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1

Deschodt, Christian, Adrian Davis, and Clarke Scholtz. "New dung beetle (Coleoptera: Scarabaeidae: Scarabaeinae) taxa from arid south-western Africa." Insect Systematics & Evolution 42, no. 3 (2011): 277–94. http://dx.doi.org/10.1163/187631211x568173.

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AbstractIn this paper we describe new dung beetle taxa (Coleoptera: Scarabaeinae) and discuss others previously described from arid south-western Africa. Descriptions are provided for a new monotypic genus (Versicorpus Deschodt, Davis and Scholtz gen.n.) and three new species, Versicorpus erongoensis Deschodt, Davis and Scholtz sp.n., Namakwanus endroedyi Deschodt, Davis and Scholtz, sp.n., and Scarabaeus (Kheper) kalaharicus Davis, Deschodt and Scholtz, sp.n. The type material of Scarabaeus (Scarabaeus) hottentorum Péringuey is examined and spelling errors on the labels are discussed. Inconsistencies between the published type locality, type locality labels, and the known geographical range are also discussed. The recently described Kheper namibicus Krajcik 2006 is synonymised with Scarabaeus (Kheper) cupreus (Castelnau 1840). Distribution records are provided for the new species described here while some closely related species are discussed.
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2

NARDI, GIANLUCA, GIOVANNI DELLACASA, and MARCO DELLACASA. "On the type species of Geophilus Gistel, 1834 (Coleoptera: Scarabaeidae: Aphodiinae)." Zootaxa 4461, no. 1 (2018): 83. http://dx.doi.org/10.11646/zootaxa.4461.1.6.

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Geophilus Gistel, 1834 (Insecta: Coleoptera: Scarabaeidae: Aphodiinae) is invalid being a junior homonym of Geophilus Leach, 1814 (Myriapoda: Chilopoda: Geophilomorpha: Geophilidae) and of Geophilus Schönherr, 1823 (Insecta: Coleoptera: Curculionidae). Scarabaeus asper Fabricius, 1775 was recently designated as type species of Geophilus Gistel, 1834, making Geophilus a junior synonym of Psammodius Fallén, 1807 (Insecta: Coleoptera: Scarabaeidae: Aphodiinae: Psammodiini: Psammodiina). However, there is clear evidence that Scarabaeus asper Fabricius, 1775 sensu Gistel, 1834 is Ptinus germanus Linnaeus, 1767, currently Rhyssemus germanus (Linnaeus, 1767) (Insecta: Coleoptera: Scarabaeidae: Aphodiinae: Psammodiini: Rhyssemina). To resolve this issue, the type species of Geophilus Gistel, 1834 is here fixed (under Article 70.3.2 of the International Code of Zoological Nomenclature) as Ptinus germanus Linnaeus, 1767, misidentified as Scarabaeus asper Fabricius, 1775 in the original paper. Scarabaeus asper Fabricius, 1775 sensu Mulsant, 1842 (= Ptinus germanus Linnaeus, 1767) is the type species of Rhyssemus Mulsant, 1842, therefore Rhyssemus Mulsant, 1842 is a junior synonym of Geophilus Gistel, 1834 (new synonymy). Although it has priority, Geophilus Gistel, 1834 is a junior homonym and therefore invalid, so Rhyssemus Mulsant, 1842 remains the valid name of this taxon.
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3

Branco, Tristão, Giovanni Dellacasa, and Marco Dellacasa. "SCARABAEUS SEROTINUS PANZER, 1799 (CURRENTLY NOBIUS SEROTINUS) A VALID NAME (NOMEN PROTECTUM) (COLEOPTERA, APHODIIDAE)." Fragmenta Entomologica 40, no. 1 (2008): 93. http://dx.doi.org/10.4081/fe.2008.106.

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Il nome <em>Scarabaeus serotinus</em> Panzer, 1799 (attualmente <em>Nobius</em> <em>serotinus</em>), ad oggi usato correntemente come nome valido, risulta sinonimo juniore di <em>Scarabaeus minu­tus</em> Herbst, 1783. Vengono dunque applicate le norme previste dall’articolo 23.9 del Codice Internazionale per la Nomenclatura Zoologica al fine di ottenere un’inversione di priorità a vantaggio del nome più recente (considerato <strong>nomen protectum</strong>) rispetto al nome più antico (considerato <strong>nomen oblitum</strong>). Si stabilisce inoltre la priorità di <em>Scarabaeus serotinus</em> Panzer, 1799 su <em>Aphodius</em> <em>serotinus</em> Creutzer, 1799.
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4

MALDANER, MARIA E., MARIO CUPELLO, DANIELA C. FERREIRA, and FERNANDO Z. VAZ-DE-MELLO. "Type specimens and names assigned to Coprophanaeus (Megaphanaeus) d’Olsoufieff, 1924, the largest New World dung beetles (Coleoptera: Scarabaeidae: Phanaeini)." Zootaxa 4272, no. 1 (2017): 83. http://dx.doi.org/10.11646/zootaxa.4272.1.4.

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Currently 13 species-group names assigned to the subgenus Megaphanaeus of Coprophanaeus are deemed available (or 'potentially available'), four of which denote valid species. In the present work we review the nomenclatural history of those names and conclude that two of them, Scarabaeus satelles Lichtenstein, 1796 and Copris ajax Sturm, 1826, are unavailable. For the other 11 names, we found type specimens of five, Scarabaeus bellicosus Olivier, 1789, Copris ensifer Germar, 1821, Phanaeus septentrionalis Pêssoa, 1934, P. vicinus Martínez, 1944, and P. vicinus var. argentinus Martínez, 1944. Two lectotypes are designated, one for Copris ensifer and other for Phanaeus septentrionalis. We were unable to locate the type series of the four names described by Castelnau (1840), i.e. Phanaeus ducalis, P. sylvanus, P. heros and P. miles, and of P. bonariensis Gory, 1844 and Scarabaeus lancifer Linné, 1767. For the latter, based on some iconotypes, we found that its type series was composite, including specimens of two different Megaphanaeus species, and, in addition, it was mixed with Copris ensifer type series. In order to fix the name to a sole species, a neotype is designated for Scarabaeus lancifer. The history of the names are presented in detail, and the application of these names to species is briefly re-discussed.
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5

Dan, Anca. "Deux scarabées d’Orgamè/Argamum." Dialogues d'histoire ancienne 37, no. 1 (2011): 9–40. http://dx.doi.org/10.3406/dha.2011.3254.

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6

Dan, Anca. "Deux scarabées d'Orgamè/Argamum." Dialogues d'histoire ancienne 37/1, no. 1 (2011): 9. http://dx.doi.org/10.3917/dha.371.0009.

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7

GÉNIER, FRANÇOIS, and PHILIPPE MORETTO. "Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae)." Zootaxa 4248, no. 1 (2017): 1. http://dx.doi.org/10.11646/zootaxa.4248.1.1.

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The taxonomy and systematics of the genus Digitonthophagus Balthasar (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini) is revised. A detailed study of the male genitalia combined with external morphology suggests that the variability, previously recognized, for D. gazella is hiding a species complex within the Afrotropical region and the Arabian Peninsula. The current study recognizes 16 species; 13 from the Afrotropical region and Arabian Peninsula and three from the eastern portion of the Saharo-Arabian region and the continental Indomalayan region. Species are organized into six species groups based on the results of the morphology-based phylogenetic analysis. The following 12 species are described as new: D. aksumensis Génier new species; D. biflagellatus Génier new species; D. dilatatus Génier new species; D. eucatta Génier new species; D. falciger Génier new species; D. fimator Génier new species; D. namaquensis Génier new species; D. petilus Génier new species; D. sahelicus Moretto new species; D. uks Génier new species; D. ulcerosus Génier new species; and D. viridicollis Génier new species. In order to stabilize nomenclature, lectotypes are designated for Scarabaeus bonasus Fabricius, 1775; Scarabaeus catta Fabricius, 1787, and Onthophagus gazella lusinganus d’Orbigny. A neotype is designated for Scarabaeus dorcas Olivier, 1789 whose status and synonymy need to be altered in order to clarify the status of Scarabaeus gazella auctorum, the widely introduced species with economic importance. A naming scheme is presented for the sclerites of the internal sac. External and male genitalia are illustrated and distribution maps are provided for each species.
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8

García Martínez, María Antonia. "Escarabeos inéditos de la provincia de Sevilla." SPAL. Revista de Prehistoria y Arqueología de la Universidad de Sevilla, no. 4 (1995): 257–63. http://dx.doi.org/10.12795/spal.1995.i4.11.

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9

Scholtz, Clarke, Catherine Sole, Shaun Forgie, and Helena Wirta. "Origin of Madagascan Scarabaeini dung beetles (Coleoptera: Scarabaeidae): dispersal from Africa." Insect Systematics & Evolution 42, no. 1 (2011): 29–40. http://dx.doi.org/10.1163/187631211x552800.

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AbstractMadagascar, the world's fourth largest island, has a long history of isolation (160 million years) and a wide range of climates and ecosystems which have in turn resulted in high levels of endemism across different taxonomic levels. Although Madagascar has a rich dung beetle fauna that belongs to various tribes only three species of the Scarabaeini are found there, namely Scarabaeus viettei, S. radama and S. sevoistra. These three species are superficially quite distinctive and have, consequently, had a variable taxonomic history since the first one was described in 1896. The morphological differences between these species resulted in them being placed in different genera at different times. However, currently, based on cladistic analyses, they are all classified in the genus Scarabaeus. In this study, two of the species, S. viettei and S. radama, were included in a phylogenetic analysis based on two mitochondrial gene regions — cytochrome oxidase I (COI) and 16S rRNA — and a 247 morphological and behavioural dataset of 23 members of the Scarabaeinae. A Bayesian phylogram supports the monophyly of the genus Scarabaeus, with the two species from Madagascar appearing sister to three species of Scarabaeus from south-west Africa. Estimated times of divergence based on published mutation rates of 0.012 and 0.0075 for COI indicate that a shared African/Madagascan origin occurred around 15.18 or 24.15 million years ago, respectively. is study is another example in support of Madagascan fauna having an African origin with colonisation having occurred via dispersal as opposed to ancient vicariant events.
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10

Branco, Tristão, and Stefano Ziani. "THE GENUS THORECTES MULSANT, 1842: A RECTIFICATION REGARDING ITS TYPE SPECIES AND SOME CONSIDERATIONS ABOUT ITS TAXONOMY (COLEOPTERA, GEOTRUPIDAE)." Fragmenta Entomologica 39, no. 2 (2007): 255. http://dx.doi.org/10.4081/fe.2007.122.

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A rectification is made to the authors’ previous statement about the reason why <em>Scarabaeus laevigatus</em> sensu Mulsant, 1842 (= <em>Scarabaeus intermedius</em> O.G. Costa, 1839) is the validly designated type species of <em>Thorectes</em> Mulsant, 1842. The history of the taxonomy of <em>Thorectes</em> Mulsant, 1842 is briefly reviewed, and reasons are adduced why the authors believe that the split of the genus, beyond eventually the recognition of two or perhaps three subgenera, is neither necessary nor desirable.
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11

ZIANI, STEFANO, and TRISTÃO BRANCO. "The authorship and date of the genus-group name Pachylomera (Coleoptera: Scarabaeidae)." Zootaxa 2803, no. 1 (2011): 63. http://dx.doi.org/10.11646/zootaxa.2803.1.7.

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In 1828, the English entomologist and clergyman William Kirby described Scarabaeus femoralis using material from the collection of another English entomologist and clergyman, Frederick William Hope. The holotype of Scarabaeus femoralis Kirby, 1828 (Fig. 1) is in the Hope Entomological Collections, Oxford University Museum of Natural History (hereafter referred to as OUMNH). The genus Pachylomera was subsequently erected for this species widespread in central and southern Africa. The purpose of this note is to demonstrate, under the rules of the International Code on Zoological Nomenclature (1999), that Griffith & Pidgeon (1831) should be credited with the authorship of the genus-group name Pachylomera.
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12

Moret, Jean-Marc. "Un groupe de scarabées italiques." Journal des savants 1, no. 1 (1995): 31–64. http://dx.doi.org/10.3406/jds.1995.1583.

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13

Vanlathem, Marie-Paule. "Scarabées de cœur in situ." Chronique d'Egypte 76, no. 151-152 (2001): 48–56. http://dx.doi.org/10.1484/j.cde.2.309162.

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14

Mortazavi, Abdolazim, and Hamidreza Hajiqanbar. "A New Podapolipid Species (Acari) On Scarabaeus (Scarabaeus) Acuticollis (Insecta: Coleoptera: Scarabaeidae) from Iran." Journal of Parasitology 98, no. 4 (2012): 746–53. http://dx.doi.org/10.1645/ge-3034.1.

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15

DANIEL, GIMO M. "The nomenclatural status of the generic nomina Epirinus Dejean, 1833 and Epirhinus Agassiz, 1846 (Coleoptera, Scarabaeidae, Scarabaeinae)." Bionomina 15, no. 1 (2019): 63–65. http://dx.doi.org/10.11646/bionomina.15.1.6.

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The genus Epirinus Dejean, 1833 was established with seven originally included species, none of which was designated as type. Reiche’s (1841) and Janssens’ (1938) designations of a type species for this genus were invalid, but Ferreira’s (1972) subsequent designation of Scarabaeus flagellatus Fabricius, 1775 is valid by virtue of Article 69.2.2, as this author had placed the nomen Scarabaeus granulatus Olivier, 1789, one of the originally included species of Epirinus, in its synonymy: the latter nominal species is therefore the type species of this nominal genus. The unjustified emendation Epirhinus Agassiz, 1846 of Epirinus Dejean, 1833 is nomenclaturally available but invalid, being its junior objective synonym. As for the spelling Epirrhinus introduced by Bedel (1903), it is an incorrect subsequent spelling, devoid of nomenclatural availability.
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16

Thomas, Donald B., Ainsley Seago, and David C. Robacker. "Reflections On Golden Scarabs." American Entomologist 53, no. 4 (2007): 224–30. http://dx.doi.org/10.1093/ae/53.4.224.

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17

Khaustov, Alexander A., and Andrey V. Frolov. "A NEW SPECIES OF PAVANIA (ACARI: HETEROSTIGMATA: DOLICHOCYBIDAE) ASSOCIATED WITH SCARABAEUS TYPHON (COLEOPTERA: SCARABAEIDAE) FROM RUSSIA." Acarina 28, no. 2 (2020): 169–76. http://dx.doi.org/10.21684/0132-8077-2020-28-2-169-176.

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A new species Pavania foliata sp. n. (Acari: Heterostigmata: Dolichocybidae), phoretic on dung beetle Scarabaeus typhon Fischer-Waldheim (Coleoptera: Scarabaeidae) from southwest Russia, is described. The updated key to the species of the genus Pavania is provided.
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18

Regier, Willis Goth. "Erasmus and Aesop." Erasmus Studies 39, no. 1 (2019): 51–74. http://dx.doi.org/10.1163/18749275-03901004.

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Abstract Erasmus was a fluent Aesopian. In books and letters he cited Aesop’s fables to explain, admonish, and insult. The Adagiorum Chiliades alludes to more than seventy different fables, including two adages about Aesop: “Ne Aesopum quidem trivisti” (2.6.27); and “Aesopicus sanguis” (2.6.63). The great adage “Scarabeus aquilam quaerit” (3.7.1) begins with Aesop’s fable. Erasmus’ own contributions to collections of fables were printed in Antwerp, Basel, Louvain, Strasbourg, Paris, and Venice. This paper examines Erasmus’ use of Aesop, identifies the fables Erasmus favored, and places his versions of fables in the history of Aesop transmission.
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19

Clayton, Peter A. "Some more ‘Fierce Lions’, and a ‘Marriage’ Scarab: The Large Commemorative Scarabs of Amenophis III." Journal of Egyptian Archaeology 82, no. 1 (1996): 208–10. http://dx.doi.org/10.1177/030751339608200125.

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20

Ratcliffe, Brett C. "Prairie Scarabs: The Nebraska Perspective." American Entomologist 36, no. 1 (1990): 28–36. http://dx.doi.org/10.1093/ae/36.1.28.

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21

Adam, Klaus-Peter. "Gillûlîm: Rollers, Scarabs in Ezekiel." Catholic Biblical Quarterly 83, no. 2 (2021): 220–40. http://dx.doi.org/10.1353/cbq.2021.0046.

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22

Hewavithana, Dishane K., Mayuri R. Wijesinghe, Chandima D. Dangalle, and H. A. S. Gayan Dharmarathne. "Habitat and dung preferences of scarab beetles of the subfamily Scarabaeinae: a case study in a tropical monsoon forest in Sri Lanka." International Journal of Tropical Insect Science 36, no. 02 (2016): 97–105. http://dx.doi.org/10.1017/s1742758416000023.

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AbstractWe studied the food and habitat preferences of beetles of the subfamily Scarabaeinae in a protected tropical monsoon forest, the Wasgomuwa National Park in Sri Lanka. While gaining an insight into the capacity of different resources to support dung beetles, we tested the hypothesis that, in a landscape with a mosaic of habitat types and a diversity of large mammals, specialization facilitates coexistence of the scarabs. Pitfall traps were laid in five distinct habitat types (forest, riverine, sandy banks, scrub and grassland), and baited with five dung types (leopard, cervid, elephant, bear and buffalo). Four hundred and fifty seven beetles of 22 species in seven genera were recorded. Significant differences were seen in the species richness and abundance of dung beetles between different habitats and dung types, indicating habitat and food preferences. The highest richness of scarabs was in the riverine habitat and in leopard dung. Niche breath values of the individual species indicated differences in the degrees of specialization, while niche overlap values indicated a greater sharing of dung beetle species between similar habitats and dung types than between dissimilar ones. These findings provide evidence that selectivity of habitat, and dung, may be important in promoting coexistence among the scarabs in a tropical environment.
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23

Costa, Mafalda, Ana Margarida Arruda, Rui Barbosa, Pedro Barrulas, Peter Vandenabeele, and José Mirão. "A Micro-Analytical Study of the Scarabs of the Necropolis of Vinha das Caliças (Portugal)." Microscopy and Microanalysis 25, no. 1 (2019): 214–20. http://dx.doi.org/10.1017/s143192761801560x.

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AbstractFive scarabs and one scaraboid found in Vinha das Caliças 4 (Beja, Portugal) were analyzed using a micro-analytical methodology in order to determine their mineralogical and chemical composition. Microstructural characterization and chemical analysis revealed that all were composed of a white body of crushed feldspathic sand covered by a lead-rich, alkaline-depleted silicate blue-green glaze showing evident signs of glass deterioration. Variable pressure scanning electron microscopy with X-ray energy dispersive spectrometry, handheld X-ray fluorescence spectroscopy, and micro X-ray diffraction results show that blue-green color of the glaze was produced by using copper ions (Cu2+) in conjunction with the lead antimonate bindheimite, a yellow-colored opacifier. The introduction of small amounts of tin in the structure of bindheimite enabled the production of a ternary Pb–Sb–Sn oxide. Tin, which was most likely added with the copper source (bronze scrapings), is known to facilitate the crystallization of bindheimite. The results are consistent with the five scarabs and one scaraboid being manufactured in Egypt. This study, the first archeometric study of scarabs found in the Iberian peninsula, has greatly contributed to the understanding of the influence of the Eastern and Central Mediterranean world in the Southwestern Iberia during the first millennium B.C.
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24

Ritner, Robert K. "The Site of the Wild Bull-Hunt of Amenophis III." Journal of Egyptian Archaeology 72, no. 1 (1986): 193–94. http://dx.doi.org/10.1177/030751338607200120.

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The toponym Štp, found on the wild bull-hunt scarabs, is identified with the well-attested name of the metropolis of the Wadi el-Natrun, Štp·t. Implications for the court residence are also discussed.
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25

TALLET, P. "Les sceaux-scarabées de deux fonctionnaires des mines." Revue d'Égyptologie 57 (December 31, 2006): 264–67. http://dx.doi.org/10.2143/re.57.0.2019414.

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26

Lorand, David. "Quatre scarabées de cœur inscrits à tête humaine." Chronique d'Egypte 83, no. 165-166 (2008): 20–40. http://dx.doi.org/10.1484/j.cde.2.309336.

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27

Jackson, Trevor A., and Michael G. Klein. "Scarabs as Pests: A Continuing Problem." Coleopterists Bulletin 60, sp5 (2006): 102–19. http://dx.doi.org/10.1649/0010-065x(2006)60[102:sapacp]2.0.co;2.

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28

Martin, Geoffrey T., Raphael Giveon, David Warburton, and Christoph Uehlinger. "Scarabs from Recent Excavations in Israel." Journal of Egyptian Archaeology 77 (1991): 206. http://dx.doi.org/10.2307/3821979.

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29

Walsh, Cabrera G., H. A. Cordo, J. A. Briano, D. E. Gandolfo, and G. A. Logarzo. "Laboratory Culture of Beneficial Dung Scarabs." Journal of Economic Entomology 90, no. 1 (1997): 124–29. http://dx.doi.org/10.1093/jee/90.1.124.

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30

KRELL, FRANK-THORSTEN. "On nomenclature and synonymy of Trichius rosaceus, T. gallicus, and T. zonatus (Coleoptera: Scarabaeidae: Cetoniinae: Trichiini)." Zootaxa 3278, no. 1 (2012): 61. http://dx.doi.org/10.11646/zootaxa.3278.1.3.

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The name Scarabaeus rosaceus Voet, 1766–1778, currently in use as one of two names for a west Palaearctic Trichius species, is unavailable because Voet’s work is not consistently binominal. The valid name for the species is Trichius gallicus Dejean, 1821 with the Sardinian and North African populations forming the subspecies T. g. zonatus Germar, 1831. The lectotype of Trichius zonatus Germar is designated.
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31

Krell, Frank-Thorsten, Zdzislawa Stebnicka, and Erik Holm. "Scarabaeus Rufus Moll, 1782 (Currently Aphodius Rufus), Scarabaeus Rufus Fabricius, 1792 (Currently Aegialia Rufa) And Scarabaeus Foetidus Herbst, 1783 (Currently Aphodius Foetidus) (Insecta, Coleoptera): Proposed Conservation Of Usage Of The Specific Names." Bulletin of zoological nomenclature. 51 (1994): 121–27. http://dx.doi.org/10.5962/bhl.part.7175.

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32

Smith, Andrew B. T., and François Génier. "Revision of the genus Holocephalus (Coleoptera: Scarabaeidae: Scarabaeinae: Coprini)." Canadian Entomologist 133, no. 6 (2001): 777–92. http://dx.doi.org/10.4039/ent133777-6.

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AbstractThe genus Holocephalus Hope is revised and now contains five species: H. cristatus (Gillet), H. eridanus (Olivier), H. julienisp.nov., H. sculptus (Gillet), and H. simonisp.nov. All taxa are described and illustrated. A key to species of Holocephalus is included. Lectotypes are designated for H. eridanus and H. sculptus. Scarabaeus eridanus Olivier is designated as the type species for the generic name Atrichius Gillet (a junior synonym of Holocephalus).
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33

Davaras, C. "A Minoan Beetle-Rhyton From Prinias Siteias." Annual of the British School at Athens 83 (November 1988): 45–54. http://dx.doi.org/10.1017/s0068245400020633.

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A terracotta Minoan figurine of a horned beetle from the Peak Sanctuary on Mt Prinias is discussed. It is shown to be a rhyton, and presumably made for cult purposes. The reasons for offerings of such beetles is investigated, and the connection with Egyptian scarabs is raised.
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34

David, Arlette. "Uninscribed Amethyst Scarabs from the Southern Levant." Bulletin of the American Schools of Oriental Research 381 (May 2019): 57–81. http://dx.doi.org/10.1086/703076.

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35

Teeter, Emily, and Andree Feghali Gorton. "Egyptian and Egyptianizing Scarabs: A Typology of Steatite, Faience and Paste Scarabs from Punic and Other Mediterranean Sites." American Journal of Archaeology 103, no. 1 (1999): 140. http://dx.doi.org/10.2307/506596.

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36

NIOGRET, JEROME, ANTOINE NICOT, ERIC DE STORDEUR, and MICHEL BERTRAND. "Combination of morphological characters and ITS-sequence to characterize a new species of Macrocheles (Acari: Macrochelidae)." Zootaxa 1386, no. 1 (2007): 19–29. http://dx.doi.org/10.11646/zootaxa.1386.1.2.

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Macrocheles seraphim sp. nov. is described from French and Moroccan populations. This species was found phoretic on three species of dung beetles: Scarabaeus laticollis, Geotrupes mutator and G. spiniger. It was identified on the basis of morphological characters as belonging to the M. glaber group, but also has similarities with M. muscaedomesticae. Its systematic position as investigated by genetic analyses using the ITS-1 marker showed that this species was closely related to M. muscaedomesticae. Its relationship with glaber group species is discussed.
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Ruiz, J. L., S. Yubero, and L. García-Cardenete. "Sobre la distribución geográfica de Scarabaeus (Scarabaeus) aegyptiacus Stolfa, 1938 (Coleoptera: Scarabaeidae): primer registro para el Sáhara Atlántico (suroeste de Marruecos)." Graellsia 73, no. 1 (2017): 056. http://dx.doi.org/10.3989/graellsia.2017.v73.176.

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Scarabaeus aegyptiacus es una especie ampliamente distribuida desde la Península Arábiga hasta Marruecos, ocupando ambientes áridos y desérticos. A pesar de su amplio rango de distribución, el número de localidades precisas conocidas es muy escaso (diez localidades) y sus poblaciones se encuentran aparentemente fragmentadas y muy distanciadas entre sí. Durante muestreos recientes en el suroeste de Marruecos, hemos hallado una población de S. aegyptiacus en una estepa pedregosa (hamada) en las cercanías de Msied (provincia de Tan-Tan, región de Guelmín-Smara), que constituye el primer registro para el Sáhara Atlántico y amplía unos 1000 km hacia el suroeste su área de distribución conocida. Los individuos estudiados fueron hallados en el interior de un aljibe (“matfiya” en lengua local), infraestructura que actúa como trampa mortal para numerosos animales, especialmente anfibios, reptiles y artrópodos epígeos, cuyo impacto negativo sobre las comunidades de invertebrados no ha sido evaluado. Se sintetiza la información corológica disponible y se aportan datos sobre el hábitat de la especie.
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Backes, Burkhard. "Summe seiner Teile?" Zeitschrift für Ägyptische Sprache und Altertumskunde 146, no. 2 (2019): 129–37. http://dx.doi.org/10.1515/zaes-2019-0012.

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Summary New interpretation of Berlin, Ägyptisches Museum 11405. The object shows traces of voluntary mutilation not dissimilar to the much older ‘reserve heads’. Some elements have been left unfinished. In the light of iconographic parallels in some reliefs, the combination of two embracing scarabs with a human head is identified as an image of Ptah as creator-god.
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Ghahari, Hassan, Marek Bunalski, Mehrdad Tabari, and Hadi Ostovan. "Scarab beetles (Coleoptera: Scarabaeidae) collected in Iranian rice fields and surrounding grasslands." Polish Journal of Entomology / Polskie Pismo Entomologiczne 80, no. 3 (2011): 465–74. http://dx.doi.org/10.2478/v10200-011-0034-z.

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Scarab beetles (Coleoptera: Scarabaeidae) collected in Iranian rice fields and surrounding grasslands Scarab beetles (Coleoptera: Scarabaeidae) comprise a group of species that are present in almost all agro-ecosystems. The fauna of these insects was studied in Iranian rice fields and surrounding grasslands from 2000 to 2005. A total of 18 species from 14 genera (Aphodius, Pleurophorus, Tropinota, Protaetia, Valgus, Amphimallon, Aplidia, Holochelus, Polyphylla, Chaetopteroplia, Asiopertha, Onthophagus, Scarabaeus, Paroniticellus) and 6 subfamilies (Aphodiinae, Cetoniinae, Melolonthinae, Rutelinae, Scarabaeinae, Coprinae) were collected. Of these, 9 species are new records for Iran.
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DORION-PEYRONNET, Caroline. "Sceaux-scarabées inédits au nom de deux vizirs d'époque tardive." Revue d'Égyptologie 60 (December 31, 2009): 201–4. http://dx.doi.org/10.2143/re.60.0.2049282.

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41

Peck, William H. "Essai de classification et datation des scarabées Menkhéperrê. Bertrand Jaeger." Journal of Near Eastern Studies 46, no. 3 (1987): 235–36. http://dx.doi.org/10.1086/373248.

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42

Martin, Geoffrey T. "Book Review: Scarabs from recent excavations in Israel." Journal of Egyptian Archaeology 77, no. 1 (1991): 206–7. http://dx.doi.org/10.1177/030751339107700132.

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43

Porter, Robert Martin. "Scarabs of Sheshi at Tell El-ajjul,ContraKempinski." Palestine Exploration Quarterly 148, no. 2 (2016): 133–45. http://dx.doi.org/10.1080/00310328.2016.1186344.

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44

Scholtz, Clarke, Armanda Bastos, and Catherine Sole. "Do individual and combined data analyses of molecules and morphology reveal the generic status of 'Pachysoma' MacLeay (Coleoptera: Scarabaeidae)?" Insect Systematics & Evolution 38, no. 3 (2007): 311–30. http://dx.doi.org/10.1163/187631207788754411.

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AbstractScarabaeus (Pachysoma) comprises 13 flightless dung beetle species endemic to the west coast of southern Africa that are biologically unique and have a variable taxonomic history at the generic level. Initially considered a valid genus Pachysoma was subsequently synonymised with the genus Scarabaeus and later assigned sub-generic status within this genus. In an attempt to resolve past taxonomic inconsistencies, morphological and molecular data partitions, and combinations thereof, were analysed and compared across different phylogenetic inference methods. Trees obtained from the individual datasets which comprised 64 morphological characters and 1197 nucleotide sites of the mitochondrial Cytochrome Oxidase I (COI) gene, were less well-resolved than those obtained from analyses in which the data partitions were combined whilst Bayesian inference generally out-performed parsimony. However, both the individual and combined data partitions, irrespective of the method of analysis, confirmed the monophyly of the Pachysoma lineage. The relative contribution of each data partition to individual nodes as assessed by positive partitioned Bremer support values was 6:5:4 for the molecular:combined:morphological data partitions. However, in terms of contribution to the resolution of terminal versus internal nodes, the morphological dataset made proportionally the greatest contribution and was crucial for recovering a sister relationship of Pachysoma to Scarabaeus, i.e. a generic level, rather than a sub-generic one. This together with the monophyly of the Pachysoma lineage across all methods of analysis and data partitions as well as the unique morphology and biology of its constituent species warrants reinstatement of Pachysoma to its former full generic status.
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Verdu, Jose R., Alfonso Diaz, and Eduardo Galante. "Thermoregulatory strategies in two closely related sympatric Scarabaeus species (Coleoptera: Scarabaeinae)." Physiological Entomology 29, no. 1 (2004): 32–38. http://dx.doi.org/10.1111/j.0307-6962.2004.0359.x.

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Mantelli, Sincero. "Quia scarabaeus uel cantharus uermis est stercoris. Una glossa erasmiana nel Commentario ad Abacuc di Gerolamo." Augustinianum 50, no. 2 (2010): 443–51. http://dx.doi.org/10.5840/agstm201050216.

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In the main editions of the Commentary by Jerome on the prophet Habakkuk one can read a gloss (« quia scarabaeus uel cantharus uermis est stercoris ») which cannot be found in the manuscript that is usually consulted. The codices show different readings which in most cases quote a corrupt text. Considering that the above-mentioned note does not appear in the pre-Erasmian editions one can draw the conclusion that Erasmus himself corrected the text, adding the meaningful note. To confirm what has been said one can add other corrections, perhaps less significant but which nevertheless can be equally attributed to the hand of the Rotterdam Humanist.
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Hajiqanbar, Hamidreza, Alexandr Khaustov, and Abdolazim Mortazavi. "New species of the genera Dolichocybe and Pavania (Acari: Dolichocybidae), phoretic on scarab beetles (Coleoptera: Scarabaeidae) from Iran." Systematic and Applied Acarology 24, no. 8 (2019): 1363–82. http://dx.doi.org/10.11158/saa.24.8.3.

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In a survey of prostigmatic mites associated with insects in southern parts of Iran, a new species of the genus Dolichocybe and four new species of the genus Pavania as associates of the scarab and trogid beetles (Coleoptera: Scarabaeidae, Trogidae) were found: Dolichocybe orzueeyehiensis sp. nov. from Adoretus sp., Pavania kermaniensis sp. nov. from Aphodius cf. lividus (Olivier), P. magowskii sp. nov. from Trox sp., P. megasolenidia sp. nov. from Aphodius sp., and P. scarabaeophilus sp. nov. from Scarabaeus sp. The scarab genus Adoretus is a new host record for mites of the genus Dolichocybe, and the genera Aphodius and Trox are new host records for mites of the genus Pavania.
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KOHLMANN, BERT, and ÁNGEL SOLÍS. "New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama." Zootaxa 3193, no. 1 (2012): 28. http://dx.doi.org/10.11646/zootaxa.3193.1.2.

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The following new species of geotrupids and scarabs from Costa Rica and Panama are described: Athyreus gulesserianinew species, Ateuchus alutacius new species (the first recorded brachypterous Ateuchus species), Coprophanaeus geph-yra new species, Deltochilum acanthus new species, and Onthophagus turgidus new species. The following two speciesof Coprophanaeus are revalidated: C. kohlmanni Arnaud and C. uhleri Malý & Pokorný. Illustrations of the dorsal habitus of the new species are provided, as well as distribution maps for all species.
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Shan, Yueming, Changlong Shu, Neil Crickmore, et al. "Cultivable Gut Bacteria of Scarabs (Coleoptera: Scarabaeidae) InhibitBacillus thuringiensisMultiplication." Environmental Entomology 43, no. 3 (2014): 612–16. http://dx.doi.org/10.1603/en14028.

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50

Brandl, Baruch. "Two Scarabs and a Trapezoidal Seal from Mount Ebal." Tel Aviv 14, no. 2 (1987): 166–72. http://dx.doi.org/10.1179/tav.1986.1986.2.166.

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