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1

Sangster, Sarah. "The anatomy, functional morphology and systematics of Dimorphodon macronyx (Diapsida: Pterosauria)." Thesis, University of Cambridge, 2003. http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.620016.

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2

Cavalcanti, Vila Nova de Albuquerque Bruno. "Preparação, re-descrição e posicionamento filogenético de Cearadactylus atrox Leonardi & Borgomanero, 1985 (Archosauria, Pterosauria)." Universidade Federal de Pernambuco, 2010. https://repositorio.ufpe.br/handle/123456789/6041.

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Coordenação de Aperfeiçoamento de Pessoal de Nível Superior
Pterossauros são um dos grupos de arcossauros fósseis melhor representados no registro fóssil brasileiro. Dentro desta diversidade, uma das primeiras espécies descritas, Cearadactylus atrox, tem causado divergências de interpretação entre pesquisadores desde a sua publicação. Devido ao fato de o holótipo e único espécime conhecido estar parcialmente preparado, as tentativas de classificações taxonômicas realizadas, posicionaram-no em grupos diferentes. Estas discordâncias geraram o colapso de certos ramos da filogenia de Pterosauria. O presente trabalho teve como objetivo a preparação do holótipo de Cearadactylus atrox, sua re-descrição e o posicionamento filogenético, utilizando-se matrizes filogenéticas disponíveis para o grupo. Evidências de adulterações foram observadas durante a preparação. Dentre as características exclusivas apresentadas por Cearadactylus atrox estão uma fenda no dentário bifurcada na extremidade rostral, órbita e narina em posição elevada relativo a fenestra nasoanterorbital e menos de 15 dentes em cada lado da maxila e mandíbula. Filogeneticamente Cearadactylus atrox se mostrou como grupo irmão do clado Anhangueridae, sendo uma espécie intermediária entre as formas européias (nominalmente Ornithocheirus compressirostris) e as formas brasileiras (Tropeognathus mesembrinus e as espécies que compõem o gênero Anhanguera)
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BANTIM, Renan Alfredo Machado. "Preparação, descrição de um novo crânio de pterossauro (Reptilia, Archosauria) e considerações sobre a morfologia craniana dos Anhangueridae." Universidade Federal de Pernambuco, 2013. https://repositorio.ufpe.br/handle/123456789/10470.

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CNPq
Os pterossauros são um grupo extinto de répteis alados que surgiu há pelo menos 228 milhões de anos, sendo encontrados no Brasil em rochas triássicas do Paraná e cretáceas do Nordeste. Dentro deste registro, o clado Anhangueridae, é documentado nas concreções calcárias da Formação Romualdo, Aptiano/Albiano da Bacia do Araripe. A esse grupo atribuímos um novo espécime LPU 017, um crânio completo preservado tridimensionalmente, com alguns ossos deslocados e a presença de dentes. Após sua preparação destacou-se uma grande e alta crista sagital pré-maxilar, característica distintiva deste grupo de pterossauros, além das características exclusivas deste exemplar. Dentre elas estão a presença de 35 pares de alvéolos; uma quilha palatal relativamente robusta; uma elevada expansão em forma de colher na pré-maxila; a presença do 5º, 6º e 7º pares de dentes menores do que o 4º e 8º; forma convexa do palato na porção rostral e o tamanho e localização da crista sagital pré-maxilar. Tais características exclusivas permitiram identificar LPU 017 como um novo gênero e espécie de pterossauro atribuído ao clado Anhangueridae. Esse resultado foi corroborado pela análise filogenética, obtida com uma matriz de 54 táxons e 89 caracteres, que gerou 9 árvores igualmente parcimoniosas. Para verificar os padrões de diferenciação craniana no clado Anhangueridae, utilizamos a morfometria geométrica, que compara a forma dos organismos, levando em consideração o caráter geométrico das formas biológicas e analisa estatisticamente sua variação. Verificamos então que crânios e cristas nesse clado crescem de forma isométrica, o oposto da realidade dos membros posteriores, onde o crescimento ósseo é alométrico. Nesse clado em especial a ontogenia não interferiu nos resultados, pois crista e crânio seguiram o mesmo padrão de crescimento, inclusive em indivíduos de estágios ontogenéticos diferentes, como ocorreu no agrupamento de Anhanguera araripensis e Anhanguera blittersdorffi
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4

Aires, Alex Sandro Schiller. "Descrição de Novos Materiais Referentes à Tapejaridae (pterosauria, Pterodactyloidea) da Formação Romualdo (Bacia do Araripe, Ceará, Brasil)." Universidade Federal do Pampa, 2013. http://dspace.unipampa.edu.br:8080/xmlui/handle/riu/229.

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O clado Pterosauria é constituído por répteis alados que viveram durante a Era Mesozóica (entre 215 a 65 milhões de anos antes do presente) e habitaram diversas regiões do planeta. Pertencente a este clado, o g rupo Tapejaridae é caracterizado principalmente pela peculiar morfologia rostral e por não possuir dentes. O estudo filogenético de tapejarídeos do nordeste do Brasil é baseado na sua maioria em elementos do crânio, devido à escassez de esqueletos pós-cranianos. Este estudo descreve morfologicamente e classifica, através de análise filogenética, um novo espécime atribuído à Tapejaridae, proveniente da Formação Romualdo (Grupo Santana, Bacia do Araripe, CE). O material compreende dez vértebras articuladas (as três últimas cervicais e também as sete primeiras dorsais que se fusionam, formando o notário), o escapulocoracóide direito, a escápula e o coracóide esquerdos, um fragmento de esterno, o úmero direito parcialmente preservado, um fragmento da 4a falange do dígito alar, a porção distal do femur direito e a porção proximal da tíbia/fíbula . Foram geradas três árvores na análise filogenética, sendo que em duas delas AMNH 22567 foi posicionado dentro de Thalassodrominae . Contudo, na outra árvore o espécime se pocisionou como Tapejaridae basal. Mesmo assim, sua assignação como Thalassodrominae se justifica pela presença de três foramens posicionados dorsal e lateralmente ao canal neural nas vértebras cervicais; presença de notário e presença de um forâmen pneumático ventral na porção proximal do úmero. Estes materiais são os mais completos elementos pós-cranianos deste grupo de pterossauros descritos até o momento. Com relação à paleobiogeografia, Tapejaridae aparenta ter surgido no oeste da Eurásia durante o Neojurássico, migrando para sudoeste e dando origem as formas eocretácicas da Bacia do Araripe.
Pterosauria is a clade composed of flying reptiles that lived during the Mesozoic Era (215 to 65 million of years before present) and inhabited various regions of the world. Within this large clade, Tapejaridae is mainly characterized by peculiar rostral morphology and absence of tooths. The phylogenetical studies of the tapejarids from northeastern Brazil are mainly based on cranial elements, due to shortage of postcranial skeletons. This study described and classified through a phylogenetical analysis, a new specimen ascribed to Tapejaridae from Romualdo Formation (Araripe Basin, CE), housed at the American Museum of Natural History (under the acronym AMNH 22567) and comments on the paleobiogeography of this group. The material comprises ten articulated vertebrae (three cervicals and seven subsequent dorsals fusioned, ones forming the notarium), a right scapulocoracoid, a left scapula and a left coracoid, a partial sternum, a right humerus, a fragment of 4th phalanx of the alar digit, a distal fragment of right femur and a fragment of the proximal portion of the right tibia/fibula. The phylogenetical analysis generated three different topologies, two of them placing AMNH 22567 within Thalassodrominae (subclade of Tapejaridae). In the other one, the specimen was placed in a basal position within Tapejaridae. Nonetheless, the presence of three foramina lateral and dorsal to the neural canal of the cervical vertebrae, the presence of a notarium and presence of a pneumatic foramen on the ventral side of the proximal portion of the humerus allow its assignation as a Thalassodrominae indet. The specimen comprises the more complete postcranial material for this subfamily described so far. Regarding the paleobiogeography of Tapejaridae, it appears to originated in western Eurásia during the Neojurassic and later migrated to southwest and irradiated to new Eocretacic forms in Araripe Basin.
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Pinheiro, Felipe Lima. "Sobre um novo espécime de Tupandactylus Imperator (Archosauria, Pterosauria), proveniente da formação Crato (Eocretáceo) da Bacia do Araripe, Ceará, nordeste do Brasil." reponame:Biblioteca Digital de Teses e Dissertações da UFRGS, 2011. http://hdl.handle.net/10183/32684.

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No ano de 2009, em uma mineradora de calcário laminado próxima à cidade de Nova Olinda (Ceará, nordeste do Brasil), um crânio parcial de pterossauro foi encontrado. O novo material, objeto de estudo do presente trabalho, é proveniente de estratos eocretácicos da Formação Crato (Bacia do Araripe). Trata-se do quarto espécime conhecido de Tupandactylus imperator (Pterosauria, Tapejaridae) e o primeiro com mandíbula associada. O estudo deste espécime contribuiu significativamente com informações a respeito da anatomia e relações filogenéticas de T. imperator e outros pterossauros tapejarídeos, resultando na publicação de um artigo científico. No presente trabalho, além da descrição anatômica detalhada e estudo filogenético do novo exemplar, revisamos a ocorrência de pterossauros no Grupo Santana da Bacia do Araripe, com ênfase em T. imperator. Questões taxonômicas, como as relações filogenéticas de pterossauros tapejarídeos e a validade de Tupandactylus navigans são, também, discutidas.
In the year of 2009 a partial pterosaur skull was found in a limestone quarry near the city of Nova Olinda (Ceará State, northeastern Brazil). The new material, object of study of the present work, comes from the Early Cretaceous strata of Crato Formation, Araripe Basin, being the fourth known specimen of Tupandactylus imperator (Pterosauria, Tapejaridae) and the first one with associated lower jaw. The study of this specimen added new significant information on T. imperator anatomy and phylogenetic relationships, and resulted in the publishing of a scientific paper. At the present work we describe the new specimen in detail, study its phylogenetic relationships and review pterosaur occurrence in the Santana Group of Araripe Basin (with emphasis on T. imperator). Taxonomic issues as Tapejaridae relationships and the validity of Tupandactylus navigans are also discussed.
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Baron, Matthew Grant. "The origin and early evolution of the Dinosauria." Thesis, University of Cambridge, 2018. https://www.repository.cam.ac.uk/handle/1810/271890.

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For 130 years dinosaurs have been divided into two distinct clades – Ornithischia and Saurischia. This dissertation looks at the earliest evolution of the clade Dinosauria by focusing upon the interrelationships of the major subsidiary clades within it. It does this following examination, comparison and description of early dinosaur material, and by utilising modern phylogenetic analysis techniques, to rigorously and objectively test the fundamental groupings within the clade Dinosauria using a newly compiled dataset of early dinosaurs and other dinosauromorphs (= close dinosaur relatives). The current consensus on how the principal clades within Dinosauria (Theropoda, Sauropodomorpha and Ornithischia) are related to one another is challenged by the results of these analyses. This study finds, for the first time, a sister-group relationship between Ornithischia and Theropoda, here termed Ornithoscelida. Consequently, a new definition for Dinosauria is presented, as the historic definition would exclude all members of Sauropodomorpha from the clade. As well as this, I propose revisions to the definitions of each of the principal dinosaurian sub-divisions and propose a new timeframe and geographic setting for the origin of Dinosauria. These new hypotheses force re-evaluations of early dinosaur cladogenesis and character evolution, suggest the independent acquisition of hypercarnivory in multiple dinosaur groups and offers an explanation for many of the anatomical features previously regarded as striking convergences between theropods and early ornithischians. As well as presenting new anatomical data on many early dinosaurs and dinosauromorphs, including a comprehensive re-description of the postcranial anatomy of Lesothosaurus diagnosticus (Chapter 2), and a new anatomical dataset of early dinosaurs (the largest ever compiled), this thesis goes on to implement the new dataset to investigate a number of important outstanding questions about early dinosaur evolution and provides new lines of enquiry for future workers to pursue. The results of this thesis reveal the oldest known members of the dinosaurian clades Theropoda and Sauropodomorpha (Chapter 6), as well as a new clade within Ornithischia; a taxon previously thought to represent a derived theropod has been recovered as a potential ‘missing link’ between theropods and ornithischians using the new dataset. This work now provides a unique tool for the assessment of the phylogenetic affinities of early dinosaurs and dinosauromorphs and, once published, will hopefully become the benchmark dataset for palaeontologists working in this area.
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Wilkinson, Matthew Thomas. "Flight of the ornithocheirid pterosaurs." Thesis, University of Cambridge, 2003. http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.619589.

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8

Vidovic, Steven Uroš. "A discourse on pterosaur phylogeny." Thesis, University of Portsmouth, 2016. https://researchportal.port.ac.uk/portal/en/theses/a-discourse-on-pterosaur-phylogeny(4d366e00-23fa-4ee9-9ae1-104e7409dfd5).html.

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The Pterosauria is an enigmatic group of volant reptiles from the Mesozoic. Despite demonstrating conspicuous apomorphic conditions, including an enlarged humeral deltopectoral crest, elongate wing spar comprising the fourth manual digit, and unique skull bone configurations, their cladogenesis is uncertain. While the ingroup monophyly of Pterosauria is clear, their interrelationships are contentious. The earliest studies of pterosaur phylogeny were concerned with phenotypic groups. The earliest cladistic analyses of pterosaurs in the late 20th Century were simple and were superseded by more comprehensive analyses in the early 21st Century. These analyses are comparable at the genus, family, and suborder level, but the ranks between these are incongruent between authors, leading to nomenclatural disputes. This study aims to understand the differences that exist between the competing phylogenetic hypotheses, why they exist, and to establish a more robust phylogenetic hypothesis. An extensive review of pterosaur phylogeny, cladistic character coding, and data transformation is performed. A distinct cladistic analysis is developed independently of the literature, providing a reference tree for a meta-analysis of topological congruence. Furthermore, this distinct analysis is supplemented with published characters to produce a comprehensive analysis which is a robust hypothesis of pterosaur phylogeny. Taxonomic actions are taken on the wastebasket taxon Pterodactylus and the paraphyletic genus Germanodactylus, to improve the taxonomic content of the analyses. The results demonstrate that analyses sharing characters produce trees with greater levels of congruence. Although this may be due to the introduction of biases, it is far more likely that the improvement seen in the comprehensive tree at least, is due to improving the phyletic content, sampling of anatomy, and reducing the negative effects of homoplasy. This is supported by the comprehensive tree's congruence to the stratigraphic record. The comprehensive tree agrees with the common clades of other studies, but some families are found in unique positions, most notably the Anurognathidae.
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Witton, Mark P. "The palaeoecology and diversity of pterosaurs." Thesis, University of Portsmouth, 2008. http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.496607.

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The taxonomy and locomotory methods of pterosaurs have been extensively scrutinised for as long as pterosaurs themselves have been known, but comparatively little research has been conducted into their lifestyles and habits. Pterosaurs have largely been interpreted as Mesozoic equivalents of modem marine birds, principally foraging through flight-based methods of dip- or skim-feeding. Here, several lines of enquiry suggest that pterosaurs were considerably more ;ologically diverse that previously anticipated.
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O'Sullivan, Michael. "The taxonomic diversity of British Jurassic pterosaurs." Thesis, University of Portsmouth, 2015. https://researchportal.port.ac.uk/portal/en/theses/the-taxonomic-diversity-of-british-jurassic-pterosaurs(ad180383-6d6c-425b-ad01-4716e2ba112c).html.

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The taxonomic diversity of Jurassic pterosaurs is more poorly understood than that of their Cretaceous counterparts. Early to Middle Jurassic pterosaur-bearing formations are rare and suffer from a low fossil yield, resulting in an apparent Late Jurassic diversity spike. Recent studies have suggested this is a preservational bias rather than a taxonomic signal, however this is difficult to test given the low number of sizeable Jurassic pterosaur collections. The United Kingdom possesses Lower, Middle and Upper Jurassic pterosaur bearing horizons containing hundreds of fossils, making it one of the best Jurassic assemblages suitable for testing taxic diversity. Lower Jurassic deposits remain low yield but include the holotypes of Dimorphodon and Parapsicephalus. Parapsicephalus has previously been synonymised with Dorygnathus but is found to be a unique genus. An isolated humerus is identified as a campylognathoidid, marking the first occurrence of the group in Britain. The Middle Jurassic Taynton Limestone Formation (informally known as the Stonesfield Slate) has yielded more than 400 isolated pterosaur fossils with only one named genus, Rhamphocephalus. The type of Rhamphocephalus, is identified here as a teleosaurid skull table with the genus considered a nomen dubium. A new genus is erected for an isolated mandible and the Taynton Limestone Formation is found to include at least 3 pterosaur taxa. The Middle-Upper Jurassic Oxford Clay Formation is found to have a low diversity with the only named genus, Rhamphorhynchus jessoni, identified as a nomen dubium. The Upper Jurassic Kimmeridge Clay Formation has produced at least 4 pterosaurs, including a new species of Rhamphorhynchus. These findings suggest that Jurassic pterosaur diversity has been underestimated and is most likely higher than traditionally believed.
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Roberts, Brian C. "Turn performance and flight dynamics of a pterosaur and a pterosaur-inspired variable-placement vertical tail aircraft." [Gainesville, Fla.] : University of Florida, 2009. http://purl.fcla.edu/fcla/etd/UFE0024421.

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Steel, Lorna. "Studies on the palaeohistology of pterosaur bone." Thesis, University of Portsmouth, 2004. http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.416203.

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Hazlehurst, Grant Allen. "The morphometric and flight characteristics of the pterosaurs." Thesis, University of Bristol, 1991. http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.303761.

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Martin-Silverstone, Elizabeth Gwendolyn. "Insights into mass estimation, pneumaticity, and anatomy of pterosaurs : implications for locomotion." Thesis, University of Southampton, 2017. https://eprints.soton.ac.uk/416889/.

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Pterosaurs were both the first and largest vertebrates to achieve powered flight, surviving for over 150 million years, and ranging in size from 0.5-11m wingspans. This large size was achieved through a number of adaptations, including a membraneous wing supported by a single elongated finger, a strongly developed forelimb, and a pneumatised skeleton with hollow bones. This thesis aims to quantify aspects of pterosaur pneumaticity, mass estimation, and wing bone morphology and deduce how these relate to pterosaur locomotion and flight. Using computed tomography (CT), the internal structure of pterosaur bones is visible, and the degree of pneumaticity is compared to other animals, indicating large pterosaurs were among the most pneumatised animals. A large database of pterosaur wing bone geometry and cross-sections shows that small pterosaurs were biomechanical generalists, resistant to loads both common in flight and terrestrial locomotion, while most larger pterosaurs sacrificed terrestrial capabilities in favour of low mass and resisting increased loads in flight. To better understand pterosaur body mass, a study of avian mass found that the relationship between skeletal mass and total mass may be accurate in neornithine birds, but should not be expanded to pterosaurs. Related, validation tests show that CT scans can be used to accurately estimate bone mass and volume, which can be incorporated into pterosaur mass studies. Building on this, a 3D skeletal reconstruction of an individual Coloborhynchus was used in a detailed estimate of the volume and mass of a single individual, in order to compare results from different methods. Although arguably subjective, 3D reconstruction more accurately represents the soft tissue than a minimum convex hull method, and is better at estimating mass than skeletal correlates such as humeral circumference. Finally, investigation of large pneumatic foramina in the sacrum of a small-bodied pterosaur lead to the identification of spinal nerve foramina in the first study of pterosaur postcranial neurology. Differences in neural canal size reveal potential patterns in locomotion, with some animals having highly innervated hindlimbs, while others have relatively poor innervation. This thesis provides new information on mass estimation in pterosaurs, pneumaticity, wing bone geometry, and locomotory patterns. Together, these results show that while some pterosaurs were the most pneumatic animals, there is significant variability, overturning conventional wisdom. This builds on previous ideas of pterosaur locomotion, giving more evidence to the idea that most larger, more-derived pterosaurs were optimised for flight, while smaller, more-basal pterosaurs were generalists.
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Fastnacht, Michael. "Jaw mechanics of the pterosaur skull construction and the evolution of toothlessness." [S.l.] : [s.n.], 2005. http://deposit.ddb.de/cgi-bin/dokserv?idn=97781257X.

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Elgin, Ross Alexander [Verfasser], and Wolfgang [Akademischer Betreuer] Stinnesbeck. "Palaeobiology, Morphology, and Flight Characteristics of Pterodactyloid Pterosaurs / Ross Alexander Elgin ; Betreuer: Wolfgang Stinnesbeck." Heidelberg : Universitätsbibliothek Heidelberg, 2015. http://d-nb.info/1180299698/34.

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Unwin, D. M. "The morphology, systematics and evolutionary history of pterosaurs from the Cretaceous Cambridge Greensand of England." Thesis, University of Reading, 1991. http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.505670.

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Hing, Richard. "A re-examination of a specimen of pterosaur soft tissue from the Cretaceous Santana formation of Brazil." Thesis, University of Portsmouth, 2011. http://eprints.port.ac.uk/8029/.

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Multiple pterosaur specimens have revealed details of their soft tissue anatomy, including wing membranes supported internally by strengthening fibres (aktinofibrils), a hair-like integument across their bodies, necks and heads (pycnofibres), elaborate soft-tissue head crests, tail vanes and other structures. These have enabled a more comprehensive picture of them as living animals than skeletal remains alone could, though many aspects of the structure and composition of these soft tissues remain controversial. The histology of pterosaur wing membranes has been a particularly contentious issue, with much debate focused on an alleged portion of wing membrane (DGM 1475-R) from the Lower Cretaceous Santana Formation of northeast Brazil. Initially described as a portion of wing membrane from the proximal wing region by Martill and Unwin (1989), after re-examination of the same specimen Kellner (1996) argued that it represented a piece of body wall. With DGM 1475-R showing features not seen in other specimens of pterosaur wing membrane and being one of the few three-dimensionally preserved portions of pterosaur membrane known, resolving this controversy is key to understanding this important component of pterosaur flight anatomy. In this study, DGM 1475-R was re-examined in an attempt to clarify the identity of its soft-tissues. Interpretation of the humerus and several ribs associated with the soft-tissue tentatively suggests that the specimen represents a thalassodromid pterosaur. A new section was taken through the soft-tissues and acid etched to highlight histological features. Multiple layers were identified within the soft tissue, including an external epidermis and a layer of muscle. The structures between these layers - numerous objects that appear to be fibres larger than the undoubted muscle fibres - are more ambiguous. These were previously identified as aktinofibrils (Martill et al., 1990), but this is considered unlikely due to differences in the shape, density and distribution of the structures when compared to aktinofibrils in other pterosaur wings. Instead, it is tentatively suggested that they represent a further layer of muscle fibres, but this cannot be conclusively proven. Further comparisons were made between DGM 1475-R and a dissected bat wing to find additional diagnostic wing tissues, but they were found to have such fundamental histological differences that such comparisons were of little use. Aktinofibrils are the only known diagnostic features of pterosaur wings, suggesting that their absence in DGM 1475-R argues against the specimen stemming from the wing tissues. Interpretations of pterosaur wing membrane and flight based on DGM 1475-R being wing membrane are therefore suspect.
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Jones, Terry D. "New insight into the morphology and physiology of Mesozoic archosaurs." Thesis, 2000. http://hdl.handle.net/1957/33248.

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Inferences regarding the biology of extinct taxa are often founded primarily on cladistically-based phylogenetic hypotheses. However, these inferences are frequently inconsistent with the morphology and physiology of modern animals. I suggest that reference to the biology of modern animals might be a more appropriate tool to aid in the reconstruction of the morphology and physiology of extinct animals. Here, this methodology is applied to: (1) reconstruction of the respiratory anatomy and physiology of some Paleozoic archosaurs (dinosaurs, early birds, pterosaurs, and early crocodilians), (2) understanding cursorial locomotion in birds and bipedal dinosaurs, and (3) a reinterpretation of an enigmatic archosaur, Longisquama. The conclusions drawn from these data raise questions regarding current thinking regarding relationships between birds and dinosaurs.
Graduation date: 2000
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Fernandes, Alexandra Eugénio. "Pterosaurs from the Late Cretaceous of Angola." Master's thesis, 2020. http://hdl.handle.net/10362/103156.

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Fourteen fossil bones from the Lower Maastrichtian marine locality of Bentiaba in the Namibe Basin of Angola, Africa, identified as pterosaurs, are here described and attributed to Pteranodontia. One articulated set of bones from within the fossil assemblage can be further attributed to the Nyctosauridae, gen. et sp. nov. This is the first record of pterosaur material from the central west coast of Africa, contributing new information about the paleoecological environment of Gondwana during this time frame. Ontogenetic deductions are also drawn from preliminary histological work. This fossil assemblage provides a first glimpse of Angolan pterosaur paleobiodiversity, giving further insight into the ecosystems of the Late Cretaceous.
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Fastnacht, Michael [Verfasser]. "Jaw mechanics of the pterosaur skull construction and the evolution of toothlessness / Michael Fastnacht." 2005. http://d-nb.info/97781257X/34.

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