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1

Singh, Vijay, Sunita Grewal, Vivek Goyal, and Vinay Malik. "Taxonomic redescription of subfamily Scymninae (Coleoptera: Coccinellidae) from Haryana, India." Journal of Applied and Natural Science 16, no. 2 (2024): 842–53. http://dx.doi.org/10.31018/jans.v16i2.5461.

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The ladybird beetles of Scymninae, a subfamily of Coccinellidae are efficient biocontrol agents. Many earlier studies document their protective role associated with many crops. This subfamily is represented by 5 tribes,15 genera and 138 species in the Indian subcontinent. However there is no scientific record of taxonomic description of the subfamily Scymninae within the Coccinellidae family of beetles (Coccinellidae: Coleoptera) in Haryana, India. In the present study, seven species from three genera and two tribes of sub-family Scymninae i.e. Nephus (Bipunctatus) bipunctatus (Kugelann,1794), Nephus regularis (Sicard), Scymnus (Pullus) coccivora Ramakrishna Ayyar, 1925, Scymnus (Pullus) latemaculatus Motschulsky, Scymnus (Scymnus) nubilus Mulsant, Scymnus (Pullus) posticalis Sicard and Stethorus aptus Kapur were identified. It included the generation of keys to the tribes, genera, subgenera and species found during the study. It provided detailed taxonomy of the identified species based on various morphological characteristics such as coxal lines, antennae, mandibles, and male and female genitalia. This study marks the first taxonomic exploration of species within the Scymninae subfamily in Haryana, India. It will significantly contribute to understanding the biodiversity of beetles in the region and will lay the groundwork for further research and conservation initiatives.
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2

Бўриева, Хуршида Парда қизи. "ҚАШҚАДАРЁ ХУДУДИДА ТАРҚАЛГАН SCYMNUS (KUGELANN, 1794) АВЛОДИГА КИРАДИГАН ТУРЛАР ТАРКИБИ." THEORETICAL ASPECTS IN THE FORMATION OF PEDAGOGICAL SCIENCES 1, № 5 (2022): 259–62. https://doi.org/10.5281/zenodo.7229252.

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Ўрта Осиёда 180 тури, Ўзбекистонда эса 2 та кенжа оиласи, 25 авлодга мансуб 106 та тур ва кенжа турлари аниқланган . N.J.Vandenberg  маълумотларига кўра, дунё миқёсида кокцинеллидларнинг (Coccinellidae) 6 та кенжа оиласи (Coccidulinae, Coccinellinae, Scymninae, Chilocorinae, Sticholotidinae, Epilachninae) мавжудлиги қайд қилинган.
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3

Ashfaque, Muhammad, Farman Ullah, Muhammad Ather Rafi, and Falak Naz. "Taxonomic study of subfamily Scymninae (Coleoptera: Coccinellidae) with one new record from Gilgit-Baltistan, Pakistan." Turkish Journal of Zoology 39, no. 6 (2015): 1034–40. https://doi.org/10.3906/zoo-1401-17.

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Ashfaque, Muhammad, Ullah, Farman, Rafi, Muhammad Ather, Naz, Falak (2015): Taxonomic study of subfamily Scymninae (Coleoptera: Coccinellidae) with one new record from Gilgit-Baltistan, Pakistan. Turkish Journal of Zoology 39 (6): 1034-1040, DOI: 10.3906/zoo-1401-17, URL: http://dx.doi.org/10.3906/zoo-1401-17
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4

Coutanceau, Jean-Pierre. "Scymninae nouveaux pour la faune de France (Col., Coccinellidae)." Bulletin de la Société entomologique de France 116, no. 2 (2011): 180. http://dx.doi.org/10.3406/bsef.2011.2663.

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5

Ceryngier, Piotr. "Stethorus pusillus (Coleoptera: Coccinellidae) as a host of the ectoparasitic fungus Hesperomyces coccinelloides (Ascomycota: Laboulbeniales: Laboulbeniaceae) in Poland." Polish Journal of Entomology / Polskie Pismo Entomologiczne 82, no. 1 (2013): 13–18. http://dx.doi.org/10.2478/v10200-012-0018-7.

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Abstract The laboulbenialean fungus Hesperomyces coccinelloides (THAXTER) THAXTER is known to be an obligate ectoparasite of beetles belonging to the coccinellid subfamily Scymninae. It has been reported from several regions in the world, mostly tropical and subtropical localities. This paper reports the first records of H. coccinelloides from Poland. The parasitic thalli were found growing on the elytra of Stethorus pusillus (HERBST) specimens collected in two adjacent localities in the vicinity of Warsaw.
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6

Burieva, Khurshida, Gulnara Mirzaeva, Nodira Arabova, and Makhmuda Mansurkhodjaeva. "Bioecological features of coccinellide in the southern Kashkadarya province, Uzbekistan." E3S Web of Conferences 284 (2021): 03014. http://dx.doi.org/10.1051/e3sconf/202128403014.

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This research was conducted in 2019-2020 in the northern and southern administrative districts of the southern Kashkadarya province of Uzbekistan. This study on the distribution of coccinellides in agrobiocenoses has been conducted at different experimental periods. In the agrobiocenoses of Kashkadarya province, 5 subspecies of coccinellides (Scymninae, Chilocorinae, Coccidulinae, Epilachninae, and Coccinellinae), 8 tribes, and 31 species belonging to 22 genera were found. Experiments have shown that species belonging to the genera Chilocorus (Leach, 1875) are relatively rare species and are not considered specific species for cotton agrocenosis. Coccinella septempunctata was observed as the most common species. The eggs of Coccinella septempunctata were studied separately in this research. Wintering conditions in mountainous areas were also considered and particular attention was paid to the beneficial entomophagous species of coccinellides.
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7

ASHFAQUE, Muhammad, Farman ULLAH, Muhammad Ather RAFI, and Falak NAZ. "Taxonomic study of subfamily Scymninae (Coleoptera: Coccinellidae)with one new record from Gilgit-Baltistan, Pakistan." TURKISH JOURNAL OF ZOOLOGY 39 (2015): 1034–40. http://dx.doi.org/10.3906/zoo-1401-17.

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8

Gafurova, Saodat T., Bakhtiyor R. Kholmatov, and Gulnara S. Mirzaeva. "An annotated checklist of ladybeetle species (Coleoptera, Coccinellidae) of Ferghana Valley (Uzbekistan)." Acta Biologica Sibirica 11 (February 26, 2025): 171–95. https://doi.org/10.5281/zenodo.14915341.

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The article presents the results of a study on the fauna of lady beetles (Coleoptera, Coccinellidae) based on our observations in the Fergana Valley during the spring, summer, and autumn seasons as well as the winter season from 2022 to 2024. Thirty-five species of lady beetles, belonging to 5 subfamilies (Scymninae, Chilocorinae, Ortaliinae, Coccinellinae, Hyperaspidinae), 9 tribes, and 19 genera, were found along the valley. Noteworthy species identified in the Fergana Valley during the research include <em>Novius yemenensis</em> (Raimundo &amp; F&uuml;rsch, 2006), <em>Platynaspidius saundersi</em> (Crotch, 1874), <em>Brachiacontha ursine</em> (Chevrolat in Dejean, 1837), <em>Scymnus marinus</em> (Mulsant, 1850). The species <em>Calvia muiri</em> (Timberlake, 1943), <em>Clitostethus arcuatus</em> (Rossi, 1794) were recorded for the first time in the fauna of Uzbekistan and 23 species were recorded for the first time in the Fergana Valley.&nbsp;
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9

Cargnus, Elena, Federico Tacoli, Francesco Boscutti, Pietro Zandigiacomo, and Francesco Pavan. "Side Effects of Kaolin and Bunch-Zone Leaf Removal on Generalist Predators in Vineyards." Insects 14, no. 2 (2023): 126. http://dx.doi.org/10.3390/insects14020126.

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In vineyards, kaolin application and bunch-zone leaf removal (LR) were effective in the control of leafhoppers and Lobesia botrana, but their side effects on generalist predators are still poorly understood. In north-eastern Italian vineyards, the impact of kaolin and LR on species and functional diversity of spiders, as well as the abundance of spiders and generalist predatory insects, was assessed in one vineyard for two consecutive years and in two vineyards for one year. The ecological indices of the spider community were never influenced by kaolin and only in one case were they influenced by LR. At the spider family level, kaolin reduced the abundance of Araneidae, Oxypidae and Salticidae, but only in single cases. In single cases, kaolin reduced the amount of Orius sp. anthocorids and increased that of Scymninae coccinellids, whereas LR increased the amount of Aeolothrips sp. Moderate use of kaolin and the application of LR had negligible and inconsistent impacts on generalist predatory arthropods in vineyards and was therefore compatible with IPM strategies.
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10

BELLA, SALVATORE, COSIMO BAVIERA, and ROBERTO CATANIA. "PHENOLOGY AND POPULATION DYNAMICS OF SOME SCYMNUS KUGELANN SPECIES (COLEOPTERA COCCINELLIDAE) IN CITRUS ORCHARDS WITH DIFFERENT MANAGEMENT IN SICILY (ITALY)." Redia 108 (February 14, 2025): 3–10. https://doi.org/10.19263/redia-108.25.01.

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Surveys on Scymninae (Coleoptera, Coccinellidae) in two citrus orchards in Sicily (Italy) with different management (organic vs integrated management) were carried out for three years (2017-2020). The phenology and population dynamics of species belonging to the genus Scymnus Kugelann are reported. The following eight species were studied: Scymnus apetzi Mulsant, 1846, Scymnus interruptus (Goeze, 1777), Scymnus laetificus Weise, 1879, Scymnus marginalis Rossi, 1794, Scymnus nubilus (Mulsant, 1850), Scymnus rubromaculatus (Goeze, 1777), and Scymnus subvillosus (Goeze, 1777) all aphidophagous, and Scymnus marinus Mulsant, 1850 coccidophagous. Overall, 2,477 specimens of Scymnus were collected over three years in the two sites. Of these, 63.9% were aphidophagous, while 36.1% were coccidophagous. A total of 920 specimens (7 species) were found in the organic orchard, dominated by aphidophagous species (99.1%,), with only 0.8% of the coccidophagous S. marinus. A different situation occurred at the integrated orchard, where 1,557 specimens were collected. Of them, 43% belonged to the aphidophagous species, and 56.9% to S. marinus. Key Words: Coccinellidae, Scymnus phenology, natural enemies, Citrus orchard, Sicily.
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11

Vanegas-Rico, Juan M., Rebeca Peña-Martínez, and Ana L. Muñoz-Viveros. "First record of Melanaphis donacis (Passerini, 1861) (Hemiptera: Aphididae) in México." Entomological Communications 5 (November 6, 2023): ec05031. http://dx.doi.org/10.37486/2675-1305.ec05031.

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The aphid Melanaphis donacis (Passerini, 1861) (Hemiptera: Aphididae) is recorded in Mexico in seven locations in the Valley of Mexico Basin on the Poaceae Arundo donax, one of the most common host plants. Both the aphid and the host plant are species considered exotic to Mexico; especially A. donax is referred to as one of the hundred most important invasive alien species in the world and is also officially cited in Mexico. Observations and collections of both the aphid colonies and the host plant began in October 2022 to February 2023. The work includes the taxonomic determination of both the aphid and the host plant. Also reviewed of websites (www.naturalista.mx) where in two sites it was referred to in a preliminary way, without taxonomic certainty by a specialist. In addition to the worldwide distribution of the aphid species and its host plants. Parthenogenetic wingless and winged forms are presented and described in vivo, as well as in mounts of microscope slides. In all the cases observed in the field, a symbiotic association with ants of the species Linepithema humile Mayr, 1868 (Hymenoptera: Formicidae) is documented. Some natural enemies were obtained, such as Harmonia axididis (Palas, 1773) (Coleoptera: Coccinellidae); Scymninae, Chamaemyiidae and Syrphidae larvae, yet to be determined. It is suggested to increase the search both in other localities and in hosts potentially present in Mexico.
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12

Ali, Muhammad, Khalil Ahmed, Shoukat Ali, et al. "An annotated checklist of Coccinellidae with four new records from Pakistan (Coleoptera, Coccinellidae)." ZooKeys 803 (December 6, 2018): 93–120. https://doi.org/10.3897/zookeys.803.22543.

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Some new ladybird (Coleoptera: Coccinellidae) records collected during the last four years across Sindh are reported. A first preliminary checklist of ladybirds from Sindh is presented, consisting of one subfamily, ten tribes, 21 genera, and 29 species including four new records, namely Bulaea lichatschovii (Hummel), Exochomus pubescens Küster, Scymnus (Pullus) latemaculatus Motschulsky, Scymnus (Pullus) syriacus Marseul, and four varieties of the species Cheilomenes sexmaculatus (Fabricius).
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13

Peng, Feng, Mingjie Tang, Xingmin Wang, and Xiaosheng Chen. "Reniscymnus gen. nov., a new genus of Scymnini (Coleoptera, Coccinellidae) from the Oriental region." ZooKeys 1226 (February 6, 2025): 87–100. https://doi.org/10.3897/zookeys.1226.130352.

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Ladybird beetles (Coleoptera, Coccinellidae) are a species-rich, ecologically diverse and economically important group of insects. A new ladybird genus of Scymnini, Reniscymnusgen. nov., is described from China and Laos, along with two new species, Reniscymnus cordatus Peng &amp; Chen, sp. nov. and Reniscymnus explanatus Peng &amp; Chen, sp. nov. This newly identified genus can be distinguished from the remaining members of the tribe Scymnini by its short antenna consisting of eight antennomeres and an inflated antennal club bearing two extremely long setae; the narrow frons; the slightly tapering apically terminal maxillary palpomere; as well as the recurved and laterally almost complete abdominal postcoxal lines that enclose a distinctly wide area. Detailed morphological descriptions, illustrations, and a key to identify these two species within the new genus are provided. Morphological similarities and relationships of the new genus with other genera of Scymnini are discussed.
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14

Peng, Feng, Mingjie Tang, Xingmin Wang, and Xiaosheng Chen. "Reniscymnus gen. nov., a new genus of Scymnini (Coleoptera, Coccinellidae) from the Oriental region." ZooKeys 1226 (February 6, 2025): 87–100. https://doi.org/10.3897/zookeys.1226.130352.

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Ladybird beetles (Coleoptera, Coccinellidae) are a species-rich, ecologically diverse and economically important group of insects. A new ladybird genus of Scymnini, <i>Reniscymnus</i> gen. nov., is described from China and Laos, along with two new species, <i>Reniscymnus cordatus</i> Peng &amp; Chen, sp. nov. and <i>Reniscymnus explanatus</i> Peng &amp; Chen, sp. nov. This newly identified genus can be distinguished from the remaining members of the tribe Scymnini by its short antenna consisting of eight antennomeres and an inflated antennal club bearing two extremely long setae; the narrow frons; the slightly tapering apically terminal maxillary palpomere; as well as the recurved and laterally almost complete abdominal postcoxal lines that enclose a distinctly wide area. Detailed morphological descriptions, illustrations, and a key to identify these two species within the new genus are provided. Morphological similarities and relationships of the new genus with other genera of Scymnini are discussed.
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15

Poorani, J. "Two new species of Scymnini (Coleoptera: Coccinellidae) from Karnataka, India." Biodiversity Data Journal 3 (June 22, 2015): e5296. https://doi.org/10.3897/BDJ.3.e5296.

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The Scymnini (Coleoptera: Coccinellidae) of the Indian region is rich and highly speciose, with nearly 90 described species and scores of undescribed species (Poorani 2002). There is a dire need to systematically revise the genera and species of this tribe from the Indian region. Due to paucity of representative collections covering the entire region and lack of access to types, it is difficult to identify most of the Scymnini of the Indian region to species. As a result, many economically important species remain poorly characterized, or worse, unnamed. Two economically important and unique species of Scymnini (Coccinellidae) belonging to <i>Horniolus</i> Weise (1900) and Scymnus (Pullus) Mulsant (1846) from the Southern Indian state of Karnataka that have remained unnamed for long are treated in this paper. These species are externally similar to other known species and often misidentified. <i>Horniolus sororius</i> sp. n. and Scymnus (Pullus) rajeshwariae sp. n. (Coleoptera: Coccinellidae) are described here and illustrated with notes on their biology and related species.
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16

Booth, R. G. "A new genus and species of coccinellid (Coleoptera: Coccinellidae) predator on Rastrococcus spp. (Homiptera: Pseudococcidae) from India." Bulletin of Entomological Research 81, no. 2 (1991): 147–50. http://dx.doi.org/10.1017/s000748530005121x.

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AbstractAponephus lentiformis, a new genus and species of Coccinellidae, is described from southern India and placed in the tribe Scymnini. It was found during searches for natural enemies of Rastrococcus spp.
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17

SZAWARYN, KAROL, and JAROSLAV VĚTROVEC. "A new species of Horniolus Weise with remarks on the systematic position of the genus (Coleoptera: Coccinellidae)." Zootaxa 4861, no. 4 (2020): 573–80. http://dx.doi.org/10.11646/zootaxa.4861.4.5.

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Horniolus Weise is an Asian genus of ladybird beetles up to now comprising 15 species with the greatest diversity in China. A new species, H. minutus sp. nov., is described and illustrated. The position of the genus within the modern classification of Coccinellidae is discussed, and its transfer from the tribe Scymnini to Platynaspini is proposed.
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18

VIDYA, C. V., and HASEENA BHASKAR. "Scymnini (Coleoptera:Coccinellidae) associated with major sucking pests of Kerala." Journal of Biological Control 31, no. 4 (2018): 212. http://dx.doi.org/10.18311/jbc/2017/18618.

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19

Ali, Muhammad, Rukhsana Perveen, Arif-Un-Nisa Naqvi, Khalil Ahmed, Ghulam Raza, and Ishtiaq Hussain. "The Tribe Scymnini (Coccinellidae: Coleoptera) From Sindh Province, Pakistan." Journal of Insect Science 15, no. 1 (2015): 146. http://dx.doi.org/10.1093/jisesa/iev105.

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20

Poorani, J. "Two new species of Scymnini (Coleoptera: Coccinellidae) from Karnataka, India." Biodiversity Data Journal 3 (June 22, 2015): e5296. http://dx.doi.org/10.3897/bdj.3.e5296.

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21

Giorgi, José Adriano, and Guillermo González. "A new species of Scymnobius Casey (Coleoptera, Coccinellidae, Scymnini) from Pernambuco, Brazil." Revista Brasileira de Entomologia 58, no. 4 (2014): 316–18. http://dx.doi.org/10.1590/s0085-56262014005000008.

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22

PENG, FENG, J. POORANI, ROGER G. BOOTH, XINGMIN WANG, XIAOSHENG CHEN, and C. ANURADHA. "Slipinskiscymnus gen. nov., a new ladybird genus of Scymnini (Coleoptera, Coccinellidae) and notes on the taxonomic status of Keiscymnus Sasaji." Zootaxa 5325, no. 1 (2023): 97–115. http://dx.doi.org/10.11646/zootaxa.5325.1.6.

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Slipinskiscymnus gen. nov. (Coleoptera: Coccinellidae) is proposed to accommodate two anomalous species of Scymnini, namely, Scymnus pallidicollis Mulsant, 1853 (= Slipinskiscymnus pallidicollis (Mulsant), comb. nov.) and Scymnus saciformis Motschulsky, 1858 (=Slipinskiscymnus saciformis (Motschulsky), comb. nov.) and five new species, Slipinskiscymnus confertus Peng et Chen sp. nov., S. siculiformis Peng et Chen sp. nov., S. spiculatus Peng et Chen sp. nov., S. interstricus Peng et Chen sp. nov. and S. keralensis Poorani sp. nov., described from China and India. A lectotype is designated for Scymnus saciformis Motschulsky, 1858 (lectotype designation). Descriptive accounts of these species with illustrations of adult habitus, male genitalia and other diagnostic characters are given with a key to species. Notes on the status of the genus Keiscymnus Sasaji, 1971, are also provided.
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Soares, António, Isabel Borges, Hugo Calado, and Paulo Borges. "An updated checklist to the biodiversity data of ladybeetles (Coleoptera: Coccinellidae) of the Azores Archipelago (Portugal)." Biodiversity Data Journal 9 (December 16, 2021): e77464. https://doi.org/10.3897/BDJ.9.e77464.

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A recently-published review from 2021 presents a comprehensive checklist of ladybeetles of Portugal, including the Azores and Madeira Archipelagos. Until then, the available information was very scattered and based on a single revision dating back to 1986, a few international catalogues and databases, individual records and studies on communities of agroecosystems. However, no information was available on faunal composition across the Azorean islands and their habitats, using standardised inventories. Here, we present data about the biodiversity of ladybeetles and their distribution and abundance in five Islands of the Azores (Faial, Graciosa, Pico, São Jorge and São Miguel). Surveys included herbaceous and arboreal habitats from native to anthropogenic-managed habitats: ruderal road vegetation, vegetable garden, mixed forest of endemic and non-native host plants, coastal prairies, coastal mixed vegetation, cornfields and urban areas. We aimed to contribute to the ongoing effort to document the terrestrial biodiversity of Portugal, including the Archipelago of the Azores, within the research project AZORESBIOPORTAL–PORBIOTA (ACORES-01-0145-FEDER-000072).In this study, a total of 1,487 specimens of Coccinellidae belonging to 19 species are reported for several habitats. The listed species are from one single sub-familiy (Coccinellinae) and six tribes; Chilocorini (one species), Coccidulini (three species), Coccinellini (six species), Noviini (one species), Scymnini (seven species), Stethorini (one species). The number of species collected per island differed; Faial (10 species), Graciosa (four species), Pico (seven species), São Jorge (seven species) and São Miguel (12 species). For six species, new island records are given. Currently, the number of species known to occur in the Azores are 32, including two doubtful records. The majority of species are Scymnini, being Scymnus (Scymnus) interruptus (Goeze, 1777) and Scymnus (Scymnus) nubilus Mulsant, 1850, the most abundant species (relative abundance 71.1%). This database will be the baseline of a long-term monitoring project allowing assessment of the impact of ongoing global changes in the distribution and abundance of ladybeetles.
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Chen, Xiaosheng, Xiufeng Xie, Shunxiang Ren, and Xingmin Wang. "A taxonomic review of the genus Horniolus Weise from China, with description of a new species (Coleoptera, Coccinellidae)." ZooKeys 623 (October 11, 2016): 105–23. https://doi.org/10.3897/zookeys.623.10191.

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Five species of the genus Horniolus Weise, 1901 from China are revised, including the description of a new species, Horniolus hainanensis Chen &amp; Ren, sp. n. Horniolus sonduongensis Hoàng, 1979 is reported from China for the first time. A key to the species from China is provided. Nomenclatural history, diagnoses, detailed descriptions, illustrations, and distribution for each species have been provided. A checklist of all known species of this genus is also presented.
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Al Ansi, Amin, Areej A. Alkhalaf, Hassan Fadl, Iftekhar Rasool, and Hathal Al Dhafer. "An annotated checklist of Coccinellidae (Insecta, Coleoptera) with eight new records from the Kingdom of Saudi Arabia." ZooKeys 1006 (December 21, 2020): 35–89. http://dx.doi.org/10.3897/zookeys.1006.59123.

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The members of Coccinellidae are among the most important biological control agents being used throughout the world. The objective of this study was to provide the current scenario of this family in Saudi Arabia. Collection surveys of ladybird individuals were conducted throughout Saudi Arabia. A total of 5400 specimens of Coccinellidae were collected belonging to 51 species in 19 genera and seven tribes: Sticholotidini (2 genera / 10 species), Coccidulini (1/3), Scymnini (4/24), Diomini (1/1), Hyperaspidini (1/3), Chilocorini (3/8), Noviini (1/3), Coccinellini (8/11), Epilachnini (2/2). One genus, Clitostethus, and eight species and subspecies, Cheilomenes lunata lunata (Fabricius, 1775), Clitostethus arcuatus (Rossi, 1794), Nephus ornatulus Korschefsky, 1931, N. nigricans (Weise, 1879), Pharoscymnus fleischeri (Weise, 1883), Novius yemenensis Raimundo &amp;amp; Fürsch, 2006, Scymnus (Scymnus) scapuliferus Mulsant, 1850, and Stethorus endrodyi Fürsch, 1970, are reported as new records to Saudi Arabia. Two endemic species, Scymnus agrumi and S. arabicus, were recorded. This study also describes the geographical distribution for each species and the diagnostic characters for new records.
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Li, Xinghao, Nan Song, and Heng Zhang. "Comparative and phylogenomic analyses of mitochondrial genomes in Coccinellidae (Coleoptera: Coccinelloidea)." PeerJ 9 (December 9, 2021): e12169. http://dx.doi.org/10.7717/peerj.12169.

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The Coccinellidae are one of the most familiar beetle families, the ladybirds. Despite the great ecological and economic significance, the phylogenetic relationships of Coccinellidae remain poorly understood. One of the reasons is that the sequenced mitogenomes available for this family are very limited. We sequenced complete or nearly complete mitogenomes from seven species of the tribe Coccinellini with next-generation sequencing. All species have the same gene content and gene order as the putatively ancestral insect mitogenome. A large intergenic spacer region (&gt; 890 bp) was found located between trnI and trnQ. The potential for using secondary structures of the large and small ribosomal subunits for phylogenetic reconstruction was predicted. The phylogenetic relationships were explored through comparative analyses across more than 30 coccinellid species. We performed phylogenetic analyses with both concatenation methods (Maximum Likelihood and Bayesian Inference) and multispecies coalescent method (ASTRAL). Phylogenetic results strongly supported the monophyly of Coccinellidae. Within Coccinellidae, the Epilachnini and the Coccinellini including Halyziini were monophyletic, while the Scymnini and Coccidulini were non-monophyletic.
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Al, Ansi Amin, Areej A. Alkhalaf, Hassan Fadl, Iftekhar Rasool, and Dhafer Hathal Al. "An annotated checklist of Coccinellidae (Insecta, Coleoptera) with eight new records from the Kingdom of Saudi Arabia." ZooKeys 1006 (December 21, 2020): 35–89. https://doi.org/10.3897/zookeys.1006.59123.

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The members of Coccinellidae are among the most important biological control agents being used throughout the world. The objective of this study was to provide the current scenario of this family in Saudi Arabia. Collection surveys of ladybird individuals were conducted throughout Saudi Arabia. A total of 5400 specimens of Coccinellidae were collected belonging to 51 species in 19 genera and seven tribes: Sticholotidini (2 genera / 10 species), Coccidulini (1/3), Scymnini (4/24), Diomini (1/1), Hyperaspidini (1/3), Chilocorini (3/8), Noviini (1/3), Coccinellini (8/11), Epilachnini (2/2). One genus, Clitostethus, and eight species and subspecies, Cheilomenes lunata lunata (Fabricius, 1775), Clitostethus arcuatus (Rossi, 1794), Nephus ornatulus Korschefsky, 1931, N. nigricans (Weise, 1879), Pharoscymnus fleischeri (Weise, 1883), Novius yemenensis Raimundo &amp; Fürsch, 2006, Scymnus (Scymnus) scapuliferus Mulsant, 1850, and Stethorus endrodyi Fürsch, 1970, are reported as new records to Saudi Arabia. Two endemic species, Scymnus agrumi and S. arabicus, were recorded. This study also describes the geographical distribution for each species and the diagnostic characters for new records.
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28

Chen, Xiaosheng, Xiufeng Xie, Shunxiang Ren, and Xingmin Wang. "Discovery of a New World ladybird beetle Nephaspis indus Gordon, 1996 (Coleoptera: Coccinellidae: Scymnini) on the Island of Taiwan." Biodiversity Data Journal 4 (November 17, 2016): e10537. http://dx.doi.org/10.3897/bdj.4.e10537.

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29

Chen, Xiaosheng, Xiufeng Xie, Shunxiang Ren, and Xingmin Wang. "Discovery of a New World ladybird beetle Nephaspis indus Gordon, 1996 (Coleoptera: Coccinellidae: Scymnini) on the Island of Taiwan." Biodiversity Data Journal 4 (November 17, 2016): e10537. https://doi.org/10.3897/BDJ.4.e10537.

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<i>Nephaspis indus</i> Gordon, 1996 was imported into Taiwan from Hawaii in 1990 as a biological control agent for the spiralling whitefly, <i>Aleurodicus dispersus</i> Russell, 1965 (Hemiptera: Aleyrodidae). However, its establishment was not known prior to this study. <i>Nephaspis indus</i> Gordon, 1996, a natural enemy of <i>Aleurodicus dispersus</i> Russell (Hemiptera: Aleyrodidae) native to the Neotropical region, is recorded as established in Taiwan for the first time. The present paper provides a detailed further description and illustrations of the adult. Diagnostic characters for the genus and species are given and the nomenclature of this species is also discussed.
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30

Soares, António Onofre, Hugo Renato Calado, José Carlos Franco, et al. "An annotated checklist of ladybeetle species (Coleoptera, Coccinellidae) of Portugal, including the Azores and Madeira Archipelagos." ZooKeys 1053 (August 2, 2021): 107–44. http://dx.doi.org/10.3897/zookeys.1053.64268.

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A comprehensive annotated checklist of the ladybeetle species of Portugal, including the Azores and Madeira archipelagos, is presented. The Coccinellidae fauna comprises a total of 101 species: 83 from the Mainland, 39 from Madeira, and 32 from the Azores. The listed species are distributed among 2 sub-families and 13 tribes: within the subfamily Microweiseinae, Madeirodulini (1 species), Serangiini (2 species), and within the subfamily Coccinellinae, Azyini (1 species), Chilocorini (4 species), Coccidulini (7 species), Coccinellini (30 species), Epilachnini (4 species), Hyperaspidini (7 species), Noviini (2 species), Platynaspini (1 species), Scymnini (37 species), Stethorini (3 species), and Sticholotidini (2 species). The Portuguese fauna comprises 10 exotic species: 5 present in the Mainland, 7 in Madeira, and 6 in the Azores. Harmonia axyridis (Pallas, 1773) from Madeira, Propylea quatuordecimpunctata (Linnaeus, 1758) from the Azores, Delphastus catalinae (Horn, 1895) from the Azores and Madeira, Nephus (Geminosipho) reunioni (Fürsch, 1974) and Nephus (Nephus) voeltzkowi Weise, 1910 from Madeira and Microserangium sp. from the Mainland, are reported for the first time. Some species are considered doubtful records, as explained in the text. These results were obtained by compiling information on the available literature regarding ladybeetle species on the Portuguese mainland and insular territories, and original data.
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Iqbal, Zafar, Rashid Azad, Xiao-Sheng Chen, et al. "A New Species of Scymnus (Coleoptera, Coccinellidae) from Pakistan with Mitochondrial Genome and Its Phylogenetic Implications." Insects 15, no. 5 (2024): 371. http://dx.doi.org/10.3390/insects15050371.

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In this study, a new species of the subgenus Pullus belonging to the Scymnus genus from Pakistan, Scymnus (Pullus) cardi sp. nov., was described and illustrated, with information on its distribution, host plants, and prey. Additionally, the completed mitochondrial genome (mitogenome) of the new species using high-throughput sequencing technology was obtained. The genome contains the typical 37 genes (13 protein-coding genes, two ribosomal RNAs, and 22 transfer RNAs) and a non-coding control region, and is arranged in the same order as that of the putative ancestor of beetles. The AT content of the mitogenome is approximately 85.1%, with AT skew and GC skew of 0.05 and −0.43, respectively. The calculated values of relative synonymous codon usage (RSCU) determine that the codon UUA (L) has the highest frequency. Furthermore, we explored the phylogenetic relationship among 59 representatives of the Coccinellidae using Bayesian inference and maximum likelihood methods, the results of which strongly support the monophyly of Coccinellinae. The phylogenetic results positioned Scymnus (Pullus) cardi in a well-supported clade with Scymnus (Pullus) loewii and Scymnus (Pullus) rubricaudus within the genus Scymnus and the tribe Scymnini. The mitochondrial sequence of S. (P.) cardi will contribute to the mitochondrial genome database and provide helpful information for the identification and phylogeny of Coccinellidae.
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32

Soares, António Onofre, Hugo Renato Calado, José Carlos Franco, et al. "An annotated checklist of ladybeetle species (Coleoptera, Coccinellidae) of Portugal, including the Azores and Madeira Archipelagos." ZooKeys 1053 (August 2, 2021): 107–44. https://doi.org/10.3897/zookeys.1053.64268.

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A comprehensive annotated checklist of the ladybeetle species of Portugal, including the Azores and Madeira archipelagos, is presented. The Coccinellidae fauna comprises a total of 101 species: 83 from the Mainland, 39 from Madeira, and 32 from the Azores. The listed species are distributed among 2 sub-families and 13 tribes: within the subfamily Microweiseinae, Madeirodulini (1 species), Serangiini (2 species), and within the subfamily Coccinellinae, Azyini (1 species), Chilocorini (4 species), Coccidulini (7 species), Coccinellini (30 species), Epilachnini (4 species), Hyperaspidini (7 species), Noviini (2 species), Platynaspini (1 species), Scymnini (37 species), Stethorini (3 species), and Sticholotidini (2 species). The Portuguese fauna comprises 10 exotic species: 5 present in the Mainland, 7 in Madeira, and 6 in the Azores. Harmonia axyridis (Pallas, 1773) from Madeira, Propylea quatuordecimpunctata (Linnaeus, 1758) from the Azores, Delphastus catalinae (Horn, 1895) from the Azores and Madeira, Nephus (Geminosipho) reunioni (Fürsch, 1974) and Nephus (Nephus) voeltzkowi Weise, 1910 from Madeira and Microserangium sp. from the Mainland, are reported for the first time. Some species are considered doubtful records, as explained in the text. These results were obtained by compiling information on the available literature regarding ladybeetle species on the Portuguese mainland and insular territories, and original data.
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33

YADAV, G. A. KAVYA, JAYALAXMI NARAYAN HEGDE, C. M. KALLESHWARASWAMY, et al. "Fauna of predaceous coccinellids (Coleoptera: Coccinellidae) associated with mealybugs (Hemiptera: Pseudococcidae) infesting horticultural crops." Journal of Biological Control, April 22, 2025, 47–53. https://doi.org/10.18311/jbc/2025/46238.

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Mealybugs (Hemiptera: Pseudococcidae) are highly polyphagous pests that cause serious damage especially in horticultural crops that are hard to kill by chemicals because of their protective waxy coating. Hence, relying on natural enemies like insect predators is one of the safest and most effective alternative solutions to manage these species. A total of 16 species of predatory coccinellids belonging to 13 genera and five tribes under three subfamilies viz., Coccinellinae, Chilocorinae and Scymninae were found feeding on 11 mealybug species infesting different horticultural crops of Karnataka. These include Anegleis cardoni (Weise), Brumoides suturalis (Fabricius), Cryptolaemus montrouzieri Mulsant, Coccinella transversalis Fabricius, Cheilomenes sexmaculata (Fab.), Chilocorus nigrita (Fab.), Coelophora sp., Jauravia pallidula (Motschulsky), Harmonia octomaculata (Fab.), Hyperaspis maindroni Sicard, Nephus sp., Propylea sp., Scymnus latifolius Poorani, Scymnus coccivora Ayyar, Scymnus sp.1 and Scymnus sp.2. Scymnus coccivora was associated with six diverse mealybug species with highest species composition (29.76%). The horticultural crop ecosystems were found to have a higher per cent species composition of predaceous coccinellids, which naturally regulates mealybug populations by devouring them. Therefore, utilizing these predatory beetles as a biological tool for the management of mealybugs has great potential.
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34

Pervez, Ahmad, Meena Yadav, and Moina Khan. "DIVERSITY OF PREDACEOUS COCCINELLID BEETLES (COLEOPTERA: COCCINELLIDAE) IN UTTARAKHAND, NORTH INDIA." Journal of Mountain Research 15, no. 1 (2020). http://dx.doi.org/10.51220/jmr.v15i1.2.

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A field survey was conducted to explore the diversity of predaceous ladybird beetles (Coleoptera: Coccinellidae) by sampling adults and larvae from five geographical habitats of Uttarakhand, North India. As a result, eighteen predaceous ladybird beetle species were identified belonging to 15 genera and 3 subfamilies: Chilocorinae, Coccinellinae, and Scymninae. These species were: Platynaspis saundersi, Adalia hexaspilota, Alloneda dodecaspilota, Calvia albida, Menochilus sexmaculatus, Coccinella septempunctata, Coccinella transversalis, Harmonia eucharis, Harmonia sedecimnotata; Hippodamia variegata, Illeis cincta, Illeis confusa, Micraspis discolor, Oenopia sauzeti, Propylea dissecta; Propylea luteopustulata; Palaeoneda auriculata and Scymnus posticalis. Of these, six species ladybird beetles are the first time reported from Uttarakhand. Their distinctive characters, distribution, host plant-prey range, active period, and ecological importance have been described. A few of these species may be potentially utilized as biological agents to control many phytophagous insect pests of agriculture and horticulture crops. A disparity was observed in the ladybird-abundance, as both the number of individuals of ladybirds and their frequency in the population differed significantly. The dominance of a few ladybird beetles, particularly C. septempunctata, M. sexmaculatus, C. transversalis and H. variegata was confirmed by the low value of Simpson’s Dominance Index (0.1903) further indicating the disparity in the populations of different ladybird species in a community
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35

Kairo, M. "Cryptolaemus montrouzieri (Mulsant) (Coccinellidae: Scymninae): a review of biology, ecology, and use in biological control with particular reference to potential impact on non-target organisms." CAB Reviews: Perspectives in Agriculture, Veterinary Science, Nutrition and Natural Resources 8, no. 005 (2013). http://dx.doi.org/10.1079/pavsnnr20138005.

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36

Wang, Chaoxin, Ivan Grijalva, Doina Caragea, and Brian McCornack. "Detecting common coccinellids found in sorghum using deep learning models." Scientific Reports 13, no. 1 (2023). http://dx.doi.org/10.1038/s41598-023-36738-5.

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AbstractIncreased global production of sorghum has the potential to meet many of the demands of a growing human population. Developing automation technologies for field scouting is crucial for long-term and low-cost production. Since 2013, sugarcane aphid (SCA) Melanaphis sacchari (Zehntner) has become an important economic pest causing significant yield loss across the sorghum production region in the United States. Adequate management of SCA depends on costly field scouting to determine pest presence and economic threshold levels to spray insecticides. However, with the impact of insecticides on natural enemies, there is an urgent need to develop automated-detection technologies for their conservation. Natural enemies play a crucial role in the management of SCA populations. These insects, primary coccinellids, prey on SCA and help to reduce unnecessary insecticide applications. Although these insects help regulate SCA populations, the detection and classification of these insects is time-consuming and inefficient in lower value crops like sorghum during field scouting. Advanced deep learning software provides a means to perform laborious automatic agricultural tasks, including detection and classification of insects. However, deep learning models for coccinellids in sorghum have not been developed. Therefore, our objective was to develop and train machine learning models to detect coccinellids commonly found in sorghum and classify them according to their genera, species, and subfamily level. We trained a two-stage object detection model, specifically, Faster Region-based Convolutional Neural Network (Faster R-CNN) with the Feature Pyramid Network (FPN) and also one-stage detection models in the YOLO (You Only Look Once) family (YOLOv5 and YOLOv7) to detect and classify seven coccinellids commonly found in sorghum (i.e., Coccinella septempunctata, Coleomegilla maculata, Cycloneda sanguinea, Harmonia axyridis, Hippodamia convergens, Olla v-nigrum, Scymninae). We used images extracted from the iNaturalist project to perform training and evaluation of the Faster R-CNN-FPN and YOLOv5 and YOLOv7 models. iNaturalist is an imagery web server used to publish citizen’s observations of images pertaining to living organisms. Experimental evaluation using standard object detection metrics, such as average precision (AP), AP@0.50, etc., has shown that the YOLOv7 model performs the best on the coccinellid images with an AP@0.50 as high as 97.3, and AP as high as 74.6. Our research contributes automated deep learning software to the area of integrated pest management, making it easier to detect natural enemies in sorghum.
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37

Janakiraman, Poorani, and Roger G. Booth. "Nomenclatural notes on three species of Scymnini (Coleoptera: Coccinellidae) from Ceylon." Oriental Insects, April 5, 2020, 1–13. http://dx.doi.org/10.1080/00305316.2020.1748134.

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38

Soares, António, Isabel Borges, Hugo Calado, and Paulo Borges. "An updated checklist to the biodiversity data of ladybeetles (Coleoptera: Coccinellidae) of the Azores Archipelago (Portugal)." Biodiversity Data Journal 9 (December 16, 2021). http://dx.doi.org/10.3897/bdj.9.e77464.

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A recently-published review from 2021 presents a comprehensive checklist of ladybeetles of Portugal, including the Azores and Madeira Archipelagos. Until then, the available information was very scattered and based on a single revision dating back to 1986, a few international catalogues and databases, individual records and studies on communities of agroecosystems. However, no information was available on faunal composition across the Azorean islands and their habitats, using standardised inventories. Here, we present data about the biodiversity of ladybeetles and their distribution and abundance in five Islands of the Azores (Faial, Graciosa, Pico, São Jorge and São Miguel). Surveys included herbaceous and arboreal habitats from native to anthropogenic-managed habitats: ruderal road vegetation, vegetable garden, mixed forest of endemic and non-native host plants, coastal prairies, coastal mixed vegetation, cornfields and urban areas. We aimed to contribute to the ongoing effort to document the terrestrial biodiversity of Portugal, including the Archipelago of the Azores, within the research project AZORESBIOPORTAL–PORBIOTA (ACORES-01-0145-FEDER-000072). In this study, a total of 1,487 specimens of Coccinellidae belonging to 19 species are reported for several habitats. The listed species are from one single sub-familiy (Coccinellinae) and six tribes; Chilocorini (one species), Coccidulini (three species), Coccinellini (six species), Noviini (one species), Scymnini (seven species), Stethorini (one species). The number of species collected per island differed; Faial (10 species), Graciosa (four species), Pico (seven species), São Jorge (seven species) and São Miguel (12 species). For six species, new island records are given. Currently, the number of species known to occur in the Azores are 32, including two doubtful records. The majority of species are Scymnini, being Scymnus (Scymnus) interruptus (Goeze, 1777) and Scymnus (Scymnus) nubilus Mulsant, 1850, the most abundant species (relative abundance 71.1%). This database will be the baseline of a long-term monitoring project allowing assessment of the impact of ongoing global changes in the distribution and abundance of ladybeetles.
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39

Walton, V. M., and K. L. Pringle. "Effects of Pesticides and Fungicides Used on Grapevines on the Mealybug Predatory Beetle Nephus 'boschianus' (Coccinellidae, Scymnini)." South African Journal of Enology & Viticulture 22, no. 2 (2017). http://dx.doi.org/10.21548/22-2-2204.

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40

De La Pava, Nataly, Christian Sherley Araújo da Silva Torres, and José Maurício S. Bento. "Behavioral responses of the lady beetles Cryptolaemus montrouzieri and Tenuisvalvae notata to specific mealybug prey." Entomologia Experimentalis et Applicata, April 5, 2024. http://dx.doi.org/10.1111/eea.13447.

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AbstractLady beetles (Coleoptera: Coccinellidae) are important biocontrol agents that are considered generalist predators, although some tribes, such as Scymnini, are specialized in predation of sucking pests, such as mealybugs, aphids, and whiteflies. However, prey preference or pre‐imaginal conditioning may occur as coccinellids are subjected to large‐scale rearing. Thus, predator–prey interaction may be an outcome of conditioning to prey stimuli. To evaluate possible pre‐imaginal conditioning, population lines were created for the lady beetles Cryptolaemus montrouzieri Mulsant and Tenuisvalvae notata (Mulsant), each fed for at least eight generations the mealybugs Ferrisia dasylirii (Cockrell) or Planococcus citri Risso (Hemiptera: Pseudococcidae). Next, the behavioral response of these coccinellids was measured in arenas treated with prey volatiles or footprints, regarding walking time, walking speed, walking distance, and residence time. Finally, food preference between prey species was measured in laboratory and semi‐field conditions on infested cotton plants. Conditioned coccinellids responded similarly to volatiles released by either prey species offered. Furthermore, there were no changes in predator behavior towards footprints of either prey species. Both predator species consumed more P. citri in all tests, regardless of rearing prey indicating no pre‐imaginal conditioning. These results suggest that rearing prey may not affect the predator behavior, and both coccinellid species are effective in the biological control of mealybugs.
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