Academic literature on the topic 'Shelterwood Harvesting'

We present 27-year results from a comparison of clear-cutting and shelterwood harvesting in the boreal forest of Alaska. Three patch clear-cut and three shelterwood units were harvested in 1972; about 100 dispersed white spruce (Picea glauca (Moench) Voss) leave trees per hectare were retained in the shelterwoods. Units were mechanically scarified and an exceptionally large seed-crop was dispersed that year. Shelterwood trees were removed after 15 years. After 27 years, overstory treatment had no effect on the density or growth of the species we studied, while scarification had highly significant effects. In 1999, scarified areas were densely populated with white spruce seedlings and saplings (118 000 – 129 000 stems/ha, with spruce in 100% of plots). Unscarified areas had far fewer spruce stems but were nevertheless well stocked (11 000 – 15 000 stems/ha, with 87% frequency). Initially, spruce grew best on scarified surfaces, but by 27 years, growth of the tallest spruce saplings was significantly greater on unscarified than scarified surfaces. By 27 years, cover of the grass Calamagrostis canadensis (Michx.) Nutt. had returned to preharvest levels in all treatment types. Because criteria for evaluating forest management practices have changed since this study was begun, partial overstory retention systems for the management of Alaska's boreal forest deserve further study.

Beech (<I>Fagus orientalis</I> Lipsky) is the most industrial commercial tree species among more than 80 broadleaved trees and shrubs in Hyrcanian forests. In this study some of the silvicultural properties of beech were studied under a shelterwood cutting system in a regeneration unit of 25 years old stand in Sangdeh forests of Iran. Tree height, diameter, canopy cover, variety of species, frequency at different stages of growth (seedling, sapling, thicket and pole) and tree density at juvenile phase were recorded in 431 circular 1,000 m<sup>2</sup> sample plots before and after shelterwood cutting. Results indicated that tree density in the diameter at breast height (dbh) class of < 20 cm increased during shelterwood cutting. Canopy cover of regeneration unit was 78.5% in 1957, while in 2000 and after the shelterwood cutting it reached 59%. Growing stages of regeneration were 14% sapling, 21% thicket, 24% small pole and 41% pole. The shelterwood method has not been very successful in our study area due to a number of technical and potential problems.

In forest ecosystems, soil CO2 efflux is an important component of ecosystem respiration (RE), which is generally driven by variability in soil temperature and soil moisture. Tree harvesting in forests can alter the soil variables and, consequently, impact soil CO2 efflux. This study investigated the response of total soil CO2 efflux, and its components, to a shelterwood harvesting event of a mature temperate white pine (Pinus strobus L.) forest located in Southern Ontario, Canada. The objective was to explore the response of soil CO2 effluxes to changes in the forest microclimate, such as soil temperature and soil moisture, after shelterwood harvesting removed approximately one-third of the overstory canopy. No significant differences were found in both soil temperature and soil moisture between the pre-harvesting (2008–2011) and post-harvesting (2012–2014) periods. Despite similar soil microclimates, total soil CO2 effluxes were significantly reduced by up to 37%. Soil CO2 effluxes from heterotrophic sources were significantly reduced post-harvesting by approximately 27%, while no significant difference in the mineral-soil horizon sources were measured. An analysis of RE, measured with an eddy covariance tower over the study area, showed an increase post-harvesting. However, the overall net ecosystem carbon exchange showed no significant difference between pre- and post-harvesting. This was due to an increase in the gross ecosystem productivity post-harvesting, compensating for the increased losses (i.e., increased RE). This study highlights the complexities of soil CO2 efflux after a disturbance, such as a harvest. The knowledge gained from this study adds to our understanding of how shelterwood harvesting may influence ecosystem carbon exchange and will be useful for forest managers focused on carbon sequestration and forest conservation.

Ecosystem-based management aims to preserve old-growth forest attributes using techniques mimicking natural disturbances. One such technique is irregular shelterwood logging, but its impacts on forest floor organisms are poorly known. Our objective was to quantify the effects of three different treatments of irregular shelterwood on population structure and the body condition of the eastern red-backed salamander (Plethodon cinereus) 5–6 years following harvesting. A total of 64 sites in western Québec were sampled using artificial refugia and quadrat searches. Large salamanders (>32 mm) were more abundant in the gap treatment than in strip, uniform, or control treatments. Small salamanders (≤32 mm) followed the same pattern, although the differences were marginal. For a given treatment, small salamanders were as abundant as large salamanders. Salamander body condition differed between the 2 years of sampling but did not differ among treatments, regardless of salamander size. We conclude that environmental conditions in irregular shelterwood treatments 5–6 years following harvesting support populations of small vertebrates on the forest floor.

This long-term experiment in Maine, U.S.A., was designed to provide information on the best silvicultural practices for managing stands of mixed northern conifers in northeastern U.S.A. We evaluated growth and yield and changes in species composition, quality, and structure during the first 40 years of the experiment. Replicated treatments include the selection system, uniform shelterwood, unregulated harvesting, and diameter-limit cutting. The new cohort established under three-stage shelterwood was subsequently left untreated or precommercially thinned. Between-treatment differences in net volume growth were not significant (α = 0.10), though gross volume growth differed significantly for managed vs. unmanaged, selection vs. shelterwood, and shelterwood vs. diameter-limit treatments. A three-stage shelterwood method with precommercial thinning 10 years following final overstory removal resulted in good control of hardwoods and hemlock and a dramatic increase in spruce and fir. The selection system on a 5-year cutting cycle resulted in increased hemlock, spruce, and fir, with a decrease in hardwood species. If the primary goal were production, even-aged management would most likely be preferred. We recommend a two-stage shelterwood method as applied in this experiment with some modification to improve species composition and stand quality. Stand quality (proportion of stand volume in cull trees) and species composition was influenced by treatment.

The Montane Alternative Silvicultural Systems (MASS) partnership was formed in 1992 to test new approaches to harvesting high-elevation old-growth forests of Vancouver Island, British Columbia. Uniform Shelterwood (SW), Green Tree Retention (GT) and Patch Cuttings (PC) were tested as alternatives to Clearcutting (CC). All systems were compared to an Old Growth (OG) control. Harvesting was completed in 1993. This paper summarizes the conclusions from the operational aspects of the study and reviews some early results from studies of the biological impacts. Harvesting costs to roadside were about 12% greater for the PC and GT and 49% greater for the SW. Cumulative losses to windthrow after three years were 25% in the GT and 5% in the SW. Site disturbance, soil compaction and rates of organic matter decomposition were low across all harvested treatments. Rates of organic matter decomposition and N mineralization on SW, GT and PC were intermediate between CC and OG. Nutrient loss in soil solution was reduced with partial harvesting but lack of available nitrogen is limiting the early growth of coniferous regeneration among all systems. During spring and summer, SW and CC treatments were generally cooler than PC or GT. Damage to understory vegetation and advanced conifer regeneration was least in the SW, which also provided the greatest amounts of seedfall among all harvested treatments. Key words: Abies amabilis, Tsuga heterophylla, Thuja plicata, Chamaecyparis nootkatensis, harvesting costs, site impacts, silvicultural systems, shelterwood

Using inventory data from a long-term silviculture experiment in east-central Maine, spatial models were developed to analyze 28 years (1974–2002) of stand structural dynamics. Differences in spatial pattern, species mingling, height differentiation, and relative stand complexity index (rSCI) were compared among five treatments: commercial clear-cutting, fixed diameter-limit, 5 year single-tree selection, three-stage shelterwood (both with and without precommercial thinning), and unharvested natural areas. Regardless of treatment, regeneration events (whether induced by natural breakup of the overstory or by harvesting) increased aggregation in spatial pattern and reduced species mingling, more so in the commercial clearcut and fixed diameter-limit treatments where hardwood densities were highest. Regular spatial patterns were rare. Height differentiation values for individual trees and stand-level mean rSCI were generally highest in untreated natural areas and 5 year selection treatments, intermediate in commercial clearcut and fixed diameter-limit treatments, and lowest in three-stage shelterwood treatments. After a brief adjustment period, precommercial thinning in a shelterwood treated stand generally increased species mingling, height differentiation, and rSCI. Two untreated natural areas exhibited divergent pathways of structural development. Dynamics in uneven-aged selection treatments more closely resembled that of the untreated natural areas than did the shelterwood, commercial clearcut, or fixed diameter-limit treatments.

Less intensive harvesting methods (e.g., selection cutting, shelterwood cuttings) are recommended as alternatives to clearcutting for maintaining mature forest biodiversity in the process of forest regeneration. However, the long-term impact of low-intensity harvesting methods has rarely been studied. Our aim was to clarify the long-term effects of repeated selective cutting, thinning, and shelterwood cutting on the richness, abundance, and species composition of vascular plants, bryophytes, and lichens in Scots pine forests (Pinus sylvestris L.). Data were collected from 25 mature stands located in dry Podzols in southwestern Estonia with a known management history for the last 70 years. Altogether, 35 vascular plant, 41 bryophyte, and 78 lichen species were recorded, including five species with conservational value. Generally, the management history was not related to species richness, except a negative correlation with the species number of epiphytic lichens on conifer trees. In addition, the abundance of two lichen species from the genus Chaenotheca was lower in more frequently managed stands. Species richness and composition were most strongly affected by soil pH and light conditions. We conclude that long-term, low-intensity harvesting of Scots pine forests on nutrient-poor Podzols maintains suitable habitats for vascular plants, bryophytes, and lichens, confirming its suitability for sustainable forest regeneration.

The MASS (Montane Alternative Silvicultural Systems) trial was established in the coastal mountains of British Columbia to compare clearcut, patch cut, green tree and shelterwood systems. A number of studies were carried out at the MASS trial to determine the extent to which these variable levels of stand retention retained old-growth attributes of N cycling and associated ecological processes. Harvesting led to increases in N mineralization in the forest floor (2×) and mineral soil (10×), and fluxes of N through the upper 25 cm of mineral soil (2× to 3×). However, fluxes of N were not large (< 0.35 kg ha-1 per growing season). Nitrogen mineralized was predominantly ammonium and not nitrate in the forest floor (> 95% in all but clearcut, > 75% in clearcut) and upper mineral soil horizon (42–86%). The nitrate component came from heterotrophic decomposition of organic matter, not conversion of ammonium to nitrate by autotrophs, and nitrate increases resulted from decreased gross nitrate consumption with harvesting, rather than increased nitrate production. The increases in soil N availability resulting from harvesting were reflected in only slight increases in seedling foliar N concentrations for two to four years after logging (peak of ~ 2% for western hemlock and ~ 1.6% for amabilis fir) before decreasing to below deficiency thresholds for both species. Overall, estimated losses of N from the rooting zone after harvesting (1 kg ha-1 yr-1) were minimal in comparison to estimated N inputs (4 kg ha-1 yr-1), N exports in logs at harvesting (250 kg ha-1) and soil reserves (11 400 kg ha-1). Although unlikely to affect future site productivity, losses of N could be reduced somewhat through the use of shelterwood harvesting. Key words: alternative silvicultural systems, variable retention harvesting, nitrogen cycling, litterfall, decomposition, nitrification, N mineralization, microbial ecology, leaching, foliar N, Abies amabilis, Tsuga heterophylla, Thuja plicata

Oaks (Quercus spp.) are an important part of the forested landscape in the eastern United States. Although oak is increasing in standing volume, an oak regeneration bottleneck has occurred throughout its range in recent decades. Subsequently, as oak overstory is being harvested, rarely is oak recruited into the overstory to maintain the historic dominance of overstory oak. In the absence of fire and subsequent canopy closure, mesic species have proliferated, frequently forming a dense understory, inhibiting oak regeneration success. This study was developed to determine species dynamics between oak and oak competitors in response to silvicultural treatments in the southern Appalachian Mountains of North Carolina. The treatments were: a shelterwood treatment (25-30% basal area reduction through mid-story removal with herbicides), a prescribed fire treatment (two late dormant season fires occurred over a 9-year period), a shelterwood and burn treatment (prescribed fire 3-5 years following 30-40% basal area removal), and an unmanaged control. To determine treatment impacts on the regeneration layer, importance value and stems ha-1 were calculated at the species group and individual species level 0- and 9- years post initial treatment. A principal component analysis and an analysis of basal area by treatment 0- and 9-years post-treatment were used to determine the influence of site-specific characteristics on regeneration layer response. The greatest relative increases in importance values were 1401% and 2995% for the red oak group and yellow-poplar (Liriodendron tulipifera), respectively, in the shelterwood and burn (SWB). Change in all species groups were predominantly influenced by the smallest size-class (<0.6 m tall), with the exception of northern red oak (Q. rubra) and yellow-poplar in the SWB. The SWB significantly reduced importance values of all shade tolerant species groups and was the only treatment to decrease red maple (Acer rubrum) importance value and density over the study years. The prescribed fire (RXF) treatment increased red oak group importance value, while simultaneously decreasing yellow-poplar importance value and increasing red maple importance value. Changes in the red oak group in the SWB and the RXF were driven by northern red oak and scarlet oak (Q. coccinea), respectively. Treatments do not appear to change the competitive status of the white oak group. Elevation was closely associated with the red oak group. Yellow-poplar importance value increases, white oak group importance value increases, and site index were closely associated. Decreases in basal area were greatest in the SWB, and the SWB was the only treatment to significantly decrease overstory basal area. The RXF and SWB treatments improved the competitive status of only some oak species, but modifications to these treatments may result in better control of yellow-poplar and red maple competition, further improving oak's competitive status. Site specific factors such as elevation and site index may have impacted the regeneration layer response to treatments.<br>Master of Science<br>Oak-hickory (Quercus and Carya spp.) and oak-pine (Quercus and Pinus spp.) forest types occupy approximately 57 million and 11 million hectares of forestland in the eastern United States, respectively. Oaks are considered ecological and economic keystone species throughout the eastern U.S and maintenance of this genus in eastern U.S. forests has been a primary regional focus for decades. Historic disturbance regimes are estimated to have been much different than they are today. Fire was a common disturbance mechanism prior to fire suppression in the early 20th century. Frequent fires maintained much of the oak component historically. In the absence of fire, the species found in the understories of mature oak stands are commonly mesophytic species, such as yellow-poplar (Liriodendron tulipifera L.) and red maple (Acer rubrum L.). Over the last several decades, research has been conducted to investigate the impacts of treatments targeting the promotion of oak regeneration, but results have been varied and valuable long-term studies are rare. To determine the effects of treatments on the regeneration dynamics of oak and its competitors, four treatments were compared in the southern Appalachian Mountains. Treatments included a control, shelterwood harvest (SW), prescribed fire (RXF), and a shelterwood and burn (SWB). Stand structure and composition were monitored over a 9-year period post-treatment. Overall, results indicate the shelterwood and burn treatment has the greatest potential to improve the competitiveness of the red oak group in the regeneration layer, but yellow-poplar competition in the shelterwood and burn will need be addressed, considering its large increases in this treatment. Although increases in the red oak group were not as great as increases in the RXF treatment compared with the SWB, fire does show promise as a method to increase oak regeneration success. Changes in red oak group importance value varied with elevation, emphasizing results of treatments can be affected by site characteristics. Treatments were not successful at enhancing the competitive status of white oak (Quercus alba L.). Silvicultural treatments can be used to improve the competitive status of oak on sites in the southern Appalachian Mountains, but close monitoring of species dynamics throughout the rotation are needed to ensure long-term oak success.

The vegetation responses to seven silvicultural treatments one growing season after harvesting were examined on seven sites in the southern Appalachian mountains of Virginia and West Virginia. Treatments included: 1) control, 2) understory control by herbicide, 3) group selection, 4) high-leave shelterwood, 5) low-leave shelterwood, 6) leave tree, and 7) clearcut. The effects of harvesting were compared between treatments and between pre-harvest and post-harvest samplings. Species richness, percent cover, and local species extinctions were calculated for sample plots ranging in size from 1m2 to 2 ha. Vegetation richness and cover increased with increasing harvest intensity. Local species extinctions were similar in the control and disturbed treatments. Additional analyses were performed using the control, high-leave shelterwood, and clearcut on five of the seven sites to determine the relationships between soil, litter, and other environmental characteristics and vegetation in the herbaceous layer (<1 m in height). Multivariate analysis techniques were used to analyze average differences in species abundance between pre-harvest and post-harvest and to relate post-harvest vegetation to microsite characteristics. Regional-scale differences in site location were more important in explaining the presence of a species than were environmental characteristics. Within a region, species primarily were distributed along a light/litter weight gradient and secondarily along a soil properties and nutrient gradient.<br>Master of Science