900, H. H. Keays collected zoological specimens in Departamento Puno, southern Peru. He shipped to the American Museum of Natural History a series of amphibians purportedly obtained near Juliaca, a locality at 3824 m a.s.l., close to the Lake Titicaca. From this series, Noble (1921) described a tiny frog named as Sminthillus peruvianus. Until then, the genus Sminthillus Barbour and Noble contained only the small Cuban species S. limbatus (Cope), and the broad geographic gap between the two species, together with some morphological arguments, led Noble to consi

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LEHR, EDGAR, and ALESSANDRO CATENAZZI. "A new species of Bryophryne (Anura: Strabomantidae) from southern Peru." Zootaxa 1784, no. 1 (2008): 1. http://dx.doi.org/10.11646/zootaxa.1784.1.1.

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A new species of Bryophryne from southern Peru (Cusco Region) is described. Specimens were found in the leaf litter of cloud forest at elevations of 2350–3215 m. The new species has a maximum snout-vent length of 21.9 mm in adult females, 18.9 mm in adult males and is the smallest species of the genus. It lacks a tympanum and dentigerous processes of vomers, has dorsolateral folds, and males without vocal slits and without nuptial pads. The new species is most similar to B. bustamantei but differs in being smaller, having discontinuous dorsolateral folds, the males lacking vocal slits, and an

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Harvey, Michael B., and Patricia Ergueta S. "A New Species of Phrynopus (Anura: Leptodactylidae) from Elfin Forest in the Bolivian Andes." Journal of Herpetology 32, no. 4 (1998): 536. http://dx.doi.org/10.2307/1565207.

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De la Riva, Ignacio. "BOLIVIAN FROGS OF THE GENUS PHRYNOPUS, WITH THE DESCRIPTION OF TWELVE NEW SPECIES (ANURA: BRACHYCEPHALIDAE)." Herpetological Monographs 21, no. 1 (2007): 241. http://dx.doi.org/10.1655/07-011.1.

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DUELLMAN, WILLIAM E., and JUAN CARLOS CHAPARRO. "Two distinctive new species of Pristimantis (Anura: Strabomantidae) from the Cordillera Oriental with a distributional synopsis of strabomantids in Central Peru." Zootaxa 1918, no. 1 (2008): 12–25. http://dx.doi.org/10.11646/zootaxa.1918.1.2.

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Two new species of the genus Pristimantis (Pristimantis) that are tentatively placed in the Pristimantis unistrigatus Group are described from the Yanachaga-Chemillén region in Departamento Pasco, Peru, where 24 species of strabomantid frogs are known. Both new species inhabit the humid montane forest on the slopes of the Andean Cordillera Oriental, and one of the new species apparently is a cryptic relative of another inhabitant of the region. Among 60 species of strabomantids recorded in a two-degree wide transect from the Amazonian lowland to the crest of the Cordillera Oriental, only three

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Lehr, Edgar, Cesar Aguilar, and Gunther Köhler. "Two Sympatric New Species of Phrynopus (Anura: Leptodactylidae) from a Cloud Forest in the Peruvian Andes." Journal of Herpetology 36, no. 2 (2002): 208–16. http://dx.doi.org/10.1670/0022-1511(2002)036[0208:tsnsop]2.0.co;2.

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MAMANI, LUIS, and SERGIO MALQUI. "A new species of Phrynopus (Anura: Craugastoridae) from the central Peruvian Andes." Zootaxa 3838, no. 2 (2014): 207. http://dx.doi.org/10.11646/zootaxa.3838.2.5.

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Lehr, Edgar, Guido Fritzsch, and Anke Muller. "Analysis of Andes frogs (Phrynopus, Leptodactylidae, Anura) phylogeny based on 12S and 16S mitochondrial rDNA sequences." Zoologica Scripta 34, no. 6 (2005): 593–603. http://dx.doi.org/10.1111/j.1463-6409.2005.00212.x.

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Trueb, Linda, and Edgar Lehr. "A new species of Phrynopus (Anura, Strabomantidae) from Peru, with comments on the osteology of the genus." Phyllomedusa: Journal of Herpetology 7, no. 1 (2008): 11. http://dx.doi.org/10.11606/issn.2316-9079.v7i1p11-24.

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Lehr, Edgar, Jiri Moravec, and Juan Carlos Cusi. "Two new species of Phrynopus (Anura, Strabomantidae) from high elevations in the Yanachaga-Chemillén National park in Peru (Departamento de Pasco)." ZooKeys 235 (October 31, 2012): 51–71. http://dx.doi.org/10.3897/zookeys.235.3885.

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Lehr, Edgar. "TAXONOMIC STATUS OF SOME SPECIES OF PERUVIAN PHRYNOPUS (ANURA: LEPTODACTYLIDAE), WITH THE DESCRIPTION OF A NEW SPECIES FROM THE ANDES OF SOUTHERN PERU." Herpetologica 62, no. 3 (2006): 331–47. http://dx.doi.org/10.1655/0018-0831(2006)62[331:tsosso]2.0.co;2.

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Lehr, Edgar, Rudolf von May, Jiri Moravec, and Juan Carlos Cusi. "A new species of Phrynopus (Amphibia, Anura, Craugastoridae) from upper montane forests and high Andean grasslands of the Pui Pui Protected Forest in central Peru." ZooKeys 713 (November 2, 2017): 131–57. http://dx.doi.org/10.3897/zookeys.713.20776.

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