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1

Navarrete, María, Pablo Venegas, and Santiago Ron. "Two new species of frogs of the genus Pristimantis from Llanganates National Park in Ecuador with comments on the regional diversity of Ecuadorian Pristimantis (Anura, Craugastoridae)." ZooKeys 593 (May 26, 2016): 139–62. https://doi.org/10.3897/zookeys.593.8063.

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We describe two new species of frogs of the genus Pristimantis from the eastern slopes of the Ecuadorian Andes, at Parque Nacional Llanganates. The new species are characterized by the spiny appearance typical of several species inhabiting montane forests. Pristimantis yanezi sp. n. is most similar to P. colonensis and P. incanus but differs from both in groin coloration and by having smaller tubercles on the upper eyelids, heels, and tarsus. Pristimantis llanganati sp. n. is most similar to P. eriphus and P. chloronotus. It can be distinguished from P. eriphus by the color pattern on the scap
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2

Ortega, Jhael A., Jorge Brito, and Santiago R. Ron. "Six new species of Pristimantis (Anura: Strabomantidae) from Llanganates National Park and Sangay National Park in Amazonian cloud forests of Ecuador." PeerJ 10 (October 17, 2022): e13761. http://dx.doi.org/10.7717/peerj.13761.

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We describe six new species of rainfrogs of the genus Pristimantis (Strabomantidae) from Amazonian cloud forests in Ecuador. We also present a phylogeny showing the relationships of the new species. The phylogeny is based on mitochondrial genes 16S rRNA (16S), 12 rRNA (12S), NADH-ubiquinone oxidoreductase chain 1 (ND1) and the nuclear gene recombination-activating 1 (RAG1). We also describe the osteology of two of the new species using high-resolution x-ray computed tomography. The new species belong to two clades. The first clade is sister to the subgenus Huicundomantis and includes P. tamia
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3

Catenazzi, Alessandro, and Edgar Lehr. "Pristimantis antisuyu sp. n. and Pristimantis erythroinguinis sp. n., two new species of terrestrial-breeding frogs (Anura, Strabomantidae) from the eastern slopes of the Andes in Manu National Park, Peru." Zootaxa 4394, no. 2 (2018): 185–206. https://doi.org/10.11646/zootaxa.4394.2.2.

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Catenazzi, Alessandro, Lehr, Edgar (2018): Pristimantis antisuyu sp. n. and Pristimantis erythroinguinis sp. n., two new species of terrestrial-breeding frogs (Anura, Strabomantidae) from the eastern slopes of the Andes in Manu National Park, Peru. Zootaxa 4394 (2): 185-206, DOI: 10.11646/zootaxa.4394.2.2
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4

Lehr, Edgar, and May Rudolf von. "A new species of terrestrial-breeding frog (Amphibia, Craugastoridae, Pristimantis) from high elevations of the Pui Pui Protected Forest in central Peru." ZooKeys 660 (March 7, 2017): 17–42. https://doi.org/10.3897/zookeys.660.11394.

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We describe a new species of Pristimantis from upper montane forests and high Andean grasslands of the Pui Pui Protected Forest and its close surroundings, Región Junín, central Peru. The description of the new species is based on 34 specimens found at elevations between 3400 and 3936 m a.s.l. Pristimantis attenboroughi sp. n. is characterized by a snout–vent length of 14.6–19.2 mm in adult males (n = 21), 19.2–23.0 mm in adult females (n = 10), and is compared morphologically and genetically with other taxonomically and biogeographically relevant species of Pristimantis. The new species is ch
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5

MEANS, D. BRUCE, and JAY M. SAVAGE. "Three New Malodorous Rainfrogs of the Genus Pristimantis (Anura: Brachycephalidae) from the Wokomung Massif in west-central Guyana, South America." Zootaxa 1658, no. 1 (2007): 39–55. http://dx.doi.org/10.11646/zootaxa.1658.1.3.

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Three new species of rainfrogs of the genus Pristimantis are described from a large mesa (tepui), the Wokomung Massif, of the Pakaraima Mountains in west-central Guyana. Pristimantis dendrobatoides n. sp. is known from 1385–1411 m, P. jester n. sp. from 1411–1650 m, and P. saltissimus n. sp. from 698–1560 m elevation. The three species are syntopic at elevations around 1400 m in cloud forest. All three taxa are unusual among species of Pristimantis in the production of malodorous and distasteful skin secretions when handled, conditions that are atypical for the genus. Two of the new species (P
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6

CATENAZZI, ALESSANDRO, and EDGAR LEHR. "Pristimantis antisuyu sp. n. and Pristimantis erythroinguinis sp. n., two new species of terrestrial-breeding frogs (Anura, Strabomantidae) from the eastern slopes of the Andes in Manu National Park, Peru." Zootaxa 4394, no. 2 (2018): 185. http://dx.doi.org/10.11646/zootaxa.4394.2.2.

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We describe two new species of Pristimantis from the Kosñipata valley in the eastern slopes of the Andes near Manu National Park, Peru. The two new species are closely related but do not overlap elevationally: P. antisuyu sp. n. occurs from 1485–1823 m a.s.l., whereas P. erythroinguinis sp. n. occurs from 930–1255 m a.s.l. Both species are readily distinguished from all other species of Pristimantis but P. cruciocularis and P. flavobracatus by possessing an iris with a cruciform pattern, no tympanum, and red bright or yellow coloration on groin. We used a Maximum Likelihood approach to infer a
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7

Lehr, Edgar, and Jiří Moravec. "A new species of Pristimantis (Amphibia, Anura, Craugastoridae) from a montane forest of the Pui Pui Protected Forest in central Peru (Región Junín)." ZooKeys 645 (January 12, 2017): 85–102. https://doi.org/10.3897/zookeys.645.11221.

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A new species of frog of the genus Pristimantis is described from a montane forest between 1700 and 1800 m a.s.l. of the Pui Pui Protected Forest (Región Junín) in central Peru. Pristimantis ashaninka sp. n. is described based on five adult females (snout–vent length 23.1–26.7 mm) and ten juveniles (snout-vent length 10.6–13.4). It differs from its congeners by having the skin on dorsum shagreen with many conical tubercles giving it a spinose appearance, lacking a tympanum, having groin, anterior and posterior surfaces of thighs uniformly grayish brown, and a pale bronze iris with fine black r
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8

Reyes-Puig, Carolina, Juan Pablo Reyes-Puig, Daniel A. Velarde-Garcéz, et al. "A new species of terrestrial frog Pristimantis (Strabomantidae) from the upper basin of the Pastaza River, Ecuador." ZooKeys 832 (March 19, 2019): 113–33. https://doi.org/10.3897/zookeys.832.30874.

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We describe a new species of Pristimantis from the montane forest of the Río Zuñag Ecological Reserve, upper basin of the Pastaza River, Ecuador. Pristimantis mallii sp. n. is characterized by a snout-vent length of 11.6–21.3 mm in adult males (n = 12), 22.6–34.3 mm in adult females (n = 8), and is compared morphologically and genetically with Pristimantis miktos and with other relevant species of Pristimantis. The new species is characterized by having skin on dorsum and flanks shagreen, distinctive scapular folds, snout broadly rounded in dorsal view, upper eyelid bearing one or two subconic
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9

Ttito, Alex, and Alessandro Catenazzi. "Pristimantis achupalla sp. n., a new minute species of direct-developing frog (Amphibia, Anura, Strabomantidae) inhabiting bromeliads of the montane forest of the Amazonian Andes of Puno, Peru." PeerJ 9 (September 23, 2021): e11878. http://dx.doi.org/10.7717/peerj.11878.

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We describe a new species of bromeliad-dwelling Pristimantis from primary montane forest (2,225 m a.s.l.) in southern Peru. The type locality is near Thiuni, in the Department of Puno (province of Carabaya) in the upper watershed of a tributary of the Inambari River. Pristimantis achupalla sp. n. is characterized by a snout-vent length of 10.0–12.8 mm in adult males (n = 4), unknown in adult females, and is compared morphologically and genetically with species in the Pristimantis lacrimosus group, and with other similar species of Pristimantis. The new species is characterized by having skin o
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10

VENEGAS, PABLO J., and WILLIAM E. DUELLMAN. "Two syntopic new species of the Pristimantis orestes Group (Anura: Strabomantidae) from northwestern Peru." Zootaxa 3249, no. 1 (2012): 47. http://dx.doi.org/10.11646/zootaxa.3249.1.5.

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Two new species of Pristimantis (Anura: Strabomantidae), tentatively assigned to the P. orestes Group, are described fromhumid puna (3596 m) of Cañaris, Provincia de Ferreñafe, Región Lambayeque on the eastern slopes of the Cordillera Oc-cidental in northwestern Peru. Pristimantis mariaelenae sp. n. is a medium-sized species characterized by the dorsumvarying from brown to reddish ochre, groin and posterior surface of the thigh with white blotches, pearly white venter withbrown blotches, distinct tympanic annulus and membrane, and tubercles on the eyelid, heel, and outer edge of the tarsus.Pri
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11

Shepack, Alexander, May Rudolf von, Alex Ttito, and Alessandro Catenazzi. "A new species of Pristimantis (Amphibia, Anura, Craugastoridae) from the foothills of the Andes in Manu National Park, southeastern Peru." ZooKeys 594 (May 30, 2016): 143–64. https://doi.org/10.3897/zookeys.594.8295.

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We describe a new species of Pristimantis from the humid sub-montane forest of the Región Cusco in Peru. Pristimantis pluvialis sp. n. was collected in the Kosñipata and Entoro valleys at elevations from 740 to 1110 m a.s.l., near the borders of Manu National Park and within the Huachiperi Haramba Queros Conservation Concession. The new species can be distinguished from other members of the genus Pristimantis by its rostral tubercle, smooth dorsal skin, and by its advertisement call. Pristimantis lacrimosus and P. waoranii superficially most resemble the new species, but P. pluvialis sp. n. di
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12

LEHR, EDGAR, RUDOLF VON MAY, JIŘÍ MORAVEC, and JUAN CARLOS CUSI. "Three new species of Pristimantis (Amphibia, Anura, Craugastoridae) from upper montane forests and high Andean grasslands of the Pui Pui Protected Forest in central Peru." Zootaxa 4299, no. 3 (2017): 301. http://dx.doi.org/10.11646/zootaxa.4299.3.1.

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We describe three new species of Pristimantis from the upper montane forests and high Andean grasslands of the Pui Pui Protected Forest and its close surroundings (Región Junín, central Peru) and compare them morphologically and genetically with other taxonomically and biogeographically relevant species of Pristimantis. All three new species have the skin on dorsum shagreen with scattered tubercles, discontinuous dorsolateral folds, tuberculate flanks, and the skin on venter areolate. Pristimantis bounides sp. nov. is known from two localities outside the Pui Pui Protected Forest in upper mont
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13

Lehr, Edgar, Jiri Moravec, and Juan Carlos Cusi. "Two new species of Phrynopus (Anura, Strabomantidae) from high elevations in the Yanachaga-Chemillén National park in Peru (Departamento de Pasco)." ZooKeys 235 (October 31, 2012): 51–71. https://doi.org/10.3897/zookeys.235.3885.

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We describe two new species of <i>Phrynopus</i> from a cloud forest of the Cordillera Yanachaga, Yanachaga-Chemillén National Park in Peru and describe and document the first clutch and case of parental care for a species of <i>Phrynopus</i>. One of the new species of <i>Phrynopus </i>is described based on two females (SVL 19.1–21.0 mm) which were found in leaf litter and moss layer in a cloud forest at 2900 m elevation. This new species is most similar to <i>Phrynopus bracki</i>. The second new species of <i>Phrynopus </i>was found in the transitional formation between cloud forest and wet pu
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14

Oliveira, Elciomar Araújo De, Luis Reginaldo Rodrigues, Igor Luis Kaefer, Karll Cavalcante Pinto, and Emil José Hernández-Ruz. "A new species of Pristimantis from eastern Brazilian Amazonia (Anura, Craugastoridae)." ZooKeys 687 (August 2, 2017): 101–29. https://doi.org/10.3897/zookeys.687.13221.

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In this study a new species of Pristimantis (Anura: Craugastoridae) of the P. conspicillatus species group is described. Pristimantis latro sp. n. is known only from the municipalities of Altamira, Anapu, Brasil Novo, Medicilândia, Uruará and Aveiro (Flona Tapajós, right bank of Tapajós river), in Pará state, Brazil. Morphologically, the new species distinguishes from known congeners in the group mainly by the presence of dorsal tubercles and absence of discoidal folds, smooth belly skin, as well as the presence of supernumerary tubercles on hands. The call of the new species consists of seven
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15

Sánchez-Nivicela, Juan C., Elvis Celi-Piedra, Valentina Posse-Sarmiento, Verónica L. Urgilés, Mario Yánez-Muñoz, and Diego F. Cisneros-Heredia. "A new species of Pristimantis (Anura, Craugastoridae) from the Cajas Massif, southern Ecuador." ZooKeys 751 (April 20, 2018): 113–28. https://doi.org/10.3897/zookeys.751.20541.

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A new species of Pristimantis is described from the highland paramos on the eastern slopes of the Cajas Massif, southern Andes of Ecuador, at 3400 m. This new species is characterized by having a distinctive reddish color, cutaneous macroglands in suprascapular region and surfaces of arm and legs, and by lacking dentigerous processes of vomers. The cutaneous macroglands are similar to those exhibited by several species of the Pristimantis orcesi group, and may suggest a close phylogenetic relationship. The new species could be a latitudinal substitution of Pristimantis orcesi in the southern A
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16

Ramírez-Jaramillo, Reyes-Puig, Batallas, and Yánez-Muñoz. "Ranas terrestres en los ecosistemas surandinos de Ecuador IV: Una nueva especie de rana terrestre (Anura: Craugastoridae) de los páramos sur del Parque Nacional Sangay." ACI Avances en Ciencias e Ingenierías 10, no. 1 (2018): 1–18. https://doi.org/10.18272/aci.v1i1.841.

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We describe a new species of terrestrial frog of the genus&nbsp;<em>Pristimantis</em>, located in the south Andes of Ecuador. The new species is characterized by: presence of dentigerous processes of vomer that are triangular in outline; ulnar tubercles flattened and disposed in a longitudinal, fold-like row; and a distinctive ventral coloration light to dark brown with white warts. We describe the call of an adult male, a single note with an average frequency of 4.23 kHz and two partial harmonics, and an average duration of 207 ms. Describimos una nueva especie de rana terrestre&nbsp;<em>Pris
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17

Székely, Paul, Dan Cogălniceanu, Diana Székely, Nadia Paez, and Santiago Ron. "A new species of Pristimantis from southern Ecuador (Anura, Craugastoridae)." ZooKeys 606 (July 21, 2016): 77–97. https://doi.org/10.3897/zookeys.606.9121.

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A new species of Pristimantis is described from Reserva Buenaventura, southern Ecuador, at elevations between 878 and 1082 m. A molecular phylogeny based on nuclear and mitochondrial genes shows that the new species is closely related to P. phoxocephalus, P. riveti, and P. versicolor. The new species differs from them and other morphologically similar congeners in having a low W-shaped dermal ridge in the scapular region, a large conical tubercle on the upper eyelid and on the heel, a thin mid dorsal fold, and a longitudinal lateral fold starting behind the tympanic fold and extending along th
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18

HEINICKE, MATTHEW P., WILLIAM E. DUELLMAN, LINDA TRUEB, D. BRUCE MEANS, ROSS D. MacCULLOCH, and S. BLAIR HEDGES. "A new frog family (Anura: Terrarana) from South America and an expanded direct-developing clade revealed by molecular phylogeny." Zootaxa 2211, no. 1 (2009): 1–35. http://dx.doi.org/10.11646/zootaxa.2211.1.1.

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Three frogs of a new species found in cloud forests on two nearby mountains in Guyana were included in a molecular phylogeny of 17 nuclear and mitochondrial genes (10,739 aligned sites) that revealed that their closest relative is Terrarana (Brachycephalidae, Craugastoridae, Eleutherodactylidae, and Strabomantidae) and their next-closest relative is Hemiphractidae (marsupial frogs). We place these frogs in a new family, genus, and species which is strongly supported as the basal clade within Terrarana: Ceuthomantidae n. fam., Ceuthomantis smaragdinus n. gen, n. sp. Morphological evidence suppo
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19

Lehr, Edgar, Jiří Moravec, Juan Carlos Cusi, and Václav Gvoždík. "A new minute species of Pristimantis (Amphibia: Anura: Craugastoridae) with a large head from the Yanachaga-Chemillén National Park in central Peru, with comments on the phylogenetic diversity of Pristimantis occurring in the Cordillera Yanachaga." European Journal of Taxonomy, no. 325 (June 2, 2017). http://dx.doi.org/10.5852/ejt.2017.325.

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We describe a new minute species of the genus Pristimantis, P. boucephalus sp. nov., from the Yanachaga-Chemillén National Park, Región Pasco, Peru. The description is based on a freshly collected male specimen found at 2950 m a.s.l. in a cloud forest and four previously unidentified museum specimens consisting of two adult males, one subadult female and a juvenile from the Yanachaga-Chemillén National Park. The new species is mainly characterized by a snout–vent length of 13.4–14.5 mm in adult males (n = 3), and 12.5 mm in the only known subadult female, and is compared morphologically and ge
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