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1

Denham, K. L., and J. Long. "Occurrence of Gyrodactylus thymalli Zitnan, 1960 on grayling, Thymallus thymallus (L.), in England." Journal of Fish Diseases 22, no. 3 (1999): 247–52. http://dx.doi.org/10.1046/j.1365-2761.1999.00166.x.

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2

OLSTAD, K., A. P. SHINN, L. BACHMANN, and T. A. BAKKE. "Host-based identification is not supported by morphometrics in natural populations of Gyrodactylus salaris and G. thymalli (Platyhelminthes, Monogenea)." Parasitology 134, no. 14 (2007): 2041–52. http://dx.doi.org/10.1017/s0031182007003332.

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SUMMARYGyrodactylus salaris is a serious pest of wild pre-smolt Atlantic salmon (Salmo salar) in Norway. The closely related G. thymalli, originally described from grayling (Thymallus thymallus), is assumed harmless to both grayling and salmon. The 2 species are difficult to distinguish using traditional, morphometric methods or molecular approaches. The aim of this study was to explore whether there is a consistent pattern of morphometrical variation between G. salaris and G. thymalli and to analyse the morphometric variation in the context of ‘diagnostic realism’ (in natural populations). Sp
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3

PETTERSEN, RUBEN ALEXANDER, TOR ATLE MO, HAAKON HANSEN, and LEIF ASBJØRN VØLLESTAD. "Genetic population structure of Gyrodactylus thymalli (Monogenea) in a large Norwegian river system." Parasitology 142, no. 14 (2015): 1693–702. http://dx.doi.org/10.1017/s003118201500133x.

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SUMMARYThe extent of geographic genetic variation is the result of several processes such as mutation, gene flow, selection and drift. Processes that structure the populations of parasite species are often directly linked to the processes that influence the host. Here, we investigate the genetic population structure of the ectoparasite Gyrodactylus thymalli Žitňan, 1960 (Monogenea) collected from grayling (Thymallus thymallus L.) throughout the river Glomma, the largest watercourse in Norway. Parts of the mitochondrial dehydrogenase subunit 5 (NADH 5) and cytochrome oxidase I (COI) genes from
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4

Curtean-Bănăduc, Angela, and Doru Bănăduc. "Thymallus thymallus (Linnaeus, 1758), Ecological Status in Maramureş Mountains Nature Park (Romania)." Transylvanian Review of Systematical and Ecological Research 18, no. 2 (2016): 53–68. http://dx.doi.org/10.1515/trser-2015-0087.

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Abstract Thymallus thymallus is considered a species of significant protective importance within the Vişeu Watershed. The state of habitats characteristically inhabited by Thymallus thymallus within the Maramureş Mountains Nature Park is balanced between reduced (one third of the lotic sectors where the species was identified), average (one third) and good (one third). The excellent conservation status is currently missing for populations of this fish in the Vișeu Basin. Human impact types identified as contributing towards the decreasing state of Thymallus thymallus habitats and therefore pop
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5

Hrytsyniak, Ir, and T. Shvets. "European grayling (Thymallus thymallus Linnaeus, 1758). Thematic bibliography." Ribogospodarsʹka nauka Ukraïni., no. 2(68) (June 28, 2024): 132–63. http://dx.doi.org/10.61976/fsu2024.02.132.

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Purpose. Forming a thematic bibliographic list of English-language and Ukrainian-language publications, which highlight the issues of artificial breeding and cultivation of European grayling in Ukraine and abroad, the results of research into its population, ecological, genetic, physiological, biochemical, immunological and other characteristics. Methods. The complete and selective methods were applied in the process of the systematic search. The bibliographic core have been consisted of scientific publications in Ukrainian and English from the fund of the Institute of Fisheries NAAS Scientifi
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6

NYGREN, AXEL, BIRGIT NILSSON, and MARGARETE JAHNKE. "Cytological studies in Thymallus thymallus and Coregonus albula." Hereditas 67, no. 2 (2009): 269–74. http://dx.doi.org/10.1111/j.1601-5223.1971.tb02379.x.

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7

Plaisance, Laetitia, Tine Huyse, D. T. J. Littlewood, Tor A. Bakke, and Lutz Bachmann. "The complete mitochondrial DNA sequence of the monogenean Gyrodactylus thymalli (Platyhelminthes: Monogenea), a parasite of grayling (Thymallus thymallus)." Molecular and Biochemical Parasitology 154, no. 2 (2007): 190–94. http://dx.doi.org/10.1016/j.molbiopara.2007.04.012.

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8

MEIER, W., and T. WAHLI. "Viral haemorrhagic septicaemia (VHS) in grayling, Thymallus thymallus L." Journal of Fish Diseases 11, no. 6 (1988): 481–87. http://dx.doi.org/10.1111/j.1365-2761.1988.tb00747.x.

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9

Kodela, Tjaša, Dušan Jesenšek, and Simona Sušnik Bajec. "Reproduction parameters of hatchery reared grayling (Thymallus thymallus) females." Animal Reproduction Science 249 (February 2023): 107196. http://dx.doi.org/10.1016/j.anireprosci.2023.107196.

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10

Mallet, J. P., S. Charles, H. Persat, and P. Auger. "Growth modelling in accordance with daily water temperature in European grayling (Thymallus thymallusL.)." Canadian Journal of Fisheries and Aquatic Sciences 56, no. 6 (1999): 994–1000. http://dx.doi.org/10.1139/f99-031.

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The model of von Bertalanffy has been and is still widely used to model fish growth, mainly because of its good description of annual growth over the whole life span. However, it does not take into account a seasonal variability in growth rate, an important phenomenon that appears quite well correlated with water temperature fluctuations in temperate climates. In the present study, we demonstrated that it was possible to model such variations by including daily water temperature in the von Bertalanffy growth formula owing to the correlation between the growth coefficient k and water temperatur
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11

Sempeski, P., P. Gaudin, H. Persat, and O. Grolet. "Diet selection in early-life stages of grayling (Thymallus thymallus)." Archiv für Hydrobiologie 132, no. 4 (1995): 437–52. http://dx.doi.org/10.1127/archiv-hydrobiol/132/1995/437.

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12

Gadd, T., S. Viljamaa-Dirks, R. Holopainen, P. Koski, and M. Jakava-Viljanen. "Characterization of perch rhabdovirus (PRV) in farmed grayling Thymallus thymallus." Diseases of Aquatic Organisms 106, no. 2 (2013): 117–27. http://dx.doi.org/10.3354/dao02654.

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13

CARMIE, H., B. MORELET, G. MAISSE, L. JONARD, and R. CUINAT. "Observations sur la reproduction artificielle de l'ombre commun (Thymallus thymallus)." Bulletin Français de la Pêche et de la Pisciculture, no. 296 (1985): 2–16. http://dx.doi.org/10.1051/kmae:1985003.

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14

Калинин, А. А., И. Ф. Куприянова, A. A. Kalinin та I. F. Kupriyanova. "Мелкие млекопитающие в питании европейского хариуса (Thymallus thymallus, Thymallidae, Salmoniformes)". Зоологический журнал 95, № 6 (2016): 712–19. http://dx.doi.org/10.7868/s0044513416060106.

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15

Bardonnet, Agnès, and Philippe Gaudin. "Diel pattern of emergence in grayling (Thymallus thymallus Linnaeus, 1758)." Canadian Journal of Zoology 68, no. 3 (1990): 465–69. http://dx.doi.org/10.1139/z90-068.

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The emergence rhythm of grayling fry (Thymallus thymallus) was studied under semi-natural rearing conditions, using special traps buried in the gravel. The diel pattern of emergence proved to be mainly diurnal, with a maximum just after sunrise. The discussion emphasizes the fact that this distinguishes the grayling from other salmonids, and since eggs are buried at a reduced depth, proposes the hypothesis that light conditions during development influence the daily pattern of emergence.
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16

Ring, O., H. Jansson, T. ÖST, and T. Andersson. "Inheritance of Allozymes in the European Grayling (Thymallus Thymallus L.)." Hereditas 120, no. 2 (2004): 159–63. http://dx.doi.org/10.1111/j.1601-5223.1994.00159.x.

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17

Thomassen, Gaute, Nicola J. Barson, Thrond O. Haugen, and L. Asbjørn Vøllestad. "Contemporary divergence in early life history in grayling (Thymallus thymallus)." BMC Evolutionary Biology 11, no. 1 (2011): 360. http://dx.doi.org/10.1186/1471-2148-11-360.

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18

Swatdipong, Akarapong, Craig R. Primmer, and Anti Vasemägi. "Historical and recent genetic bottlenecks in European grayling, Thymallus thymallus." Conservation Genetics 11, no. 1 (2009): 279–92. http://dx.doi.org/10.1007/s10592-009-0031-x.

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19

BARDONNET, AGNES, PHILIPPE GAUDIN, and HENRI PERSAT*. "Microhabitats and diel downstream migration of young grayling (Thymallus thymallm L.)." Freshwater Biology 26, no. 3 (1991): 365–76. http://dx.doi.org/10.1111/j.1365-2427.1991.tb01404.x.

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20

Denys, Gaël P.J., Giulia Secci-Petretto, and André Gomes-Dos-Santos. "The complete mitochondrial genome of Thymallus thymallus (Linnaeus, 1758) (Actinopterygii, Salmonidae) obtained by long range PCRs and double multiplexing." Cybium 44, no. 2 (2020): 91–94. https://doi.org/10.26028/cybium/2020-442-001.

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Denys, Gaël P.J., Secci-Petretto, Giulia, Gomes-Dos-Santos, André (2020): The complete mitochondrial genome of Thymallus thymallus (Linnaeus, 1758) (Actinopterygii, Salmonidae) obtained by long range PCRs and double multiplexing. Cybium 44 (2): 91-94, DOI: 10.26028/cybium/2020-442-001, URL: http://dx.doi.org/10.26028/cybium/2020-442-001
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21

Kucheruk, A., I. Hrytsyniak, A. Mruk, and I. Velykopolsky. "PECULIARITIES OF RAISING YOUNG-OF-THE-YEAR EUROPEAN GRAYLING (THYMALLUS THYMALLUS L.)." Ribogospodarsʹka nauka Ukraïni, no. 2 (June 10, 2015): 31–40. http://dx.doi.org/10.15407/fsu2015.02.031.

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22

Persat, Henri, Karin Mattersdorfer, Stéphanie Charlat, Tamara Schenekar, and Steven Weiss. "Genetic integrity of the European grayling (Thymallus thymallus) populations within the Vienne River drainage basin after five decades of stockings." Cybium 40, no. 1 (2016): 7–20. https://doi.org/10.26028/cybium/2016-401-001.

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Persat, Henri, Mattersdorfer, Karin, Charlat, Stéphanie, Schenekar, Tamara, Weiss, Steven (2016): Genetic integrity of the European grayling (Thymallus thymallus) populations within the Vienne River drainage basin after five decades of stockings. Cybium 40 (1): 7-20, DOI: 10.26028/cybium/2016-401-001, URL: http://dx.doi.org/10.26028/cybium/2016-401-001
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23

Persat, Henri, Karin Mattersdorfer, Stéphanie Charlat, Tamara Schenekar, and Steven Weiss. "Genetic integrity of the European grayling (Thymallus thymallus) populations within the Vienne River drainage basin after five decades of stockings." Cybium 40, no. 1 (2016): 7–20. https://doi.org/10.26028/cybium/2016-401-001.

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Persat, Henri, Mattersdorfer, Karin, Charlat, Stéphanie, Schenekar, Tamara, Weiss, Steven (2016): Genetic integrity of the European grayling (Thymallus thymallus) populations within the Vienne River drainage basin after five decades of stockings. Cybium 40 (1): 7-20, DOI: 10.26028/cybium/2016-401-001, URL: http://dx.doi.org/10.26028/cybium/2016-401-001
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24

Persat, Henri, Karin Mattersdorfer, Stéphanie Charlat, Tamara Schenekar, and Steven Weiss. "Genetic integrity of the European grayling (Thymallus thymallus) populations within the Vienne River drainage basin after five decades of stockings." Cybium 40, no. 1 (2016): 7–20. https://doi.org/10.26028/cybium/2016-401-001.

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Persat, Henri, Mattersdorfer, Karin, Charlat, Stéphanie, Schenekar, Tamara, Weiss, Steven (2016): Genetic integrity of the European grayling (Thymallus thymallus) populations within the Vienne River drainage basin after five decades of stockings. Cybium 40 (1): 7-20, DOI: 10.26028/cybium/2016-401-001, URL: http://dx.doi.org/10.26028/cybium/2016-401-001
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25

Persat, Henri, Karin Mattersdorfer, Stéphanie Charlat, Tamara Schenekar, and Steven Weiss. "Genetic integrity of the European grayling (Thymallus thymallus) populations within the Vienne River drainage basin after five decades of stockings." Cybium 40, no. 1 (2016): 7–20. https://doi.org/10.26028/cybium/2016-401-001.

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Persat, Henri, Mattersdorfer, Karin, Charlat, Stéphanie, Schenekar, Tamara, Weiss, Steven (2016): Genetic integrity of the European grayling (Thymallus thymallus) populations within the Vienne River drainage basin after five decades of stockings. Cybium 40 (1): 7-20, DOI: 10.26028/cybium/2016-401-001, URL: http://dx.doi.org/10.26028/cybium/2016-401-001
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26

Didenko, A. V., E. M. Talabishka, I. I. Velykopolskiy, F. F. Kurtyak, and A. I. Kucheruk. "Current Distribution Of The European Grayling, Thymallus Thymallus, And Huchen, Hucho Hucho, In The Transcarpatian Region Of Ukraine." Vestnik Zoologii 52, no. 1 (2018): 65–74. https://doi.org/10.2478/vzoo-2018-0009.

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Didenko, A. V., Talabishka, E. M., Velykopolskiy, I. I., Kurtyak, F. F., Kucheruk, A. I. (2018): Current Distribution Of The European Grayling, Thymallus Thymallus, And Huchen, Hucho Hucho, In The Transcarpatian Region Of Ukraine. Vestnik Zoologii 52 (1): 65-74, DOI: 10.2478/vzoo-2018-0009, URL: http://dx.doi.org/10.2478/vzoo-2018-0009
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27

Papakostas, Spiros, L. Asbjørn Vøllestad, Craig R. Primmer, and Erica H. Leder. "Proteomic Profiling of Early Life Stages of European Grayling (Thymallus thymallus)." Journal of Proteome Research 9, no. 9 (2010): 4790–800. http://dx.doi.org/10.1021/pr100507s.

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28

Nykänen, M., and A. Huusko. "Transferability of habitat preference criteria for larval European grayling (Thymallus thymallus)." Canadian Journal of Fisheries and Aquatic Sciences 61, no. 2 (2004): 185–92. http://dx.doi.org/10.1139/f03-156.

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We examined the transferability of habitat preference criteria for larval European grayling (Thymallus thymallus) to two rivers in Finland by testing whether there were significant positive rank correlations between local fish densities (shoals per square metre) and preference indices for depth, velocity, substrate, and vegetation cover or selected combinations. Two sets of preference curves, one obtained from literature for the River Pollon, France, and another for the River Kuusinkijoki, Finland, were tested. All transferability tests for water velocity criteria were successful, correlation
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29

Khandozhivska, A. "SIZE-WEIGHT INDICES OF EUROPEAN GRAYLING (THYMALLUS THYMALLUS) IN TRANSCARPATHIAN RIVERS." Ribogospodarsʹka nauka Ukraïni, no. 3(25) (September 20, 2013): 89–96. http://dx.doi.org/10.15407/fsu2013.03.089.

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30

MAISSE, G., H. CARMIE, and L. JONARD. "Influence de la température sur l'ovulation de l'ombre commun (Thymallus thymallus)." Bulletin Français de la Pêche et de la Pisciculture, no. 305 (1987): 54–60. http://dx.doi.org/10.1051/kmae:1987007.

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31

Koskinen, M. T., J. Nilsson, A. Je Veselov, A. G. Potutkin, E. Ranta, and C. R. Primmer. "Microsatellite data resolve phylogeographic patterns in European grayling, Thymallus thymallus, Salmonidae." Heredity 88, no. 5 (2002): 391–401. http://dx.doi.org/10.1038/sj.hdy.6800072.

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32

Sempeski, P., and P. Gaudin. "Size-related changes in diel distribution of young grayling (Thymallus thymallus)." Canadian Journal of Fisheries and Aquatic Sciences 52, no. 9 (1995): 1842–48. http://dx.doi.org/10.1139/f95-177.

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An approach based on direct observation from the riverbank allowed a precise description of the size-dependent distribution patterns of young grayling. The smallest larvae (15–20 mm) were mainly observed in the upper layers of the water column and very close to the riverbank. With increasing age and size (20–35 mm), old larvae and young parr began to hold positions closer to the bottom and to the edge of the main channel. From a size of 35–40 mm. an increasing number of juveniles were observed in the river channel, holding a benthic position. A diel habitat shift was also shown for individuals
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33

Vøllestad, L. A., E. Fjeld, T. Haugen, and S. A. Oxnevad. "Developmental instability in grayling (Thymallus thymallus) exposed to methylmercury during embryogenesis." Environmental Pollution 101, no. 3 (1998): 349–54. http://dx.doi.org/10.1016/s0269-7491(98)00055-4.

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34

HOFFMANN, R. W., TH FISCHER-SCHERL, and C. PFEIL-PUTZIEN. "Lymphosarcoma in a wild grayling, Thymallus thymallus L.: a case report." Journal of Fish Diseases 11, no. 3 (1988): 267–70. http://dx.doi.org/10.1111/j.1365-2761.1988.tb00548.x.

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35

Hlavová, Věra. "Reference values of the haematological indices in grayling (Thymallus thymallus linnaeus)." Comparative Biochemistry and Physiology Part A: Physiology 105, no. 3 (1993): 525–32. http://dx.doi.org/10.1016/0300-9629(93)90429-8.

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36

Surre, C., H. Persat, and J. M. Gaillard. "A biometric study of three populations of the European grayling, Thymallus thymallus (L.), from the French Jura Mountains." Canadian Journal of Zoology 64, no. 11 (1986): 2430–38. http://dx.doi.org/10.1139/z86-363.

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Three populations of grayling (Thymallus thymallus L.) from the Jura Mountains (Rhône drainage basin, France) were compared using 8 meristic and 10 morphometric characteristics; discriminant analysis clearly distinguished these populations. This result raises a number of questions about the specificity of fish populations inhabiting any given river and the usefulness of stocking that has been carried out up to now without any controls.
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37

Czeczuga, Bazyli, Andrzej Witkowski, and Mieczysław Kowalewski. "Carotenoids in fish - XXXIX. Presence of salmoxanthin in Thymallus thymallus (L.) specimens." Acta Ichthyologica et Piscatoria 15, no. 1 (1985): 73–81. http://dx.doi.org/10.3750/aip1985.15.1.04.

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38

Hayes, Daniel S., Erwin Lautsch, Günther Unfer, et al. "Response of European grayling, Thymallus thymallus, to multiple stressors in hydropeaking rivers." Journal of Environmental Management 292 (August 2021): 112737. http://dx.doi.org/10.1016/j.jenvman.2021.112737.

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39

Marić, Saša, Belma Kalamujić, Aleš Snoj, et al. "Genetic variation of European grayling (Thymallus thymallus) populations in the Western Balkans." Hydrobiologia 691, no. 1 (2012): 225–37. http://dx.doi.org/10.1007/s10750-012-1076-2.

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40

Purtscher, Ulrich, and Uwe H. Humpesch. "Egg size relationships in five species of salmonids and Thymallus thymallus (L.)." River Systems 16, no. 4 (2006): 649–65. http://dx.doi.org/10.1127/lr/16/2006/649.

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41

Nykanen, M., and A. Huusko. "Size-related changes in habitat selection by larval grayling (Thymallus thymallus L.)." Ecology of Freshwater Fish 12, no. 2 (2003): 127–33. http://dx.doi.org/10.1034/j.1600-0633.2003.00013.x.

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42

Bănăduc, Doru, Răzvan Voicu, Lee Jason Baumgartner, Saša Marić, Alexandru Dobre, and Angela Curtean-Bănăduc. "Technical Solutions to Mitigate Shifting Fish Fauna Zones Impacted by Long Term Habitat Degradation in the Bistra Mărui River – Study Case." Transylvanian Review of Systematical and Ecological Research 20, no. 3 (2018): 75–114. http://dx.doi.org/10.2478/trser-2018-0021.

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Abstract The Bistra Mărului River fish fauna has been severely impacted by man-made activities, especially through longitudinal fragmentation, over the past 40 years. Fish fauna monitoring revealed structural changes and technical methods have been proposed, in order to restore the natural connectivity and the conservation of fish species. Benefits should accrue for key species: Salmo trutta fario, Cottus gobio, Thymallus thymallus, Eudontomyzon danfordi, Eudontomyzon vladykovi, Gobio uranoscopus, Barbus meridionalis, and Condrostoma nasus.
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43

Scott, Alasdair. "Distribution, Growth, and Feeding of Postemergent Grayling Thymallus thymallus in an English River." Transactions of the American Fisheries Society 114, no. 4 (1985): 525–31. http://dx.doi.org/10.1577/1548-8659(1985)114<525:dgafop>2.0.co;2.

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44

Пономарев, В. И., та А. Б. Захаров. "Распространение и биологические особенности хариуса Thymallus thymallus (Thymallidae) на Европейском Северо-Востоке России". Вопросы ихтиологии 61, № 2 (2021): 153–66. http://dx.doi.org/10.31857/s0042875221010136.

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45

Formicki, Krzysztof, Mieczysław Kowalewski, Andrzej Sobociński, and Aleksander Winnicki. "Sperm quality of grayling (Thymallus thymallus L.) under natural conditions and in ponds." Acta Ichthyologica et Piscatoria 23, no. 2 (1993): 139–45. http://dx.doi.org/10.3750/aip1993.23.2.09.

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46

NAGIEC, M., P. CZERKIES, K. GORYCZKO, A. WITKOWSKI, and E. MURAWSKA. "Mass-marking of grayling,Thymallus thymallus(L.), larvae by fluorochrome tagging of otoliths." Fisheries Management and Ecology 2, no. 3 (1995): 185–95. http://dx.doi.org/10.1111/j.1365-2400.1995.tb00111.x.

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47

LUCZYNSKI, M., R. R. ZAPOROWSKI, and J. S. GOLONKA. "Rearing of European grayling, Thymallus thymallus L., larvae using dry and live food." Aquaculture Research 17, no. 4 (1986): 275–80. http://dx.doi.org/10.1111/j.1365-2109.1986.tb00114.x.

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48

Honkanen, J. O., A. Kostamo, and J. V. K. Kukkonen. "Toxicity of a Phytosterol Mixture to Grayling (Thymallus thymallus) during Early Developmental Stages." Archives of Environmental Contamination and Toxicology 48, no. 3 (2005): 391–96. http://dx.doi.org/10.1007/s00244-003-9238-x.

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Kalat, M., B. Mayr, P. Rab, and W. Schleger. "Heterochromatin Studies and Localization of NORs in European Grayling (Thymallus Thymallus, Pisces, Thymallidae)." Caryologia 41, no. 3-4 (1988): 245–49. http://dx.doi.org/10.1080/00087114.1988.10797865.

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Gum, Bernhard, Riho Gross, Oswald Rottmann, Wolf Schröder, and Ralph Kühn. "Microsatellite variation in Bavarian populations of European grayling (Thymallus thymallus): Implications for conservation." Conservation Genetics 4, no. 6 (2003): 659–72. http://dx.doi.org/10.1023/b:coge.0000006106.64243.e6.

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