Dissertations / Theses on the topic 'Tropical mountains'

Numerical simulations using the Penn State University/NCAR MM4 model are performed to examine a stably stratified, zonal easterly flow past large scale three-dimensional mountain ranges in a rotating, initially barotropic, atmosphere. Upstream blocking by the mountain range diverts the flow primarily to the south and around the mountain. Conservation of potential vorticity results in the formation of a horizontal jet at low levels south of the mountain. This jet is barotropically unstable and leads to a continuous production of synoptic scale vorticity maxima which separate from the mountain and propagate downstream. Numerical simulations using topography representative of the Sierra Madre in Mexico imply that this mechanism may be important in providing some of the initial disturbances which grow into tropical cyclones in the eastern North Pacific Ocean. The wave train produced in the simulations corresponds to waves with 3-7 day periods which have been identified observationally in the eastern North Pacific region. The sensitivity of this effect to the stability of the basic state and the upstream wind speed is investigated. Simulations are also performed which show that the Hoggar and Atlas mountains of west-central Africa block the low-level easterlies resulting in a barotropically unstable jet and a train of vorticity maxima which separate from the mountain and propagate downstream. The spacing of these disturbances is roughly 1600 km and they propagate to the east with a period of about 2.5 days. These characteristics correspond to those of observed waves in the Africa/Atlantic region. It will also be shown that the unique topography of north-central Africa results in a mid-tropospheric easterly jet which has a maximum between 0-10°E and 15-20°N. The location and magnitude of this jet correspond to the so-called African easterly jet which is usually attributed to the strong surface temperature gradients over the continent of Africa. The numerical simulations presented in this work suggest that the mechanical effect of the topography may provide a constant source of energy for the maintenance of the African easterly jet.

Two state-of-the-art Earth systemmodels (ESMs) were used in an idealized experiment to explore the role of mountains in shaping Earth's climate system. Similar to previous studies, removing mountains from both ESMs results in the winds becoming more zonal and weaker Indian and Asian monsoon circulations. However, there are also broad changes to the Walker circulation and El Nino-Southern Oscillation (ENSO). Without orography, convection moves across the entire equatorial Indo-Pacific basin on interannual time scales. ENSO has a stronger amplitude, lower frequency, and increased regularity. A wider equatorial wind zone and changes to equatorial wind stress curl result in a colder cold tongue and a steeper equatorial thermocline across the Pacific basin during La Nina years. Anomalies associated with ENSO warm events are larger without mountains and have greater impact on the mean tropical climate than when mountains are present. Without mountains, the centennial-mean PacificWalker circulation weakens in both models by approximately 45%, but the strength of the mean Hadley circulation changes by less than 2%. Changes in the Walker circulation in these experiments can be explained by the large spatial excursions of atmospheric deep convection on interannual time scales. These results suggest that mountains are an important control on the large-scale tropical circulation, impacting ENSO dynamics and the Walker circulation, but have little impact on the strength of the Hadley circulation.

Climate warming induced range shifts are evident globally for a wide range of taxonomic groups. However, whether such responses have taken place in tropical insect species is unclear. I provided the first such evidence that tropical insects have moved uphill over four decades of climate warming. I repeated a historical moth transect on Mount Kinabalu in Borneo in 2007, 42-years after the original survey in 1965. I duplicated the 1965 sampling strategy in 2007, and excluded sites where habitat changes were evident. I estimated that the average elevation of 102 montane moth species, in the family Geometridae, increased by a mean of 67 m. A sub-group of species retreated their upper boundaries, which may be associated with cloud cover changes and the presence of geological / vegetation transitions. Without these constraints, most species expanded their upper boundaries upwards (by an average of 152 m) more than they retreated at their lower boundaries (77 m), indicating that different ecological mechanisms may predominate in leading and trailing edge populations. Declines of local endemic species, ecological barriers constraining uphill movements, disappearing climate types and a decreasing area of land at higher elevations are the major concerns for conservation. Research of biodiversity risk under warming is urgently required in Southeast Asia. I conducted a meta-analysis of range shifts representing ~1700 species worldwide. The average rate of elevational range shift was 12.2 m/decade uphill, and latitudinal shifts averaged 17.6 km/decade poleward. These rates are faster than previously reported. Latitudinal range shifts responded to the rates of regional warming, but these were not the cases for elevational range shifts. Poikilotherms shifted their distributions faster than homeotherms. The original data and meta-analysis presented in this thesis indicate that climate change is having a pervasive impact on the distributions of species, including in the tropics.

Thesis (PhD(For))--Stellenbosch University, 2013.<br>ENGLISH ABSTRACT: Widdringtonia whytei is a tropical endemic, fire-adapted pioneer coniferous tree species within natural fire-fragmented Afromontane forest patches in a confined area on Mulanje Mountain in Malawi. Natural and anthropogenic fires within the surrounding fire-prone landscape, insect attacks, and uncontrolled harvesting of mature trees for timber threaten the survival of W. whytei. This study investigated the effects of population fragmentation on the reproductive biology of W. whytei, through four specific studies: effects of population size, tree size and crown position on viable seed output; seed rain variation among population sizes; effects of temperature and light on viability and germination of W. whytei seeds; and the influence of population size on natural regeneration patterns. The study was conducted during 2008 and 2009 on three sites using three W. whytei population sizes: small (≤10 reproductively mature cone-bearing W. whytei trees), medium (11-20 cone-bearing trees) and large (>20 cone-bearing trees). Data were collected using field and laboratory experiments and a review of secondary information. The effect of fragment size and crown position on viable seed output and seed rain pattern was tested with a nested linear mixed model. The influence of stem diameter (dbh) on viable seed output was tested using a separate data set. The variation in seed rain was explored with a generalised linear model (GLM) with a negative binomial link function. Variations in seed germination was analysed with a Weibull regression model. The influence of fragment size on seedling regeneration was tested with a GLM (binary logistic regression). Viable seed output from W. whytei cones was very low (23%) and was not affected by population size, tree diameter or crown position. W. whytei population size influenced seed rain with large fragments having higher seed rain densities. Number of dispersed seeds was generally very low and poorly dispersed. W. whytei cones are semiserotinous and depends on moderate to severe fires for heavy seedfall and wider dispersal. Such fires did not occur during the study period. Temperature is a critical factor for seed germination. Seeds germinate between 15 and 25°C with the optimum at ~ 20°C either under light or dark conditions. Regeneration density followed the seed rain pattern indicating that seed availability is a critical factor for regeneration. Regular cool fires outside the forest patches and along the edges during the hot dry season, and the thick litter layer and shady conditions inside forest patches, caused most of the seedling mortality. This study highlighted the difficulties W. whytei experiences with low output of viable seed, limited seed dispersal and low seedling regeneration in and around forest patches, i.e. factors important for conservation management of this species. It is recommended that gaps with diameter equal to canopy height and occasional spot fires would promote seed dispersal onto exposed mineral soil for rapid seedling establishment and subsequent population growth. Such natural fire disturbance events may show the true seed rain and seedling establishment patterns in this species.<br>AFRIKAANSE OPSOMMING: Widdringtonia whytei is ‘n tropiese endemiese, vuuraangepaste pionier naaldhoutagtige boomsoort binne die natuurlike vuurgefragmenteerde Afromontane woudkolle binne ‘n beperkte area op Mulanje berg in Malawi. Natuurlike en menslike vure binne die omringende vuurgeneigde landskap, insekaanvalle, en onbeheerde kap van volwasse bome vir hout bedreig die voortbestaan van W. whytei. Hierdie studie het die effek van populasiefragmentasie op die voortplantingsbiologie van W. whytei ondersoek, deur vier spesifieke studies: die effek van populasiegrootte, boomgrootte en kroonposisie op produksie van lewenskragtige saad; saadreënvariasie tussen populasiegroottes; die effek van temperatuur en lig op lewenskragtigheid en ontkieming van W. whytei saad; en die invloed van populasiegrootte op natuurlike verjongingspatrone. Die studie is gedurende 2008 en 2009 uitgevoer op drie groeiplekke met drie W. whytei populasiegroottes: klein (≤10 keëldraende (volwasse) W. whytei bome), medium (11-20 keëldraende bome) en groot (>20 keëldraende bome). Data is versamel in veld- en laboratorium eksperimente asook 'n oorsig van sekondêre inligting. Die invloed van populasie fragmentgrootte en kroonposisie van bome op saadproduksie en die saadreënpatroon is getoets met ‘n geneste liniêre gemengde model. Die invloed van boomstamdeursnee op die produksie van kiemkragtige saad is met ‘n aparte datastel getoets. Die variasie in saadreënpatroon is met 'n algemene liniêre model (GLM) met 'n negatiewe binomiese skakelfunksie getoets. Variasies in ontkiemingspersentasie van sade is met 'n Weibull regressiemodel ontleed. Die invloed van fragmentgrootte op saailingverjonging is met 'n algemene linêre model (binêre logistiese regressie) getoets. Produksie van lewenskragtige saad in W. whytei keëls was baie lag (23%) en was nie deur populasiegrootte, boomstamdeursnee of kroonposisie beïnvloed nie. W. whytei populasiegrootte het saadreën beïnvloed en groot fragmente het groter saadreëndigthede gehad. Die aantal verspreide sade was in die algemeen baie laag en swak versprei. W. whytei keëls is semi-saadhoudend (serotinous) en is afhanklik van matige tot intense vure vir massiewe saadvrystelling en wyer saadverspreiding. Sulke vure het nie gedurende die studieperiode voorgekom nie. Temperatuur is ‘n kritiese factor vir saadontkieming. Sade ontkiem goed tussen 15 en 25°C met die optimum rondom ~ 20°C onder beide lig en donker toestande. Fragmentgrootte het nie die saailingverjonging in W. whytei beïnvloed nie. Verjongingsdigtheid het die saadreënpatroon gevolg, wat daarop dui dat saadbeskikbaarheid ‘n kritiese factor is vir saailingverjonging. Gereelde koel vure buite die woudkolle en langs die woudrand gedurende die droë warm seisoen, en die dik lae blaarval (litter) en skadutoestande binne ‘n woud, het meeste van die saailingmortaliteit veroorsaak. Hierdie studie het die probleme beklemtoon wat W. whytei ondervind met die lae produksie van kiemkragtige saad, saadverspreiding en saailingverjonging binne en rondom die woudkolle, m.a.w. faktore wat belangrik is vir bewaringsbestuur van hierdie boomsoort. Die aanbeveling word gemaak dat kroonopeninge met ‘n deursnee gelyk aan minstens die kroonhoogte, en sporadiese kolbrande binne die woudkolle kan saadverspreiding op blootgestelde minerale grond vir vinnige saailingvestiging en gevolglike populasiegroei bevorder. Sodanige vuurversteuringsgeleenthede mag die werklike patrone in saadreën en saailingvestiging in hierdie species toon.

Literature review and past empirical work suggests that a resource based model of geodiversity may be a useful proxy for biodiversity within tropical mountains and could provide a valuable conservation planning tool. Here, geodiversity is defined as variation in overall resource availability, along with spatial and temporal (seasonal) variation in resource availability. Using freely available pan-tropical datasets at a 1 km resolution, a spatial model of geodiversity that is informed by an understanding of ecological processes was developed and tested before being used to address three research questions: 1. Is there a quantifiable relationship between geodiversity and biodiversity? 2. Do areas included in multiple conservation prioritisation schemes include a higher proportion of geodiversity than would be expected by chance? 3. What are the likely impacts of climate change on the spatial distribution of current geodiversity classes and what might be the implications of this with respect to the suitability of current protected area configuration in tropical mountains to protect geodiversity and thus biodiversity into the future? Some support for a relationship between geodiversity and biodiversity was found across a range of spatial aggregations and a variety of taxa, however a full validation of this relationship was not possible due to a lack of suitable validation data. It was found that areas rated as valuable on a greater number of conservation prioritisation schemes do not conserve a greater proportion of geodiversity or biodiversity than would be expected by coverage alone or from a random selection. The impacts of climate change on geodiversity were evaluated in terms of change from current conditions under three SRES scenarios and based on projected temperature and precipitation as a mean of five GCM. It was found that there will be significant changes to the current state of geodiversity by the period 2040-2060 and 2080-2100, with the most severe changes occurring by the 2040-2060 period. The implications of these findings are considered in detail for protected areas within Colombia and suggestions for climate change stable conservation strategies are made. It is concluded that the model of geodiversity proposed in this thesis has potential to become a useful conservation tool when considering the effectiveness of current protected areas, and changes in geodiversity due to climate change.

Temperature is likely to be one of the most important abiotic factors given how it affects the physiology of the whole organism and as a consequence, it has an essential role in ecology and evolution. However, how the geographical (and temporal) variation of temperature is related to physiology still raises many questions. Several macrophysiological hypotheses have been proposed to explain the variation patterns of thermal physiological parameters across ecological gradients. Among them, Janzen’s (1967) 'seasonality’ hypothesis is probably one of the most relevant as it has awakened a great interest in other areas besides physiology itself, such as biogeography, ecology, evolution and conservation biology. Janzen (1967) proposed that tropical species are specialists to thermally stable environments and therefore would be more limited to dispersing altitudinally (up or down mountain) than temperate species because of evolved physiological barriers. These biologically based dispersal constraints may be act as a selective mechanism promoting isolation in the populations and thus fuelling speciation rates in tropical mountain ranges, considered the most diverse hotspots in the world. The present thesis explores the evolution in the thermal sensitivity of amphibians across a broad tropical elevational range (4000 meters) in the tropical Ecuadorian Andes and propose the likely environmental causes (altitude and microenvironment) driving the extraordinary diversity in physiological parameters across the gradient. This information also provides essential insight for predicting which species or populations are most vulnerable to global warming. Through estimates of thermal sensitivity in larval and adult amphibians we show, through comparative methods, how thermal sensitivity and tolerance limits diverge along the elevational gradient. We demonstrate that environmental variation at the individual scale is important when testing some of the main macrophysiological hypotheses being better predictors of thermal physiological diversity in amphibians. We demonstrate that amphibians’ thermal physiology is strongly influenced by their thermal environment but also some of its variation may be limited by inherent constraints. A further main finding is the empirical demonstration that physiological barriers in tropical mountains are ‘higher’ upwards than downwards, refining Janzen´s paradigm that altitude functions as a barrier through cold tolerance whereas warm evolution occurs contrarily only transversally (horizontally) through habitat selection. Finally, this thesis suggests that lowland tropical amphibians are more vulnerable to an increase of temperature than their upland counterparts, because they are currently experiencing environmental temperatures close to their physiological optima and heat tolerance. Yet, the use of microclimatic information predicts how, in some cases, highland species may be also vulnerable to suffer heat stress and will therefore need to search for thermal shelters to avoid extreme heat events.<br>La temperatura es probablement un dels factors abiòtics més importants, ja que afecta la fisiologia de tot l'organisme i, com a conseqüència, té un paper essencial en ecologia i evolució. No obstant això, com la variació geogràfica (i temporal) de la temperatura està relacionada amb la fisiologia encara planteja moltes preguntes. S'han proposat diverses hipòtesis macrofisiològiques per explicar els patrons de variació dels paràmetres fisiològics tèrmics a través de gradients ecològics. Entre elles, la hipòtesi de "estacionalitat" de Janzen (1967) és probablement una de les més rellevants, ja que ha despertat un gran interès en altres àrees més enllà de la pròpia fisiologia, com la biogeografia, l'ecologia, l'evolució i la biologia de la conservació. Janzen (1967) va proposar que les espècies tropicals són fisiològicament especialistes ja que viuen en ambients de temperatures estables i, per tant, estarien menys capacitades que les espècies temperades a dispersar-se altitudinalment (muntanya amunt o avall), a causa de les barreres fisiològiques a les que estan evolutivament limitades. Aquesta restricció en la seva dispersió es considera un mecanisme selectiu que promou l'aïllament de les poblacions i, per tant, incrementa les taxes d'especiació en les zones muntanyoses tropicals, considerades les zones més biodiverses del món. Aquesta tesi explora l'evolució de la sensibilitat tèrmica dels amfibis al llarg d'un ampli gradient altitudinal tropical (4000 metres) als Andes equatorians tropicals i proposa algunes de les possibles causes ambientals (altitud i microambient) que promouen la extraordinària diversitat en els paràmetres fisiològics. Aquestes dades ens proporcionen, també, una informació essencial per predir quines espècies o poblacions seran més vulnerables a l'escalfament global. Mitjançant estimacions de sensibilitat tèrmica en larves i amfibis adults mostrem, a través de mètodes comparatius, com els límits de sensibilitat tèrmica (CTmax i CTmin) divergeixen al llarg del gradient altitudinal. Demostrem que la variació ambiental a escala individual és molt important quan es comproven algunes de les principals hipòtesis macrofisiològiques, ja que són un millor predictor de la diversitat fisiològica tèrmica en els amfibis. Demostrem que la fisiologia tèrmica dels amfibis està fortament influenciada per la temperatura a la que estan exposats però, també, algunes de les seves variacions poden estar limitades inherentment. Altres resultats principals són la demostració empírica que les barreres fisiològiques en les muntanyes tropicals són "més altes" cap amunt que cap avall, de manera que es redefineix el paradigma de Janzen: les barreres altitudinals funcionen a través de l'adaptació al fred (per tant cap amunt), mentre que l'evolució al calor es produeix de forma horitzontal, a través de la selecció d'hàbitats. Finalment, aquesta tesi suggereix que els amfibis tropicals de zones baixes, en comparació als de zones altes, són més vulnerables al augment de les temperatures, ja que actualment experimenten temperatures ambientals properes al seu òptim fisiològic (Topt) i tolerància a la calor (CTmax). Tanmateix, l'ús de la informació microclimàtica prediu que, en alguns casos, les espècies d’alta muntanya podrien ser també vulnerables a patir estrès tèrmic i, per tant, hauran de buscar refugi tèrmics per tal d’evitar els extrems de calor.

Understanding biological responses to climate change is a primary concern in conservation biology. Of the ecosystems being rapidly impacted by climate change, those in the high-elevation tropics are among the most poorly studied. The tropical Andean biosphere includes record elevations above 5000 meters, where extreme environmental conditions challenge many organisms. In the Cordillera Vilcanota of southern Peru, frogs including Pleurodema marmoratum and Telmatobius marmoratus have expanded their ranges to 5244 – 5400 m into habitats created by glacial recession, making them among the highest recorded amphibians on Earth. To understand how hydrologic alterations from loss of glacial meltwater and climatic fluctuations affect these amphibians, I conduct a 36-month field study of reproductive phenology and develop a method to distinguish glacial meltwater-fed ponds and precipitation-fed ponds utilizing natural variation in stable isotopes of water (18O, 2H, and d-excess). My results suggest that some ponds critical for breeding populations may have lost their connection to glacial runoff. Ongoing deglaciation may transform these ponds from permanent to ephemeral habitats, leading to the extirpation of the fully aquatic species, T. marmoratus. The 2015/2016 El Niño delayed the onset of the 2015 wet season and shortened the P. marmoratum breeding and tadpole development period in ephemeral ponds. I examine regional patterns of amphibian occupancy and prevalence of the deadly amphibian pathogen Batrachochytrium dendrobaditis in unexplored high-elevation zones that were until recent decades covered by permanent ice. Next, I examine adaptive strategies that allow these two frog species to persist in the harsh high-elevation environment. Pleurodema marmoratum withstands the daily freeze-thaw cycle by utilizing a wide thermal tolerance range (from below 0ºC to CTmax > 32ºC) and I report the first evidence of frost tolerance in a tropical frog. My research compares divergent strategies allowing two anuran species to persist through disease and variable, extreme conditions in high-mountain environments, providing a better understanding of responses to and consequences of climate change for some of the world's highest life forms.

As a strategy to conserve tropical rainforests of the East Usambara block of the Eastern Arc Mountains, Tanzania, I developed a set of models that can identify above-average tree species richness areas within the humid forests. I developed the model based on geo-referenced field data and satellite image-based variables from the Amani Nature Reserve, the largest forest sector in the East Usambara. I then verified the model by applying it to the Nilo Forest Reserve. The field data, part of the Tanzanian National Biodiversity Database, were collected by Frontier-Tanzania between 1999 and 2001, through the East Usambara Conservation Area Management Program, Government of Tanzania. The field data used are rapidly collectible by people with varied backgrounds and education. I gathered spectral reflectance values from pixels in the Landsat Enhanced Thematic Mapper (Landsat ETM) image covering the study area that corresponded to the ground sample points. The spectral information from different bands formed the satellite image-based variables in the dataset. The best satellite image logistic regression and discriminant analysis models were based on a single band, raw Landsat ETM mid-infrared band 7 (RB7). In the Amani forest, the RB7-based model resulted in 65.3% overall accuracy in identifying above average tree species locations. When the logistic and discriminant models were applied to Nilo forest sector, the overall accuracy was 62.3%. Of the rapidly collectible field variables, only tree density (number of trees) was selected in the logistic regression and the discriminant analysis models. Logistic and discriminant models using both RB7 and number of trees recorded 76.3% overall accuracy in Amani, and when applied to Nilo, 76.8% accuracy. It is possible to apply and adapt the current set of models to identify above-average tree species richness areas in East Usambara and other forest blocks of the Eastern Arc Mountains. Potentially, managers and researchers can periodically use the model to rapidly assess, monitor, update, and map the tree species rich areas within the forest. The same or similar models could be applied to check their applicability in other humid tropical forest areas.<br>Ph. D.

Thesis (Ph. D.) -- University of Maryland, College Park, 2007.<br>Thesis research directed by: Dept. of Geography. Title from t.p. of PDF. Includes bibliographical references. Published by UMI Dissertation Services, Ann Arbor, Mich. Also available in paper.

The peatlands are ecosystems extremely sensitive to changes in hydrology, and are considered as faithful \"natural archives of ecological memory\". In the Serra do Espinhaço Meridional, Minas Gerais State, Brazil, mountain peatlands has been studied by soil scientists, but until now multi-proxy studies are almost absent. The location of these peatlands is ideal because they are in an area influenced by the activity of the South America Monsoon Systems (SAMS), which controls the amount and distribution of annual rainfall. The aim of this work was to reconstruct the environmental changes occurred throughout the late Pleistocene and Holocene, both at the local and regional scale by using a multi-proxy approach (stratigraphy, physical properties, 14C and OSL datings, pollen and geochemistry). However, determining of the processes involved in the genesis and evolution of peatlands soils was also necessary step. The physico-chemical properties and elemental composition of five peat cores (PdF-I, PdF-II, SJC, PI and SV) from four selected mires (Pau de Fruta, São João da Chapada, Pinheiros and Sempre Viva) seem to have responded to four main processes: relative accumulation of organic and mineral matter, linked to the evolution of the catchment soils (local erosion); deposition of dust from distant/regional sources; preservation of plant remains; and long and short-term peat decomposition. The combination of proxies of PdF-I core defined six main phases of change during the Holocene: (I) 10-7.4 cal kyr BP, wet and cold climate and soil instability in the mire catchment; (II) 7.4-4.2 cal kyr BP, wet and warm with catchment soils stability and enhanced deposition of regional dusts; (III) 4.2-2.2 cal kyr BP, dry and warm and a reactivation of soil erosion in the catchment; (IV) 2.2-1.2 cal kyr BP, dry and punctuated cooling, with enhanced deposition of regional dusts; (V) 1.2 cal kyr-400 cal yr BP, sub-humid climatic and the lowest inputs of local and regional dust and the largest accumulation of peat in the mire; and (VI) <400 cal yr BP, sub-humid conditions but both local and regional erosion largely increased. For the late Pleistocene, a combination of proxies applied to the PI core also defined six main phases: (I) 60-39.2 cal kyr BP, from sub-humid to dry amid colder conditions than today, and high soil instability in the mire catchment; (II) 39.2-27.8 cal kyr BP, dry and warm with cooling events under still high local erosion rates; (III) 27.8-16.4 cal kyr BP, wet and very cold with a decreased in soil erosion in the catchment; (IV) 16.4-6.6 cal kyr BP, very wet and very cold conditions with low intensity of local erosion; (V) 6.6-3.3 cal kyr BP, very dry and warm with increasing rates of local erosion; and (VI) <3.3 cal kyr BP, from dry and warm to sub-humid climate, with local erosion trend similar to the previous period. The climate is seen as the most important driving force of environmental change, but human activities are likely to have been at least partially responsible for the significant changes recorded over the past 400 years. Given the value as environmental archives, mires from Serra do Espinhaço Meridional should be fully protected.<br>As turfeiras são ecossistemas extremamente sensíveis às mudanças da hidrologia, e são por excelência consideradas como \"arquivos naturais da memória ecológica\". Na Serra do Espinhaço Meridional, Minas Gerais, Brasil, as turfeiras de montanha vem sendo estudadas pelos cientistas do solo, mas até então estudos multi-proxy são quase ausentes. A localização destas é ideal pois estão em uma área influenciada pela atividade do Sistema Monçônico da América do Sul (SMAS), que controlam a quantidade e distribuição de precipitação anual. O objetivo deste trabalho foi reconstruir as mudanças ambientais ocorridas através do Holoceno e Pleistoceno Tardio, tanto sob escala local quanto regional, usando uma abordagem multiproxy (estratigrafia, propriedades físicas, datações 14C e LOE, pólen e geoquímica). No entanto, determinação dos processos envolvidos na gênese e evolução dos solos das turfeiras também foi um passo necessário. As propriedades físico-químicas e composição elementar de cinco testemunhos de turfa (PdF-I, PdF-II, SJC, PI e SV) de quatro turfeiras selecionadas (Pau de Fruta, São João da Chapada, Pinheiros e Sempre Viva) parecem ter respondido a quatro processos principais: acumulação relativa de matéria orgânica e material mineral, ligados à evolução dos solos das bacias das turfeiras (erosão local); deposição de poeira de fontes distantes/regionais; preservação de restos de plantas; e decomposição da turfa em longo e curto prazo. A combinação de proxies de PdF-I definiu seis principais fases de mudanças durante o Holoceno: (I) 10-7,4 mil anos cal AP, clima úmido e frio e instabilidade do solo na bacia da turfeira; (II) 7,4-4,2 mil anos cal AP, úmido e quente com solo na bacia estável e aumento de deposição de poeiras regionais; (III) 4,2-2,2 mil anos cal AP, seco e quente e reativação da erosão do solo na bacia; (IV) 2,2-1,2 mil anos cal AP, seco e resfriamentos pontuais, com aumento de poeiras regionais; (V) 1,2 mil anos-400 anos cal AP, sub-úmido e com os mais baixas entradas de poeiras local e regionais e as maiores acumulações de turfa; e (VI) <400 anos cal AP, sub-úmido com forte erosão local e regional. Para o Pleistoceno tardio, uma combinação de proxies aplicada para PI também definiu seis principais fases: (I) 60-39,2 mil anos cal AP, de sub-úmido para seco em meio à temperaturas mais frias que o atual, e alta instabilidade do solo na bacia da turfeira; (II) 39,2-27,8 mil anos cal AP, seco e quente com alguns resfriamentos e ainda sob elevadas taxas de erosão local; (III) 27,8-16,4 mil anos cal AP, úmido e muito frio com redução da erosão do solo na bacia; (IV) 16,4-6,6 mil anos cal AP, muito úmido e muito frio com baixa intensidade de erosão local; (V) 6,6-3,3 mil anos cal AP, muito seco e quente com taxas crescentes de erosão local; e (VI) <3,3 mil anos cal AP, de seco e quente para sub-úmido, com tendência de erosão local semelhante ao período anterior. O clima é visto como o forçante mais importante das mudanças ambientais, mas é provável que atividades humanas tenham sido parcialmente responsáveis pelas mudanças significativas registradas ao longo dos últimos 400 anos. Dado o valor como arquivos ambientais, as turfeiras da Serra do Espinhaço Meridional devem ser completamente protegidas.

Both tectonics and climate profoundly influence orogenisis, but specifics regarding the forcings, interactions and feedbacks are still largely unclear. This study addresses the evolution of the Cordillera Blanca Mountain Range of northern Peru: an elevated, high-relief, 200 km long string of glaciated peaks comprising the spine of the Andes. Extension along the Cordillera Blanca Detachment Fault (CBDF) actively produces relief along the western flank of the range, exceeding several kilometers in many areas. Abundant records of past glaciations span from >440 ka to the present in the form of moraine and bog deposits. Thus, tectonics (active faulting) and climate (glacial erosion) are operating in tandem to produce some of the highest topography in the western hemisphere. Understanding these processes in the Cordillera Blanca will provide an invaluable perspective into tectonic and climatic effects on orogenisis.Through the combined use of cosmogenic 10Be, low temperature thermochronology and digital terrain analysis, I explore the erosional and morphologic history of the range over a variety of spatial and temporal scales. Significant variations in modern range elevation (maximum, mean, modal and minimum), relief (local and within basins) and slope (maximum, mean and minimum) exist along the strike of the range, potentially reflecting the combined effects of variable displacement along the CBDF and varying degrees of glacial erosion. The morphology of the adjacent supradetachment basin varies as well, containing zones with distinct styles of faulting and basin growth, likely defined in part by the segmentation history of the CBDF. By combining Holocene-scale 10Be basin-averaged erosion rates with new thermochronologic data, I expand the known denudation history of the range. These two datasets constrain the exhumation and erosional history of the range-forming Cordillera Blanca Batholith from the late Miocene to Holocene, and expand the thermal history of the range southwards to include the older Carhuish Stock. Two new vertical 10Be exposure age transects allow comparison of fluvial incision rates within the Cordillera Blanca Batholith and the older Coastal Batholith. Incision rates from the site in the Cordillera Blanca record uplift on the order of ~1 mm yr-1, and potentially place a minimum constraint on CBDF slip rates at this location. Incision rates from the Coastal Batholith are twice as fast (~2 mm yr-1) possibly representing large-scale regional uplift. Exhumation rates and erosion rates generally fall between 0.01 and 0.5 mm/yr, suggesting a fairly continuous state of erosion over long-term (10^6 yrs) to recent (<10^3 yrs) time scales. Aside from a seemingly isolated zone of elevated erosion rates, no trends are observed along the strike of the mountain range. As the CBDF is believed to display variable slip-rates along strike, it seems that associated base level lowering is not a first order control on basin-averaged erosion rate.<br>Bien qu'il soit établi que la tectonique et le climat influencent profondément l'orogénèse, plusieurs questions demeurent en ce qui a trait aux forçages, interactions et rétroactions. Cette étude se penche sur l'évolution de la cordillère Blanche, dans le nord du Pérou : une chaîne de sommets glacés à haut relief de 200 km de longueur, épine dorsale des Andes. L'extension le long de la faille de détachement de la cordillère Blanche (FDCB) cause la formation de relief sur le flanc ouest de la chaîne sur plusieurs kilomètres dans certaines zones. D'abondantes traces de glaciations passées, datant de plus de 440 ka jusqu'à la période actuelle, sont présentes sous forme de dépôts de moraines et de tourbières. Donc, la tectonique (mouvements de failles actives) et le climat (érosion glaciaire) opèrent en tandem pour produire l'une des topographies les plus accidentées de l'hémisphère ouest. Une évaluation pointue des procédés en cause dans la cordillère Blanche permettra une meilleure compréhension des effets de la tectonique et du climat sur l'orogénèse. En utilisant une combinaison de datation par l'isotope cosmogénique 10Be, de thermo-chronologie de basse température et d'analyse de modèles numériques de terrain, cette étude explore l'historique d'érosion et morphologique de la chaîne sur plusieurs échelles spatiales et temporelles. De variations significatives d'altitude moderne de la chaîne (maximale, moyenne, modale et minimale), de relief (local et en bassin) et de pente (maximale, moyenne et minimale) existent le long de la chaîne et sont potentiellement dues aux effets combinés de mouvements variables le long de la FDCB et de différents degrés d'érosion glaciaire. La morphologie du bassin de supra-détachement adjacent varie également et contient des zones dotées de styles distincts de faille et de développement de bassin, probablement définis en partie par l'historique de segmentation de la FDCB. En combinant les rythmes d'érosion (âges 10Be, moyennes de bassin à l'échelle holocène) avec de nouvelles données thermo-chronologiques, ces travaux augmentent l'étendue connue de l'historique de dénudation de la chaîne. Ces deux bases de données documentent l'historique d'exhumation et d'érosion du batholithe de la cordillère Blanche depuis la fin du Miocène jusqu'à l'Holocène et étendent l'historique thermal de la chaîne vers le sud pour inclure le plus ancien Carhuish Stock. Deux coupes d'âges d'exposition verticale par 10Be permettent une comparaison des rythmes d'incision fluviale au sein du batholithe de la cordillère Blanche et du plus ancien batholithe côtier. Les rythmes d'incision au site de la cordillère Blanche indiquent un soulèvement d'environ 1 mm par an et suggèrent un glissement minimal de la FDCB de cet ordre à cet endroit. Les rythmes d'incision au batholithe côtier sont deux fois plus rapides (~2 mm par an), possiblement dû à un soulèvement d'échelle régionale. Les rythmes d'exhumation et d'érosion sont généralement de 0.01 à 0.5 mm par an, indiquant une érosion continue sur le long (10^6 années) et court terme (<10^3 années). Outre une zone isolée de rythmes d'érosion apparemment élevés, aucune tendance n'est observée le long de la chaîne de montagnes. Comme la FDCB est connue pour ses rythmes de glissement variables le long de la chaîne, il semble que l'abaissement du niveau de base associé ne soit pas un contrôle de premier ordre sur les rythmes d'érosion moyens à l'échelle du bassin.

The tropical montane forest of Ecuador is one of the ‘hot spots’ of species diversity. Despite this great species diversity, there are forests in our study area, namely in the Zamora-Chinchipe province in southern Ecuador, which are being depleted at an alarming rate. For example, large areas are being permanently deforested for use as ‘pastureland ’. This development is typically characterized by intensive slash and burn activities for vegetation clearance. After clear cut and slash burning, pasture species are planted of which Setaria sphacelata and Melinis minutiflora, are the most common. However, forest clearing by slash burn for pasture production occurs in the region simultaneously with subsequent invasion of bracken fern (Pteridium arachnoideum) and re-colonization of secondary succession vegetation on abandoned pasture land. In this study, we will examine the effects that the above phenomena have on ecosystems, namely the effects associated with the conversion of natural forestland to pasture land, and the succession that occurs as a result. We will also examine how land use change affects the nutrient status of the soil. This study focuses specifically on the area called the ‘San Francisco Valley’, (3°58’ 30”S latitude, 79°4’ 25”W longitude), which lies between Loja and the Zamora-Chinchipe provinces in Ecuador. Field work for this study was carried out at 1,798 and 2,226 m a.s.l in the following sites: 1) the San Francisco Natural Forest (this area refers to the eastern part of Podocarpus National Park); 2) active pastures Type I (this area refers to the pastureland in front of the San Francisco Scientific Station ECSF, and are pastures that have existed for 50 years or more); 3) active pastures Type II (this area refers to the pastureland in Sabanilla sites, and are pastures that have existed for approximately 17 years); and 4) abandoned pastureland (affected by succession over a period of at least twenty years). Each land-use type consists of five plots of approximately 20 x 20m2. Five points in each plot were then chosen and later sampled (two sub-samples) according to the defined horizons and depth units, namely the organic layer and mineral top soil horizons. Organic layer (only on forest and succession sampling plots) were designated as LOf1, Of2/Oh1 and mineral top soil was sampled to a depth of a 0-10, 10-20, 20-30 cm. To quantify the effects of the impact that land use change has on the chemical characteristics of soil in the selected areas, we analyzed the following bio-elements: the pH value, soil organic carbon (SOC), the total nitrogen (TN), the effective cation exchange capacity (ECEC), as well as stocks of total and available macronutrients. In addition, we examined the biological characteristics such as carbon and nitrogen microbial biomass (MBC, MBN), basal respiration (BR), and nitrogen mineralization (Nmin) in organic layers and top mineral soils (0-30 cm) of the following: the San Francisco Natural Forest, ECSF pastureland, Sabanilla pastureland, and abandoned pastures affected by succession. The main results of this study can be summarised as follows: The soil pH value increased after forest-to-pasture conversion and tended to decrease with soil depth. Moreover, pastureland in the study that was fifty years or more, showed evidence of a decrease in pH values. This decrease in pH value of the soil can be attributed to the reduction in exchangeable cations . However, they still remained higher than the pH values for the forest sites in the study. After the abandonment of pastureland, we observed a re-acidification in the soil of succession sites. This resulted in a decrease of base saturation . On the other hand, the total exchangeable base cation stocks were significantly higher for pasture soils compared with forest and succession soils. This was because of the addition of basic cations by ‘slash and burn activity ’. In this study, we observed that the greater values of soil organic carbon stocks occurred in forest sites. This can be explained by their high SOC value in the organic layer (73.9 Mg ha-1). However, the mineral layer (0 to 30 cm depth) of forest had a lower value than pasture and succession sites. This higher SOC stock of the mineral layer of our pasture sites are partly a result of carbon input from the former standing biomass. Likewise, in the mineral layers, we found that MBC content increases in a similar way to the SOC content. For example, when we compared the MBC content of forests with the pasture sites, we observed that pastures had approximately three times as much MBC. The results of nitrogen stored in the Microbial Biomass (MBN) showed the same tendency as the MBC values. Moreover, for organic layers, the total nitrogen stocks of forest were higher compared to succession sites. On the other hand, for mineral soil (0 to 30 cm depth), nitrogen stocks increased after forest to pasture conversion. This increase is partly due to the burning of aboveground biomass and the subsequent death of roots. Furthermore, the values of nitrogen stocks decreased again in succession sites (4.2 Mg N. ha-1), with similar values those of forests (4.4 Mg N. ha-1 ). Our results show that the slash-and-burn practice leads to a significant increase of P stock. We found that stocks of total phosphorus were significantly higher in the mineral topsoil (0–30 cm) of 50 year-old pastures (ECSF) than in the 17 year-old pastures (Sabanilla). It is important to note that the P stocks in the mineral soil of the abandoned pasture (20 year-old pasture sites) tend to return to forest values (399.9. Kg.ha-1). Nevertheless, the results show very low values of available P on both pastures and succession sites compared with forest sites. In the organic layers, our results show significant differences in the values of basal respiration between forest and succession sites. This indicates that the level of CO2 was greater in the selected forestland due to an increase of organic material. This essentially means that there was an increase of micro-organisms in the soil and subsequently an improved nutrient cycle . For the mineral soil, however, the results only showed a significant difference of 0-10 cm depth in the ECSF pastures and forests. In our study, we did not find any significant differences in the net nitrogen mineralization values in the four studied areas. Nevertheless, the results show that net nitrogen mineralization values decrease systematically according to the depth of the land uses. Summarizing, after the conversion of forest to pasture, there was an increase of the value of bio-elements in the mineral layers at both pasture sites. However, this increase was higher in 50 year old pastures (ECSF) than in the 17 year old pastures (Sabanilla). In addition, we noticed that after 20 years of the abandonment of pastures, most measured soil properties returned to the old-growth forest levels<br>El bosque montano tropical de Ecuador es uno de los \"puntos calientes\" de diversidad de especies. Pero el bosque primario en el área de investigación (la provincia de Zamora Chinchipe al Sur de Ecuador) se está perdiendo a un ritmo alarmante. Grandes áreas están siendo permanentemente deforestadas para su uso como tierras de pastoreo. Este proceso se caracteriza por el uso intensivo de fuego para desmonte de la vegetación. Después de la quema y roza, se plantan especies de pastos entre las que Setaria sphacelata y Melinis minutiflora son las más comunes. Sin embargo, la quema y tala del bosque para la conversión a zonas de pastos ocurre simultáneamente con la subsecuente invasión del helecho común (Pteridium arachnoideum), produciéndose. La difusión de esta maleza y la decreciente productividad de los pastos (especialmente Setaria spacelata y Melinis minutiflora) conducen a que las tierras se utilizan en pastoreo hasta que la tierra se agota de nutrientes y luego se abandonan. En este estudio, se examinan los efectos que los fenómenos anteriormente mencionados tienen sobre los ecosistemas; a saber, los efectos asociados a la conversión de bosques naturales en tierras de pastoreo y la posterior sucesión de los pastizales; además de cómo el cambio de uso del suelo afecta el estado nutricional de los suelos del sur del Ecuador. El estudio se sitúa en la zona del valle de San Francisco (3°58’ 30”S latitud, 79°4’ 25”W longitud), entre Loja y Zamora Chinchipe, provincias que se encuentran en el sur de Ecuador. El trabajo de campo se llevó a cabo a una altura entre 1,798 y 2,226 m s.n.m.; en: 1) las áreas de bosque natural San Francisco (esta área se refiere a la parte oriental del Parque Nacional Podocarpus); 2) pasto activos Tipo I (esta área se refiere a los pastos en frente de la Estación Científica San Francisco ECSF, y son pastizales que tienen más de 50 años de edad); 3) pastos activos Tipo II (esta área se refiere a los pastos en los sitios de Sabanilla, y son pastizales que existen desde hace aproximadamente 17 años); y 4) pastos abandonados bajo vegetación de sucesión (más de 20 años de edad). Cada tipo de uso de la tierra consistió en cinco parcelas de aproximadamente 20 x 20 m2; se eligieron cinco puntos en cada parcela y se tomaron muestras (dos sub-muestras) de acuerdo con los horizontes orgánico y mineral. Las capas Orgánicas (solo presentes en el bosque y en los sitios de sucesión) se identificaron como LOf1, Of2/Oh1 y las capas minerales se muestrearon hasta los 0-30 cm de profundidad del suelo. Para cuantificar los efectos del impacto del cambio del uso de la tierra en las características químicas del suelo, se analizó los siguientes bio-elementos: pH, carbono orgánico del suelo (COS), nitrógeno total (Nt), la capacidad de intercambio catiónico efectiva (CICE), así como las reservas del contenido total y disponibilidad de los macro nutrientes. Además, las características biológicas, tales como el carbón (MBC) y nitrógeno (MBN) de la biomasa microbiana, la respiración basal (Rb) y la mineralización de nitrógeno (Nmin) en las capas orgánicas y en la capa mineral del suelo (hasta -30 cm) de los bosques naturales, pastos y pastizales abandonados bajo vegetación de sucesión. Los principales resultados del estudio se describen a continuación: El valor de pH del suelo indica una elevada acidez, después de la conversión de bosque a pastos y tiene una tendencia general a disminuir con la profundidad del suelo, cuando los pastos se hacen más viejos (más de 50 años de edad); los valores de pH del suelo disminuye como consecuencia de la lixiviación de cationes intercambiables, pero siguen siendo superiores a los valores de pH de los sitios del bosque. Tras el abandono de los pastos se observó una re-acidificación en el suelo de los sitios de sucesión, lo que resulta en una disminución de la saturación de bases. Los resultados además indican que las capas orgánicas, especialmente en los suelos de los bosques, almacenan una cantidad importante de potasio, calcio y magnesio. Sin embargo, las reservas totales de cationes básicos intercambiables fueron significativamente mayores en los suelos de los pastizales que en los bosques y que los suelos de sucesión, debido a la adición de cationes básicos producidos por la tala y quema usada en la conversión de bosques a pastizales. En este estudio, se encontró que los mayores valores de reservas de COS se producen en zonas forestales, que se corresponde con un alto valor en la capa orgánica (73,9 Mg C ha-1). Sin embargo, la capa mineral (hasta - 30 cm de profundidad) de los suelos del bosque tiene un valor menor en comparación con los valores de los pastos y sucesión. Este alto contenido de COS en los pastos, se debe en parte al ingreso de carbono desde la biomasa. Asimismo, en las capas minerales, se encontró que el contenido de carbono en biomasa microbiana (CBM) aumenta de una manera similar al contenido de carbono orgánico del suelo (COS). Por ejemplo, cuando comparamos el contenido de CBM de los bosques con los sitios de pastoreo, se observó que los pastos tenían aproximadamente tres veces más MBC. El resultado de nitrógeno almacenado en la biomasa microbiana (NBM) mostró la misma tendencia que los valores de CBM. En las capas orgánicas, las reservas totales de nitrógeno de los bosques fueron mayores en comparación con los sitios de sucesión. Por otro lado, en el suelo mineral (hasta -30 cm de profundidad) las reservas de nitrógeno aumentan después de la conversión de bosques a pastizales. Este incremento es parte debido a la quema de la biomasa superior y de la subsecuente muerte de las raíces. Además, los valores de las reservas de nitrógeno disminuyen de nuevo en los sitios de sucesión (4.2 Mg N. ha-1), a valores similares a las del bosque (4.4 Mg N. ha-1 ). Nuestros resultados muestran que las prácticas de quema y tala incrementan significativamente las reservas de fosforo, encontramos que las reservas de fosforo total fueron significativamente altas en las capas minerales (- 30 cm) de los pastos de 50 años (ECSF), seguido de los pastos de 17 años de edad (Sabanilla). Es importante notar que las reservas de fosforo en las capas minerales de los suelos de pastos abandonados (20 años de edad) tienden a retornar a los valores del bosque (399.9. Kg.ha-1). Sin embargo, los resultados muestran valores muy bajos de fosforo disponible en ambos sitios de pastos y sucesión comparado con el bosque. En las capas orgánicas, los resultados revelan diferencias significativas en los valores de la respiración basal (potencial) entre el bosque y la sucesión. Esto indica que el nivel de CO2 fue mayor en el los sitios de bosque debido a un aumento de la materia orgánica. Esto implica un aumento de los microorganismos en el suelo y, posteriormente, una mejora en el ciclo de nutrientes. En el suelo mineral, los resultados sólo muestran una diferencia significativa en la profundidad de 0-10 cm entre los pastos ECSF con los bosques. En nuestro estudio, no se encontró diferencias significativas en los valores de mineralización neta de nitrógeno entre las cuatro áreas estudiadas. Sin embargo, los resultados muestran que los valores netos de mineralización del nitrógeno disminuyen sistemáticamente con la profundidad en todos los usos de los suelos estudiados. En resumen, después de la conversión de bosques en pastizales, se produjo un incremento del valor de los bio-elementos en las capas minerales de ambos sitios de pastoreo. Sin embargo, este aumento fue mayor en los pastos de 50 años de edad (ECSF) que en los pastos de 17 años de edad (Sabanilla). Además, los resultados muestras que después de 20 años del abandono de los pastos, la mayoría de los parámetros medidos retornan a valores similares a los de los sitios de bosque<br>Im globalen Kontext stellt der tropische Bergregenwald in Süd-Ecuador ein „Hotspot“ der Biodiversität dar. Im Untersuchungsgebiet ist die enorme Artenvielfalt durch die massive Zerstörung der natürlichen Waldökosysteme gefährdet. Der Wald wird durch intensive Brandrodung großflächig in Weideland umgewandelt. Im Laufe der Weidenutzung kommt es auf den Weideflächen zu einer zunehmenden Ausbreitung und Dominanz des tropischen Adlerfarns (Pteridium arachnoideum). Die Farnausbreitung und die abnehmende Produktivität der Weidegräser insbesondere von Setaria sphacelata und Melinis minutiflora, führt zum Verlassen der Weiden und zur Etablierung neuer Weideflächen durch fortgesetzte Brandrodung des Naturwaldes. Aufgelassene ehemalige Weideflächen unterliegen einer sekundären Sukzession. Innerhalb dieser Studie wurden die Effekte der Umwandlung des Naturwaldes in Weideland, der Weidenutzung sowie der Wirkungen der sekundären Sukzession nach dem Verlassen unproduktiver Weideflächen auf die Nährstoffsituation der Böden untersucht. Die Untersuchung erfolgte im Gebiet des „San Francisco Tales“, (3°58’ 30”S, 79°4’ 25”W), welches sich zwischen den beiden Provinzen Loja und Zamora-Chinchipe in Ecuador befindet. Feldarbeiten innerhalb dieser Studie wurden in einer Höhe zwischen 1798 und 2225 m NN in den folgenden Bereichen ausgeführt: 1) Naturwald in San Francisco; 2) aktive Weidefläche Typ I (seit circa 50 Jahren in Nutzung); 3) aktive Weidefläche Typ II (seit circa 17 Jahren in Nutzung); und 4) verlassenes Weideland (bestimmt durch sekundäre Sukzession seit mindestens 20 Jahren). Jeder Landnutzungstyp besteht aus fünf Plots mit einer Ausdehnung von circa 20 x 20 m. Fünf Punkte innerhalb eines jeden Plots wurden zur Beprobung ausgewählt. Die organische Auflage (Naturwald, verlassenes Weideland) wurde nach Auflagehorizonten (LOf1 und Of2/Oh) und der Mineralboden nach Tiefenstufen (0-10 cm, 10-20 und 20-30 cm) getrennt beprobt. Um die Effekte und den Einfluss des Landnutzungswandels auf chemische Bodenkennwerte zu quantifizieren, wurden die folgenden Indikatoren untersucht: pH, organischer Kohlenstoffgehalt des Bodens (SOC), Gesamtstickstoff (TN), effektive Kationenaustauschkapazität (CECeff) sowie Vorräte der gesamten und pflanzenverfügbaren Makronährelemente. Zusätzlich wurden bodenbiologische Indikatoren wie Kohlenstoff und Stickstoff der mikrobiellen Biomasse (MBC, MBN), Basalatmung (BR) und Stickstoffmineralisation (Nmin) in den organischen Auflagehorizonten und dem mineralischen Oberboden (0-30 cm) wie folgt untersucht: Naturwald in San Francisco, ECSF Weideflächen, Sabanilla Weideflächen und aufgelassene, von der Sukzession beeinflusste Weiden. Die hauptsächlichen Resultate der Untersuchung können wie folgt zusammengefasst werden: Der pH-Wert des Bodens erhöhte sich nach der Umwandlung von Wald zu Weide und zeigt mit zunehmender Tiefe einen abnehmenden Trend. Darüber hinaus nahmen die pH-Werte von der 17 Jahre alten zur 50 Jahre alten Weide ab. Diese Abnahme kann auf den Rückgang austauschbarer Kationen zurückgeführt werden. Dennoch verblieb der pH-Wert in den 50 Jahre alten Weiden oberhalb des pH-Wertes der im Wald beprobten Flächen. Nach dem Auflassen der Weiden wurde eine erneute Versauerung des Bodens im Stadium der Sukzession beobachtet. Dieser pH-Rückgang wird durch einen Abnahme der Basensättigung begleitet. Auf der anderen Seite waren die Vorräte an austauschbaren Basen der CECeff auf den Weideflächen signifikant über denen der Wald- und Sukzessionsflächen. Ursächlich dafür war die Freisetzung basischer Kationen während der Brandrodung des Naturwaldes. Innerhalb dieser Studie wurden die höchsten Vorräte an Bodenkohlenstoff im Naturwald ermittelt. Das kann durch hohe Vorräte an SOC in der organischen Auflage (73.9 Mg ha-1) erklärt werden. Allerdings waren die Vorräte im Mineralboden (0-30 cm) des Waldes geringer als auf den Weide- und Sukzessionsflächen. Diese höheren SOC-Vorräte im Mineralboden der Weideflächen sind teilweise das Resultat der höheren Kohlenstoffzufuhr durch Reste der oberirdischen Biomasse. Der Gehalt an mikrobiellem Biomassekohlenstoff nahm in ähnlicher Art und Weise wie der des bodenbürtigen organischen Kohlenstoffs zu. Beispielsweise wurde beim Vergleich von MBC und MBN-Gehalten des Waldes mit denen der Weide der dreifache Gehalt in der Weide gefunden. Außerdem waren die Stickstoffvorräte der organischen Auflagen im Wald höher als auf den Sukzessionsflächen. Andererseits nahmen die Stickstoffvorräte nach der Umwandlung von Wald zu Weide im Mineralboden (0-30 cm) zu. Diese Zunahme beruht zum Teil auf dem Abbrennen der oberirdischen Biomasse und dem anschließenden Absterben der Wurzeln. Des Weiteren verringerten sich die Stickstoffvorräte im Stadium der Sukzession (4.2 Mg N ha-1) und glichen sich wieder denen des Waldes (4.4 Mg N ha-1) an. Die Resultate zeigen, dass die Brandrodung zu einem signifikanten Anstieg der Phosphorvorräte führte. Vorräte des Gesamtphosphors waren in den mineralischen Oberböden (0-30 cm) der 50 Jahre alten Weide, gefolgt von der 17 Jahre alten Weide signifikant am höchsten. Zu betonen ist, dass die P-Vorräte im Mineralboden der aufgelassenen Weide dazu tendieren auf Werte des Waldes abzusinken (399.9 kg ha-1). Trotzdem zeigten die Ergebnisse sehr geringe Werte für pflanzenverfügbaren Phosphor in Weide und Sukzession im Vergleich zum Wald. In den organischen Auflagehorizonten wurden signifikante Unterschiede in der mikrobiellen Aktivität (Basalatmung) zwischen Wald und Weide gefunden. Das zeigt, dass das Niveau an C-Mineralisierung in der ausgewählten Waldfläche größer ist, was auf die erhöhte Verfügbarkeit der organischen Substanz zurückgeführt werden kann. Das bedeutet im Wesentlichen einen Anstieg der Mikroorganismen in den Böden und darauffolgend einen verbesserten Nährstoffkreislauf. Für den Mineralboden zeigten die Ergebnisse jedoch nur einen signifikanten Unterschied in einer Tiefe von 0-10 cm zwischen Wald und Weide. Signifikante Unterschiede in der Nettostickstoffmineralisation der vier untersuchten Gebiete konnten nicht gefunden werden. Nichtsdestotrotz zeigten die Ergebnisse, dass die Nettostickstoffmineralisation mit zunehmender Tiefe in den unterschiedlichen Landnutzungen abnimmt. Zusammenfassend kann festgestellt werden, dass nach der Brandrodung die Gehalte an Bioelementen im Mineralboden beider Weideflächen zunahmen. Dieser Anstieg war in den 50 Jahre alten Weideflächen (ECSF) sogar ausgeprägter als in den 17 Jahre alten Weideflächen (Sabanilla). Zusätzlich zeigte sich in den 20 Jahre alten Sukzessionsflächen, dass durch die Regeneration einer organischen Auflage ein positiver Effekt hinsichtlich erhöhter Vorräte an SOC und pflanzenverfügbarem Phosphor festgestellt werden konnte. Diese allmähliche Zunahme an Nährstoffen im Boden der Sukzession setzte sich fort, bis das Ausgangsniveau des Waldes wieder erreicht war

Tropical mountain forests contribute greatly to carbon sequestration and reduction of carbon levels in the atmosphere. In order to achieve the “Reducing Emission from forest deforestation and degradation” (REDD +) objectives, reliable information on carbon sequestration of all forest management areas is required. So, this study aims to determine the amount of above ground carbon bound in trees and how carbon sequestration capacity of trees varies with altitudinal change in the Magamba Nature Forest Reserve. A random stratified coordinate by elevation method was used, measuring approximately 2636 tree of 5&lt;diameter at breast height (DBH) distributed across 29 plots with a total area of 2.03ha. The effect of altitude was tested using ANOVA and Pearson correlation test. The results show that there is a significant (p&lt;0.05) positive correlation between aboveground carbon sequestered and elevation, and that there was significant difference (p&lt;0.02) in the amount of aboveground carbon stored between low elevation (147.5 tones ha-1) mid elevation (244.86 tonnes ha-1) and high elevation (245.4 tones ha-1). The lower amount of carbon at low elevation may be due to anthropogenic activities including logging and fuel production, since it is easier to reach lower altitude than higher altitude forest zone. To improve carbon sequestration capacity of tropical Mountain protected forests, participatory protective measure at lower elevation zones are of importance. These should even be included in management plans, detailing the involvement of adjacent community.

Quantificar a biomassa florestal tropical e caracterizar a estrutura da floresta em escalas finas é fundamental para compreender melhor o papel dos ecossistemas tropicais no ciclo global de carbono. O sensoriamento remoto por LiDAR é uma ferramenta poderosa para avaliar a estrutura da vegetação em 3D e estimar a biomassa acima do solo, considerando que as medições do LiDAR penetram na vegetação densa da floresta gerando estimativas precisas da topografia e das alturas das árvores. Florestas tropicais com dossel denso apresentam vários desafios para o sensoriamento remoto por LiDAR, especialmente em áreas de topografia acidentada, onde grande parte das remanescentes da Mata Atlântica se encontra. Dados de LiDAR aerotransportado foram adquiridos sobre o Parque Estadual da Serra do Mar - SP, Brasil, em uma área montanhosa com declives íngremes cobertas por floresta tropical densa. Modelos digitais de terreno derivados do LiDAR foram comparados com 35 pontos de controle medidos com receptores GNSS de alta precisão, pós-processados com correção diferencial. Os dois modelos de terreno obtidos do LiDAR mostraram-se extremamente precisos, com erros médios de 0,19 m ($\pm 0,97$ m) e 0,18 m ($\pm 0,95$ m) em relação aos pontos de controle. Redução aleatória da densidade original de pontos de LiDAR (20 pontos/$m^{2}$) resultou em diminuição da precisão dos modelos de terreno: os erros médios subiram para 0,38 m ($\pm 1,32$ m), 1,12 m ($\pm 2,04$ m), 1,59 m ($\pm 3,13$ m) e 3,21 m ($\pm 3,12$ m) com as densidades de 8, 4, 2 e 1 pontos/$m^{2}$, respectivamente. As diferenças de elevação entre o modelo de terreno e os pontos de controle em áreas submontanas foram consistentemente mais altas do que as diferenças em áreas montanas, possivelmente refletindo a complexidade variável do terreno e os efeitos da altura do vôo. A altura do dosse1 calculada a partir dos dados de LiDAR com densidade reduzida diferiu também significativamente da altura do dossel estimada a partir dos dados com densidade original. A altura média da superfície do dossel diminuiu de 3\%, 8\%, 16\% e 25\% com densidades de 8,4,2 e 1 pontos/$m^{2}$,, respectivamente. A magnitude da mudança na altura do dos sel foi maior nas parcelas submontanas (entre 0,79-6,08 m) do que nas montanas (entre 0,60-4,24 m) dentro de todas as classes de densidade de dados. A baixa variação das elevações do modelo da superfície da vegetação com dados reduzidos indicou que a diminuição da altura do dosse1 foi devido a dificuldade de caracterizar a topografia e não a incapacidade de capturar as alturas máximas das árvores. Métricas de estrutura do dossel e características do terreno derivadas dos dados de LiDAR (densidade original) foram bons preditores de componentes da biomassa acima do solo em parcelas permanentes de 1 hectare dentro da área de estudo. A fração de aberturas no dossel foi fortemente relacionada a perda de biomassa acima do solo ($R^{2}$ = 0,87) e a variação líquida ($R^{2}$ = 0,91), enquanto que a altura média da superfície do dos seI explicou 43\% da variação na biomassa total acima do solo e 68\% da variação na área basal total. A fração de aberturas no dossel e as taxas de mortalidade foram menores em terrenos íngremes do que em encostas suaves, sugerindo que encostas mais íngremes podem proporcionar condições mais favoráveis (de nutrientes, de água e disponibilidade de luz) para o crescimento das árvores e acúmulo de biomassa. Dada a crescente ênfase sobre o uso de LiDAR aerotransportado para manejo florestal e esforços de conservação (REDD+), os resultados deste estudo destacam a importância do planejamento cuidadoso de levantamentos futuros com amostragem consistente para a quantificação precisa dos estoques e dinâmica da biomassa acima do solo. Em terreno montanhoso coberto com densa vegetacão tropical, como a Mata Atlântica da Serra do Mar, dados de LiDAR de baixa densidade vão subestimar a biomassa e podem não caracterizar verdadeiramente a heterogeneidade espacial da estrutura da floresta.<br>Quantification of tropical forest biomass and characterization of forest structure at fine scales is critical for a better understanding of the role of tropical ecosystems in the global carbon cycle. LiDAR remote sensing is a powerful tool for assessing 3D vegetation structure and estimating aboveground forest biomass, provided that LiDAR measurements penetrate dense forest vegetation to generate accurate estimates of surface topography and canopy heights. Dense tropical forest canopies present various challenges for LiDAR remo te sensing, especially in areas of steep topography where much of the remaining Atlantic Forest is concentrated. Airbome LiDAR data were acquired from a commercial provider for a region of the Serra do Mar State Park in the state of São Paulo, Brazil, a mountainous area with steep slopes covered by mature tropical dense forest. Digital terrain models (DTMs) derived from all LiDAR data were compared to 35 ground control points measured with survey grade GNSS receivers, post-processed with differential correction. The two LiDAR-based terrain mo dels were extremely accurate, with mean signed errors of 0.19 m ($\pm 0.97$ m) and 0.18 m ($\pm$ 0.95 m) compared to ground points. Random thinning of the original LiDAR point density (20 points/$m^{2}$) decreased the accuracy of the terrain mode1s, with signed errors rising to 0.38 m ($\pm$ 1.32 m), 1.12 m ($\pm$ 2.04 m), 1.59 m ($\pm$ 3.13 m) and 3.21 m ($\pm$ 3.12 m) as point density was reduced to 8, 4, 2 and 1 points/m-, respective1y. Offsets between LiDAR DTMs and ground data in submontane areas were consistently higher than those in montane areas, possibly reflecting the varying complexity of the terrain and the effects of variable ranging distance. Canopy heights ca1culated from the thinned LiDAR data also differed significantly from canopy heights estimated with the full LiDAR density. Mean canopy surface height decreased by 3\%, 8\%, 16\% and 25\% as retum density was reduced to 8, 4, 2 and 1 points/m-, respectively. The magnitude of change in canopy height was greater in submontane plots (range of 0.79-6.08 m) than in montane plots (range of 0.60-4.24 m) within all data density classes. Low variation in vegetation surface model elevations with reduced data density indicated that the decrease in canopy heights was due to the difficulty to characterize topography precisely and not the inability to capture the top heights of trees. Metrics of canopy structure and terrain characteristics derived from the full-density LiDAR data were significantly related to aboveground biomass components in 1-ha permanent plots at the study site. Canopy gap fraction showed close correspondence with aboveground biomass loss ($R^{2}$ = 0.87) and net change ($R^{2}$ = 0.91), while mean canopy surface height explained 43\% of the variation in total aboveground biomass and 68\% of the variation in total basal area. Both canopy gap fraction and rates of mortality were lower on steep terrain than on gentler slopes, suggesting that steeper slopes might provi de more favorable conditions (nutrient, water and light availability) for tree growth and biomass accumulation than flat terrain. Given the growing emphasis on the use of airbome LiDAR for forest management and conservation efforts (REDD+), the results of this study highlight the importance of careful survey planning and consistent sampling frames for accurate quantification of aboveground biomass stocks and dynamics. In mountainous terrain under closed-canopy tropical forest, such as the Atlantic Forest of the Serra do Mar, low-density LiDAR coverage will underestimate biomass and might not characterize truthfully the spatial heterogeneity of forest structure.

Latin America has pioneered the concept of Payment for Ecosystem Services (PES) as a strategy to improve the management of ecosystem services. Ecuador is not an exception, where many PES schemes have been implemented to protect the tropical mountain ecosystem “páramo” and the water resources these areas are generating for downstream societies. A successful PES scheme needs to achieve both targeted bio-physical objectives and at the same time benefit local conditions while not risking to sacrifice the local demand for ecosystem services. This balance is explored here in a case study focusing on the Río Grande watershed in the highlands in the northern parts of Ecuador by exemplifying community participation in the public PES scheme Socio Bosque (PSB) starting in 2009. The water resource distribution (precipitation, discharge, actual evapotranspiration and potential evapotranspiration) in the watershed was evaluated over the last decades. The local perception of the PSB and its impacts on local and regional water resources were also studied and characterized. The results showed that the annual discharge in the Río Grande watershed has decreased significantly from 1967-2014 and that the annual discharge was significantly lower between 1997-2015 compared to 1979-1997. Since precipitation did not decrease significantly during this period, the changes of the annual discharge are more likely depended on factors controlling the seasonal distribution of discharge and evapotranspiration in the watershed. For example, large scale land use changes coupled with a significantly warmer climate in the region could be a possible driver. Of course, this would not exclude other important factors such as changes in water demand and the supply of freshwater from the Río Grande watershed to downstream societies. The results of this case study showed that it is likely too early to see any impacts in the water balance components as a direct response to the implemented PSB scheme. Clearly, this motivates a need for continued evaluation of the local perception and the water resources to ensure that the need and demand for ecosystem services in a long-term perspective are maintained.<br>Latinoamérica ha sido pionera en el concepto de Pago por Servicios Ecosistémicos (PSE o PES en lassiglas en inglés) como estrategia para mejorar la gestión de servicios ecosistémicos. En Ecuador, se han implementado muchos PSE para proteger el ecosistema montañoso tropical de El Páramo así como los recursos acuíferos que dichas áreas generan para las sociedades que habitan cuenca abajo. Un esquema de PSE exitoso requiere alcanzar los objetivos biofísicos y respetar las necesidades locales de servicios ambientales. Este equilibrio se ha analizado tomando como objeto de estudio la cuenca hidrográfica del Río Grande en las tierras altas del norte de Ecuador y la participación comunitaria en el programa de PSE denominado Socio Bosque (PSB) iniciado en 2009. Se estudiaron la distribución del agua (precipitación, descarga del agua, evapotranspiración actual y evapotranspiración potencial) en la cuenca hidrográfica durante las últimas décadas. También se estudiaron los impactos locales y regionales del PSB en los recursos hídricos y los percepción local con respecto a la implementación de PSB. Los resultados muestran que la descarga anual de la cuenca hidrográfica ha decrecido significativamente durante el período comprendido entre 1967 y 2014, particularmente, señalan un decrecimiento considerablemente mayor entre 1997 y 2015 con respecto al período 1979-1997. La precipitación no se redujo durante el período estudiado y, en consecuencia, los cambios en la descarga anual dependen presumiblemente de factores que controlan la distribución estacional de la descarga y la evapotranspiración en la cuenca. Como ejemplo, los intensos cambios en el uso del suelo junto a un clima regional marcadamente más cálido pueden ser dos condicionantes del fenómeno. Esto no excluye otros factores como los cambios en la demanda y abastecimiento de agua potable en la cuenca del Río Grande en las comunidades que se encuentran distribuidas a lo largo del río. Los resultados muestran que es aún temprano para observar impactos en los componentes del balance hídrico como resultado directo de la implementación de un esquema de PSB. Esto motiva la necesidad de una evaluación continua de la percepción local y un monitoreo los recursos hídricos para garantizar que las necesidades y demandas de servicios ecosistémicos en la región se mantengan a largo plazo.

A GIS-based method has been applied for assessing the spatial soil erosion risk and sediment yield in Mae Rim watershed, Northwest Thailand. The gross soil erosion in each watershed cell was estimated using the Revised Universal Soil Loss Equation (RUSLE) by thoroughly determining its various parameters. RUSLE results showed mean annual soil loss rate of 31 tonnes ha-1 yr-1, while the mean annual suspended sediment yield was 7.4 tonnes ha-1 yr-1. From the results of the spatial analysis between controlling factors and soil erosion, it indicated that bare land, field crop land and high steep slope were linked to extreme soil erosion (> 150 tonnes ha-1 yr-1). With respect to soil erosion under land use and rainfall change, it was revealed that the conversion from deciduous forest to field crop area has very serious implications for soil erosion in the Mae Rim watershed. Indeed, transition from forest to agriculture may lead to erosion increase despite reduced rainfall. The results obtained from analysing scenario sensitivities identify synergistic effects on soil erosion hazard if bare land, field crop land and rainfall erosivity are increased simultaneously in the future.

Global change phenomena, such as forest disturbance and land-use change significantly affect elemental balances as well as the structure and function of terrestrial ecosystems. Inappropriate land management often causes nutrient losses and finally soil degradation and loss of soil functioning. Especially in tropical ecoregions, soil degradation by nutrient losses is widely abundant. Soil microorganisms are the proximate agents of many processes performed in soils and are regarded as sensitive bio-indicators. However, the incorporation of microbial responses to the definition of critical soil conditions is not intensively developed. In the present thesis, several data analyses of the relationships between ecosystem disturbance and land-use change (natural forest, pastures of different ages, secondary succession) and a diverse set of soil ecological characteristics in the tropical mountain rainforest region of southern Ecuador were compiled. In particular, it was tested whether soil microbial biomass and community functioning were sensitive to land-use change effects. Furthermore, an information-theoretic approach was applied to find the factors that regulate soil microbial biomass and community function. Finally, in a nutrient enrichment experiment the above- and belowground responses to N and P additions were examined. The tested research questions and results were linked to the theory of ecological stoichiometry in order to connect the research to a sound and unifying scientific basis. Soil and microbial stoichiometry were affected by both land-use change and soil depth. After forest disturbance, significant decreases of soil C:N:P ratios at the pastures were fol-lowed by increases during secondary succession. Microbial C:N ratios varied slightly in response to land-use change, whereas no fixed microbial C:P and N:P ratios were observed. Shifts in microbial community composition were associated with soil and microbial stoichiometry. Strong positive relationships between PLFA-markers 18:2n6,9c (saprotrophic fungi) and 20:4 (animals) and negative associations between 20:4 and microbial N:P point to land-use change affecting the structure of soil food webs. Significant deviations from global soil and microbial C:N:P ratios indicated a major force of land-use change to alter stoichiometric relationships and to structure biological systems. Data analysis reveals a strong impact of land-use change on soil microbial biomass, C-mineralization, gross-NH4-consumption and –production rates. According to the results of the IT-approach, combined models better describe effects of land-use change on soil microorganisms than single explanation models. Microbial resources and soil chemical environment were important pre-dictors for soil microbial biomass and community functioning. Little is known about the environmental drivers of the catalytic properties of EHEs (e.g., pH, nutrients) and their functional link to the structure of soil microbial communities. The activities of the six hydrolytic enzymes were tested. Microbial production of AP responded to the low P status of the sites by a higher investment in the acquisition of P compared to C. Three major drivers of enzyme activities were found to be significant for enzyme production: 1.) Microbial demand for P regulated the production of AP, provided that N and C were available. At the natural forest site the two-fold higher specific activity of AP pointed to a high microbial P-demand, whereas the production of AP was constrained by the availability of N and DOC after pasture abandonment. 2.) Microbial biomass that was controlled by pH and resource availability was the main driver for CBH, BG and NAG activities. 3.) Substrate induction due to increased litter inputs of herbaceous plant species seemed to regulate AG and XYL activities during secondary succession. The enzymes’ affinity to substrate, as a potentially critically enzyme kinetic parameter is understudied. The data analysis suggests that microbial communities adapted to environmental changes, demonstrated high flexibility of extracellular enzyme systems and selected for enzymes with higher catalytic efficiency compared with pure cultures. Under in situ conditions, enzyme-specific environmental drivers of the Km, e.g., the pH for XYL, the C:N ratio for AP, and the C availability for NAG were found. The data demonstrated that the higher substrate affinity of XYL and AP was associated with more abundance of Gram(-) bacteria. The catalytic efficiency of enzymes decomposing cellulose, hemicellulose, and starch positively correlated with the relative abundance of Gram(-) bacteria. The turnover rate of the tested substrates was three to four times faster at the young pasture site compared with the longterm pasture and secondary succession sites. Nutrient inputs by atmospheric deposition are known to affect terrestrial ecosystems. However, little is known about how N and P co-limited ecosystems respond to single nutrient enrichment. In this work the susceptibility of above- and belowground ecosystem compo-nents and of their linkages in an N and P co-limited pasture to N- and P-enrichment was assessed. It was tested if the plants´responses can be explained by the concept of serially linked nutrients introduced by Ågren (2004). In this concept, the control of the growth rate by one nutrient is assumed to depend on the control of a different cellular process by another nutrient. The responses of shoot and root biomass and C:N:P stoichiometry of the grass Setaria sphacelata (Schumach.) to moderate N, P, and N+P application over five years were investigated. In addition, the effects of nutrient enrichment on soil nutrient pools, on arbuscular mycorrhizal fungi (AMF) as well as on microbial biomass, activity, and community structure were tested. In order to evaluate the importance of different factors explaining microbial responses, a likelihood-based information-theoretic approach was applied. The application of N+P increased aboveground grass biomass. Root biomass was stimulated by P-treatment. Grass C:N:P stoichiometry responded by altering the P-uptake or by translocating P from shoot to root. In particular, root C:N and C:P stoichiometry decreased in P- and in N-treatment. Extractable fractions of soil C, N, and P were significantly affected by nutrient enrichment. P application increased the biomass of Gram-positive bacteria and the abundance of AMF, however, results of the IT-approach suggested indirect effects of nutrient enrichment on microbes. The responses of the N and P co-limited pasture to particular nutrient enrichment support the concept of serially linked nutrients. The present study provides evidence for the fundamental importance of P for controlling resource allocation of plants in responses to nutrient enrichment. Resource allocation of the grass rather than direct effects of nutrient additions drives changes in AMF, microbial biomass, community structure, and activity<br>Seit dem Übergang vom Holozän zum Anthropozän greift der Mensch immer stärker in globale und regionale Stoffkreisläufe ein. Durch die Zerstörung von Naturwäldern und Landnutzungswandel werden die Strukturen und die Funktionen der Ökosysteme stark verändert. Unangepasste Landnutzung führt zu Nährelementverlusten, die mittel- bis langfristige zur Bodendegradation und zur Reduktion von Bodenfunktionen führen. Solche Veränderungen sind insbesondere in den Tropen zu beobachten. Bodenmikroorganismen spielen in den Stoffkreisläufen eine zentrale Rolle. Zudem sind sie sensitive Bioindikatoren für den Zustand von Ökosystemen. Im Gegensatz dazu, werden die Bodenmikroorganismen noch nicht ausreichend für die Zustandsbewertung von Ökosystemen verwendet. In der vorliegenden Dissertation werden verschiedene Datenanalysen zu den Beziehungen von Landnutzungswandel (Naturwald, Weiden verschiedener Alter, sekundäre Sukzession) und den Eigenschaften der Bodenmikroorganismen in einer tropischen Bergregenwaldregion Süd-Ecuadors zusammengefasst. Ein besonderer Fokus lag darauf zu prüfen, ob die mikrobielle Biomasse und die Funktionen die von der mikrobiellen Gemeinschaft geleistet werden (z.B. Enzymaktivitäten) durch den Landnutzungswandel beeinflusst werden. Ein informations-theoretischer Ansatz wurde verwendet um verschiedene Erklärungsansätze der steuernden Faktoren vergleichend zu testen. Darüber hinaus wurden in einem Weidedüngungsexperiment die Reaktionen der ober- und der unterirdischen Ökosystemkomponenten auf die Anreicherung mit N und P getestet. Um die Ergebnisse auf eine breite wissenschaftliche Basis zu stellen wurde die Untersuchungen in den Kontext der Theorie die Ökologischen Stöchiometrie eingeordnet. Die C:N:P Stöchiometrie im Boden und in den Mikroorganismen veränderte sich durch den Landnutzungswandel und mit der Bodentiefe. Mit der Weideetablierung nahmen die C:N:P Verhältnisse im Boden deutlich ab, stiegen dann nach dem Verlassen der Weiden im Zuge der sekundären Sukzession wieder an. Das mikrobielle C:N Verhältnis variierte nur leicht, dagegen zeigten das C:P und N:P Verhältnis deutliche Veränderungen durch den Landnutzungswandel. Mit diesen Veränderungen in der Boden- und Organismenstöchiometrie waren auch Veränderungen in der Struktur der mikrobiellen Gemeinschaften verbunden. Deutliche positive Beziehungen existierten zwischen den saprotrophen Pilzen und den Protozoen. Die steigenden Mengen von Protozoen waren wiederrum mit sinkendem mikrobiellen N:P verbunden. Diese Muster weisen auf Veränderungen in den Bodennahrungsnetzten durch Landnutzungsänderungen hin. Sehr deutliche Abweichungen von globalen Mustern der C:N:P Stöchiometrie deuten darauf hin, dass der Landnutzungswandel signifikanten Einfluss auf die C:N:P Stöchiometrie ausübt. Der Landnutzungswandel beeinflusste auch die mikrobielle Biomasse, die Basalatmung, sowie die mikrobielle Aufnahme und Produktion von NH4-N im Boden. Dabei zeigten kombinierte Erklärungsansätze die adäquateren Beschreibungen der Muster. In den kombinierten Modellen zur Erklärung der mikrobiellen Biomasse und der mikrobiellen Leistungen überwogen Prädiktoren der mikrobiellen Ressourcen und der bodenchemischen Umwelt. Ein weiterer Schwerpunkt der Untersuchungen lag auf der Erfassung der Effekte des Land-nutzungswandels auf die Aktivität von extrazellulären Bodenenzymen. Bisher ist wenig darüber bekannt, welche Faktoren die katalytischen Eigenschaften steuern und beispielsweise, ob es Zusammenhänge zur mikrobiellen Gemeinschaftsstruktur gibt. Um diese Fragen näher zu beleuchten wurden sechs hydrolytische Enzyme basierend auf MUF-Substraten untersucht. Die mikrobielle Produktion von AP stand dabei in Zusammenhang mit dem niedrigen P-Status der untersuchten Böden. Das wurde besonders durch die hohe AP Produktion im Vergleich zu BG belegt. Im Allgemeinen konnten drei verschiedene Mechanismen festgestellt werden, die die Produktion der untersuchten EHEs vermutlich steuerten. 1.) Der P-Bedarf der Mikroorganismen regulierte die Produktion von AP, vorausgesetzt, dass ausreichend N und C zur Enzymsynthese zur Verfügung standen. 2.) Die Höhe der mikrobiellen Biomasse hat sich als wichtiger Faktor für die Produktion von CBH, BG und NAG gezeigt. Das deutet auf die konstitutive Produktion dieser Enzyme hin. 3.) Die substratinduzierte Produktion von Enzymen ist vermutlich entscheidend für die Aktivität von AG und XYL. Die Berücksichtigung der Enzymkinetiken, insbesondere der Michaelis-Menten-Konstante lieferte weitere Aufschlüsse über relevante Faktoren. Im Allgemeinen so scheint es, haben sich die mikrobiellen Gemeinschaften an die starken Umweltgradienten, die durch den Landnutzungswandel erzeugt worden angepasst. Im Vergleich zu den verfügbaren Daten aus Reinkulturen, wiesen die mikrobiellen Gemeinschaften der untersuchten Böden in der Regel eine deutlich höhere katalytische Effizienz auf. Auch für die Michaelis-Menten-Konstante sind die Faktoren enzymspezifisch. So ist für die Km von XYL der Boden-pH-Wert, für AP das C:N Verhältnis und für NAG die DOC-Menge entscheidend. Darüber hinaus haben sich deutliche Beziehungen zwischen der Menge an Gram(-)-Bakterien und der Substrataffinitäten von XYL und AP ergeben. Je höher die Gram(-)-Abundanz, desto höher war die Substrataffinität der Enzymsysteme. Gegenüber alter und degradierter Weiden, war der Umsatz der untersuchten Substrate im Oberboden der aktiv genutzten Weide drei- bis vierfach erhöht. In einem 5-jährigen Düngeexperiment in der Bergregenwaldregion der Anden Süd-Ecuadors wurden die Reaktionen des auf dieser Fläche N/P co-limitierten Grases (Setaria sphacelata), der Arbuskulären Mykorrhiza (AMF) sowie der Bodenmikroorganismen auf moderate N, P und N+P-Düngung untersucht. Die Zugabe von N+P erhöhte die oberirdische Biomasse (+61%) wohingegen die Wurzelbiomasse durch die Zugabe von P (+45%) anstieg. Die C:N:P Verhältnisse weisen auf veränderte P-Aufnahme oder Translokation von P in die Wurzeln hin. Im Besonderen verengte sich das Wurzel C:N and C:P in der P- und der N-Zugabe. Die aus dem Boden extrahierbaren C, N und P-Fraktionen wurden deutlich beeinflusst. Die Zugabe von P stimulierte die Biomasse Gram-(+)-Bakterien (+22%), die Abundanz der AMF (+46%) und die Brutto-N-Mineralisierung. Die Auswertungen deuten darauf hin, dass die Nährstoffanreicherung indirekt über die Veränderungen der Graswurzeln auf die Bodenorganismen wirkte. Die Ergebnisse bestätigen, dass N und P in den Reaktionen von co-limitierten Pflanzen eng miteinander verbunden sind. Vor allem aber steuert P grundlegend die Allokation von Ressourcen und wirkt damit auf andere Ökosystem-komponenten, z.B. auf die Struktur und Aktivität der Bodenmikroorganismen

Les îles coralliennes d’atolls et les zones basses des îles hautes tropicales sont considérées comme comptant parmi les territoires les plus vulnérables aux impacts des événements météo-marins extrêmes et aux conséquences attendues du changement climatique. Néanmoins, alors que les littoraux métropolitains ont bénéficié d’un effort significatif d’évaluation de leurs évolutions passées de long terme, les espaces insulaires tropicaux ultramarins sont les moins documentés à l’échelle du territoire français. Partant de ce constat, cette thèse propose d’investiguer le rôle des facteurs météo-marins (cyclones, houles) et anthropiques (rigidification du trait de côte, remblaiement) dans les évolutions passées d’îles coralliennes d’atolls de Polynésie française et de plages d’îles hautes du nord des Petites Antilles.Ces travaux se basent sur un protocole méthodologique couplant l’apport de la géomatique et de la photo-interprétation assistée par ordinateur et des relevés réalisés sur les terrains étudiés. À partir d’une analyse multiscalaire (spatiale et temporelle), ils détectent et attribuent les évolutions planimétriques qu’ont connues les systèmes sédimentaires côtiers des îles étudiées. Les résultats obtenus sur les îles coralliennes abondent dans le sens des études antérieures qui établissaient que la majorité des îles a été stable ou en extension à l’échelle des dernières décennies. Ils permettent de proposer des modèles conceptuels de trajectoires d’évolutions pluridécennales et interrogent les contributions respectives des différents facteurs d’évolution considérés. Sur les plages des îles hautes, ces travaux mettent en évidence que les caractéristiques locales des sites étudiés expliquent la très forte variabilité longitudinale des évolutions détectées dans différents contextes météo-marins. Au-delà, ces travaux contribuent à alimenter l’échantillon mondial d’îles étudiées à ce jour et participent à enrichir la connaissance sur les évolutions passées des outremers insulaires français<br>Low-lying reef islands and coastal areas of tropical mountainous islands are highly vulnerable to the impacts of tropical cyclones and the expected effects of climate change. However, while the French metropolitan coasts have benefited from a significant effort to assess their long-term changes, French overseas islands are the least documented areas in French Territory. Based on this observation, this thesis examines the respective contribution of natural and anthropogenic drivers in the past evolution of reef islands (French Polynesia) and mountainous island beaches (Northern Lesser Antilles). This study relies on a combined approach between geomatic and fieldwork. From a multi-scalar (spatial and temporal) analysis, we detect and attribute the planimetric changes experienced by the sedimentary systems of the studied islands. Results on reef islands are comparable to those of previous studies which established that most of the islands have been stable or in expansion over the last decades. They allow to suggest conceptual models of long-term trajectories and examine the respective contribution of the drivers considered in this study. On mountainous island beaches, this study shows that local settings explain the high longitudinal variability detected in various climatic situations. In addition, this study contributes to the global samples of studied islands and to move forward on the understanding of past coastal changes in French overseas islands

I. An original methodological discussion is proposed on the problem of the typology of tropical rain forest’s plant communities, based on the study of forest types across gradients of continentality and elevation, within Atlantic central Africa. These investigations were based on the statement that the main problems in forest typology are related to the non-zonal or zonal character of the different vegetation types and to non considering the relations and differences between forest strata.<p><p>II. Field data consisted in phytosociological homogeneous sample plots localized within different recognized phytogeographical entities, in a region of tropical Africa where these entities are known to be well conserved. A total of 37 such plots were inventoried in the region extending from the littoral forests of Ndoté, Equatorial Guinea, which are wet evergreen forests, to the continental forests of the Dja, Cameroon, known as evergreen seasonal forests. The studied region also included the oriental Atlantic forests of Equatorial Guinea, known as moist evergreen forests or caesalp forests. In various parts of this continentality gradient, some plots were localized within climax non-zonal formations, namely the submontane rain forests. The emphasis was put on the vegetation of the Monte Alén National Park.<p><p>The sampling methodology was willing to be as "complete ", including all strata, "quantitative ", enumerating all individuals, and "representative ", within each stratum, as possible. These multi-layers plots were realised using nested sub-plots, with a sampling size of 100 individuals for every ligneous stratum recognized (dominant trees, dominated trees and shrubs) and a sampling size of 200m² for the herbaceous and suffrutex stratum.<p><p>Forest types were defined independently for each stratum and the differences were analysed. A method was proposed for the simultaneous analysis of all floristic data, converting and standardizing the values from ligneous strata, on the one hand, and from understorey strata, on the other hand.<p><p>III. Ten forest types were described using IndVal and discussed in the general context of the guineo-congolian region, from a syntaxonomic view point (agglomerative classification) and from a phytogeographical view point (divisive classification). Homologies between these two approaches are described. The proposed phytogeographical system is based on an "open " conception of hierarchical classifications, combining advantages of agglomerative and divisive classifications. In concrete terms, the non-zonal criteria, for example the submontane variants, are categorised separately and in analogy with the zonal criteria, related to the usual phytochoria.<p><p>Analysis of ecological relationships for the 10 communities showed that the main variables related to the floristic variability in our mainland rain forests are elevation, rainfall, hygrometry (estimated using bryophytes cover levels) and distance to the ocean. The two extremes on the vertical microclimatic gradient, dominant trees stratum and herbaceous stratum, give similar typologies, however canonical analysis showed that for the herbaceous layer, non-zonal variables (hygrometry and elevation) were gaining more importance when the influence of the two zonal variables was attenuated. In every case, spatial autocorrelation was less important than the environment in explaining floristic variability but its role increased in the spatial arrangement of understorey species, whose dispersal capacity is generally lower than canopy trees. The phytosociological, phytogeographical and ecological description of forest types is accompanied by a physiognomical description using biological types spectrum, as well as architectural models, leaf sizes, etc.<p><p>With regard to diversity, we have demonstrated that species richness was higher from upper to lower strata because of the accumulation in lower strata of species from various strata. On the other hand, the proper stratum diversity, i.e. the structural set, decreased from dominant trees to shrubs. The proper diversity of the herb layer showed relatively high figures mainly due to the higher individual density in relation to the existence of microstrata. Within the 37 sample plots, 1,050 taxa have been identified to species or morpho-species levels, for a total of 25,750 individuals. These taxa represent 442 genus among 104 families. The richest forest type is found on the foothills of the Niefang range, on the windward side. This forest type is also characterised by a high number of oligotypic genus and by species belonging to functional types indicators of glacial refuges. These functional types are defined on the basis of the dispersal capacity and on kind of stand needed for effective germination. We formulated the hypothesis that this kind of "foothills refuge ", characterised by his zonal nature, could have been one of the rare refuges for species from mainland rain forests, while montane and fluvial refuges would mainly have preserved species from non-zonal forest types: (sub)montane and riverine.<p><p>Based on indicator species of submontane forests, a potential distribution map of this forest type has been realised at the Atlantic central African scale. More than 400 submontane forest localities have been mapped. These forests begin at 400m of altitude near the ocean, and progressively at higher altitude for increasing distance to the ocean. Many lowland localities also comprised submontane species, which could indicate the existence of ecological transgressions. These transgressions would allow migratory tracks for submontane species between isolated mountain ranges, not only during glacial periods, through heights at the northern and southern borders of the congo basin, but also contemporarily through the lowland riverine forest network, in the centre of this basin. Finally, a special attention has been attributed to littoral forests and to some cases of choroecological transgressions, coupled to the ecological equalization phenomenon.<p><br>Doctorat en sciences agronomiques et ingénierie biologique<br>info:eu-repo/semantics/nonPublished

One of the profound realizations in Earth science during the last several decades has been that the solid earth and climate system interact through mountain belt evolution. Tectonic forces generate topography, and erosion, driven largely by the climate, destroys topography. Perturbations to the competition between these processes may, for example, have driven the transition from greenhouse to icehouse climate during the Cenozoic. Erosion is the ultimate connection between the climate and solid earth system, and because landscapes are shaped by erosion, they hold in their form information about climatic and tectonic forcings. Reading climatic and tectonic processes from the landscape requires an understanding of how these processes drive erosion. One way that climate influences erosion is by setting the elevation at which glaciation occurs. It has been thought for over a century that erosion by glaciers can limit the height of cold, heavily glaciated mountains. In this thesis, I argue that the prevalence of this phenomenon is underappreciated, and that glacial erosion has imposed an upper limit on the growth of warm, tropical mountains. The argument is premised on a combination of field observations from two (sub)tropical mountain ranges in Costa Rica and Taiwan (including 10Be and 3He surface exposure ages), a new method of topographic analysis that identifies previously unrecognized patterns of landscape rearrangement introduced by high elevation glaciation, and a study of ten tropical mountain ranges that reveals a widespread glacial control on their height. The results of this thesis demonstrate the efficacy of glacial erosion even in the warmest mountains, and challenge the hypothesis that quickly uplifting and eroding landscapes have approached a steady state balance between rock uplift and fluvial erosion during the Pleistocene.

碩士<br>國立中央大學<br>大氣物理研究所<br>101<br>First of all, this research has successfully established a tool to generate those wanted scenarios of balanced I.C.s for idealized WRF simulations and use WRF high resolution nested domains to figure out the rainfall patterns on the terrain. There are two experiments, one is to control vortex parameters, the other is to control terrain parameters. In the controlling vortex parameters experiment, we use a fixed terrain comparable to CMR with width 100km and length 300km. We find that the speed of steering flow cause the negative effect on the tropical cyclones track deflection and accumulated rainfall amount and the results are the same as previous studies. There is an interesting phenomenon that the patterns of rainfall are in phase between U1 group and U2 group neglecting the amount of rainfall. The results of the controlling terrain parameters experiments show that there is obvious evidence for that the direction of the tropical cyclones track deflection is dominated by the terrain length L_y and the degree of track deflection is dominated by the terrain altitude H. The tropical cyclones will turn left for enough large value of Ly. For example, case W1L3H. (about 3000km with H=1500m, critical value of non-dimensional parameter R_mw/L_y=0.027). Overall, the terrains make the cyclones speed down for 96-hr average translation speed both of two experiments and the faster the steering flow is, the more deceleration the cyclones will be. While only considering the upstream average translation speed of cyclones, U1 group is an exception. The terrain makes the U1 group most cases speed up. The upstream de-acceleration value is about 20% of the steering flow speed. With the vorticity budget analysis, the advection term dominate the voriticity tendency upstream, and the stretching term dominate when the cyclones passing through the mountain with strong blocking. Tilting term also has much contribution to the vorticity tendency while the cyclones across the terrain. The results of vorticity budget analysis are the same as Lin et al (2011). According to the results of terrain steepness experiments, the upstream track deflection is not sensitive to the steepness of the terrain (parallel to the moving direction of cyclones) of the mountain. At last, the direction of cyclones track deflection has a lot of to do with the horizontal asymmetric wind field. The cyclone tends to turn right with the maximum wind appearing at the east side of the cyclone, and tend to turn left with the maximum wind appearing at the west side of the cyclone. This can also be applied to explain the phenomenon of the upstream anti-cyclone track found by Yeh and Elsberry (1993).

<p>Global climate change can cause shifts in species distributions, and increases in some of their competitors, predators, and diseases that might even cause their extinction. Species may respond to a warming climate by moving to higher latitudes or elevations. Shifts in geographic ranges are common responses in temperate regions. For the tropics, latitudinal temperature gradients are shallow: the only escape for species may be to move to higher elevations. There are few data to suggest that they do, and our understanding of the process is still very limited. Yet, the greatest loss of species from climate disruption may be for tropical montane species. To better understand the potential process of elevational range shifts in the tropics and their implications we have to: 1) Build theoretical models for the process of range shifting, 2) Evaluate potential constraints that species could face while moving to higher elevations, 3) Obtain empirical evidence confirming the uphill shift of species ranges, 4) Determine the number of extinctions that could arise from elevational range shifts (mountain top extinctions) and 5) Identify vulnerable species and areas, and determine their representation by the Protected Areas Network. The purpose of this dissertation is to address these issues, by applying novel methods and collecting empirical evidence. </p><p>In the second chapter I incorporated temperature gradients and land-cover data from the current ranges of species in a model of range shifts in response to climate change. I tested 4 possible scenarios of amphibian movement on a tropical mountain and estimated the constraints to range shifts imposed by each scenario. Confirming the occurrence of elevational range shifts with empirical data is also essential, but requires historical data as a baseline for comparison. I repeated a historical transect in Peru, sampling birds at the same locations they were sampled 40 years ago, and compared their elevational ranges between sampling occasions to evaluate if they were moving uphill as a response to warming temperatures. Finally, based on the results from this comparison, I estimated the potential extinctions derived from elevational range shifts, using information on the species distribution, the topography and land cover within the ranges and surrounding areas. I evaluated the extent of mountain top extinctions for 172 bird species with restricted ranges in the northern Andes. I also considered how Colombia's protected Area Network represents species and sites that are vulnerable in the face of climate change.</p><p>More than 30% of the range of 21 of 46 amphibian species in the tropical Sierra Nevada de Santa Marta is likely to become isolated as climate changes. More than 30% of the range of 13 amphibian species would shift to areas that currently are unlikely to sustain survival and reproduction. Combined, over 70% of the current range of 7 species would become thermally isolated or shift to areas that currently are unlikely to support survival and reproduction. The constraints on species' movements to higher elevations in response to climate change can increase considerably the number of species threatened by climate change in tropical mountains.</p><p>In the comparison of bird distributions in the Cerrros del Sira, in Peru, I found an average upward shift of 49 m for 55 bird species over a 41 year interval. This shift is significantly upward, but also significantly smaller than the 152 m one expects from warming in the region. The range shifts in elevation were similar across different trophic guilds. Endothermy may provide birds with some flexibility to temperature changes and allow them to move less than expected. Instead of being directly dependent on temperature, birds may be responding to gradual changes in the nature of the habitat or availability of food resources, and presence of competitors. If so, this has important implications for estimates of mountaintop extinctions from climate change. </p><p>The estimated number of mountain top extinctions from climate disruption in the northern Andes is low, both the absolute number (5 species) and the relative number (less than 0.5% of Colombian land birds). According to future climate predictions these extinctions will not likely occur in this century. The extent of species loss in the Andes is not predicted by absolute mountaintop extinctions modeled by the kind of processes most other studies use. Rather, it is highly contingent -- the species will survive or not depending on how well we protect their much reduced ranges from the variety of other threats.</p><br>Dissertation

Complex interactions between topographic heterogeneity and steep gradients in climate and environmental conditions are commonly assumed to promote biotic diversification. Using tropical frogs as a model, I investigate the nature of these interactions that disrupt migration between populations, causing genetic divergence and speciation. I determine the role of several putative factors that affect gene flow (Euclidean distances, Least Cost Path (LCP) distances, topographic complexity, and elevation difference) and promote genetic structure (FST) between populations of three tropical Andean frog species. Moreover, I investigate, from an intraspecific perspective, whether montane frog species display on average larger genetic distances per kilometer relative to lowland species. Finally, I test if recent genetic divergence caused by topographic barriers to gene flow is paralleled by independent character systems such as acoustics and morphological traits in the high Andean frog Dendropsophus labialis. Even though the effect of geographic features on migration (and conversely, FST) was species-specific, LCP and Euclidean distances had the strongest effect on migration rate. Topographic complexity also reduced migration rate whereas elevation difference did not have an effect. I found that indeed highland species show larger genetic distances per kilometer between haplotypes than do lowland species. Also, genetic divergence is strongly associated with topographic heterogeneity, which is an intrinsic characteristic of montane regions. Finally, I found that acoustic variation in D. labialis diverges according to genealogical history, but external morphology does not follow this relationship. Stochastic processes due to genetic drift appear to be a better explanatory mechanism for the divergence in calls than adaptive variation. The strong and congruent divergence observed in acoustic and genetic characters indicates that these two groups correspond to morphologically cryptic parapatric species. Overall, the results of this study suggest some of the mechanisms that allow tropical mountains to promote intraspecific genetic divergence. The combined effect of ridges (promoting allopatric differentiation) and environmental gradients across elevation (promoting parapatric differentiation) are effective forces that are present mostly in highland biomes. Unfortunately, such biomes are critically threatened by habitat destruction and climate change, possibly more than any other biome on earth.<br>text

O bioma Mata Atlântica é mundialmente reconhecido como uma das regiões de maior diversidade biológica do planeta, abrigando elevada riqueza de espécies e um elevado número de espécies endêmicas, se estendendo por cerca de 1.300.000 km2 ao longo da costa brasileira, entre as latitudes 3º e 33º S e longitudes 35º e 57º L. Como resultado do incremento das atividades humanas de uso da terra e, consequentemente, de ampla modificação da paisagem ao longo dos séculos, cerca de apenas 10-15% do bioma Mata Atlântica ainda se encontra em estado natural ou próximo ao natural, sendo considerados áreas prioritárias para conservação. Devido à grande variedade climática e geomorfológica, o bioma Mata Atlântica é um complexo mosaico de diferentes ecossistemas. Dentre estes, a floresta com Araucária (Floresta Ombrófila Mista), a floresta nebular (Floresta Ombrófila Densa Altomontana) e os campos de altitude ocupam as médias e altas altitudes da Serra do Mar, que se estende por cerca de 1000 km paralela à costa, do sul ao sudeste brasileiro. A distribuição da floresta com Araucária está relacionada ao clima úmido e relativamente frio, entre 400 e 1400 m s.n.m. no sul do Brasil e em fragmentos menores entre as altitudes de 1400 a 1800 m s.n.m. no Sudeste. Atualmente, está reduzida a não mais do que 7% da sua distribuição original. A floresta nebular se estende nas encostas do alto da Serra do mar, normalmente acima de 1100 m s.n.m. no Sul e acima de 1500 m s.n.m. no sudeste do Brasil, nos pequenos vales e sítios protegidos. Os campos de altitude são uma vegetação tipicamente herbácea, restrita aos cumes e picos da serra e aos platôs mais elevados. Estudos paleoecológicos demonstraram que, apesar deste mosaico de ecossistemas ter persistido durante o Holoceno, a perpetuação dos campos de altitude é muito frágil. Como a vegetação campestre se expande em condições climáticas mais frias e secas e parece ser adaptada ao fogo, sugere-se que a presente área de campos de altitude é maior do que esperada sobre as condições climáticas atuais, especialmente em locais mais quentes em altitudes mais baixas. Além disso, estudos de mudanças climáticas preveem um clima mais quente e úmido durante o século 21, que provavelmente irá intensificar a migração da floresta atlântica para maiores altitudes, em detrimento da vegetação campestre. Nesta pesquisa, as relações passadas e presentes do mosaico de campos de altitude e florestas altomontanas (floresta com Araucária e floresta Atlântica nebular) são exploradas por meio de análises palinológicas. Inicialmente, foi investigada a correlação atual entre cobertura vegetal e produção de pólen. Observou-se que os taxa arbóreos são superestimados no conjunto de pólen de campos de altitude, constituindo uma área muito maior de captação de pólen do que no conjunto de pólen arbóreos. Sendo assim, o conjunto de pólen que caracteriza a vegetação de campos de altitude apresenta uma grande proporção de taxa de vegetação arbórea. Posteriormente, um sedimento de quase 10.000 anos foi analisado, demonstrando que, apesar dos taxa representantes da floresta altomontana estarem presentes na região de estudo durante todo o Holoceno, a vegetação florestal expandiu majoritariamente durante o Holoceno Tardio. Até cerca de 1350 cal a AP, a vegetação de campos de altitude ocupava áreas mais extensas. Em geral, os resultados demonstraram que o aumento de temperatura e precipitação ao longo do Holoceno favoreceram a migração da floresta para altitudes mais elevadas. Além disso, a pesquisa indicou que o fogo já estava presente na região antes da chegada dos primeiros humanos no Sudeste do Brasil, implicando na adaptação da vegetação campestre ao fogo. Por último analisou-se a dinâmica da vegetação nos últimos sete séculos. Os resultados indicaram que interferências antropogênicas como fogo, pastoreio e exploração madeireira desempenharam um importante papel na relação campos-floresta na Serra do Mar do Sudeste do Brasil. Com base nestes estudos, sugere-se que a manutenção do mosaico de campos de altitude e floresta no clima presente e futuro depende tanto de um manejo ativo quanto da mudança de foco da conservação de ambientes florestais para ambientes campestres.

Zur Rekonstruktion der jungquartären Landschaftsentwicklung am Kilimanjaro werden Sedimente aus Paläoböden am Mt Kilimanjaro untersucht, um die lokale und regionale Ökosystem-, Klima-, Feuerdynamik in einem größeren Rahmen zu verstehen. Desweiteren soll die Reaktion der Ökosysteme auf Umweltveränderungen erforscht werden, um die Beziehungen verschiedener Ökosysteme und ihre Rolle hinsichtlich der Entwicklung von „Biodiversity Hot Spots“ in Ostafrika aufzudecken. Die Ergebnisse tragen zu einem besseren Verständnis der heutigen und zukünftigen Dynamik von Ökosystemen bei. Sie sind außerdem eine wichtige Grundlage im Naturschutz, da sie bedeutende Informationen für die Aufrechterhaltung und das Management der hohen Biodiversität in den ostafrikanischen Hochländern liefern. Die Durchführung von paläoökologischen Untersuchungen ist unerlässlich, da die Prozesse der Vergangenheit eine wichtige Rolle bei der Entwicklung der Ökosysteme und Biodiversität spielen. Die vorliegende Arbeit erforscht die spätquartäre Vegetation-, Klima- und Feuerdynamik am Kilimanjaro, um das Verständnis der dortigen Ökosystem zu vertiefen. Sowohl Pollen als auch weitere Klima-Proxies von zwei Sedimentkernen werden analysiert, um die Vegetationsdynamik vom frühen Spätglazial bis heute zu rekonstruieren und die jeweiligen Einflußfaktoren aufzudecken. Die Ergebnisse vom Maundi Krater, welcher auf 2780 m Höhe am südostlichen Hang des Kilimanjaros gelegen ist, stellen eines der längsten, terrestrischen Klimaarchive im tropischen Ostafrika dar, und ermöglichen den Einblick in fast 100 000 Jahre Vegetations- und Klimageschichte. Das WeruWeru Paläobodenprofil aus dem montanen Waldgürtel am Kilimanjaro ermöglicht die detaillierte Rekonstruktion der Reaktionsdynamik der Vegetation auf Veränderungen in der Umwelt. Die Ergebnisse beider Untersuchungen zeigen, dass Klimaveränderungen zu einer Verschiebung der montanen Vegetationsgürtel entlang des Höhengradienten des Mt Kilimanjaro geführt haben. Das Pollenarchiv des WeruWeru Profils dokumentiert starke Veränderungen in den höchsten Vegetationgürteln, dem Erikagürtel und dem oberen montanen Wald. Trotz markanter Klimaveränderungen scheint Mt Kilimanjaro aber auch als eiszeitliches Refugium für Waldarten gedient zu haben. Feuerereignissen spielten hinsichtlich der Ausbildung des Erikagürtels eine entscheidende Rolle. Hinweise auf verstärkte menschliche Aktivitäten können an keinem der beiden Untersuchungsstandorte festgestellt werden. Die Pollenregen-Studie entlang des Höhengradienten am Kilimanjaro zeigt, dass es sehr entscheidend ist, die quantitative Beziehung zwischen Pollen-Niederschlag und moderner Vegetation zu untersuchen, um die Rekonstruktionen der Vergangenheit zu kalibrieren. Die Ergebnisse ermöglichen eine weitaus präzisere Interpretation der Vegetations- und Klimarekonstruktionen im tropischen Ostafrika.

Tropical mountain rain forests are among the most species rich regions in the world. The RBSF (Reserva Biológica San Francisco) area in southern Ecuador is a hotspot of biodiversity. To characterise this exceptional high diversity it is essential to investigate the complex interactions between the organisms and their abiotic environment. A number of groups of organisms, including birds, moths, orchids and mosses, have been investigated, whereas knowledge on others, including soil invertebrates, is minute. This study focuses on (1) the characterisation of the hidden diversity of soil animal species, in particular that of oribatid mites (Oribatida), (2) the composition and function of soil microarthropods including their trophic relationship, (3) decomposition processes including altitude (temperature, moisture), litter type (low and high quality) and microarthropods (small and large mesh of litterbags). (1) At the study site 193 species of 48 families of oribatid mites were determined which is high but compared to temperate forest ecosystems not extraordinary divers. The proportion of not described species is assumed to be around 40 %. Nine new species of ptyctimous oribatid mites were described. Many species are restricted to tropical regions indicating specific community structures in the tropics. Furthermore, the oribatid mite community differed along the elevation gradient from 1850 to 2270 m indicating distinct oribatid mite communities at different altitudes. (2) The high diversity of oribatid mites and decomposer soil animals in general, is one of the great riddles in soil ecology. Whereas in deciduous forests the soil microarthropods are well characterised, taxonomical knowledge in tropical rainforests and especially in mountain regions is low. One aim of this study was to investigate the microarthropod soil community. The density of microarthropods and oribatid mite communities at three horizons (L, F/H, Ah) and on the bark of adjacent trees along an elevation gradient (1850, 2020, 2200 and 2270 m) in the RBSF forest were studied. Oribatid mites were the most abundant group followed by Collembola and Gamasina. The compared to temperate forests generally low number of microarthropods declined with elevation in the order 1850 > 2020 > 2270 ~ 2200 m and with soil depth (L, F/H, Ah). Microarthropods on bark were less abundant than in soil. Declining numbers of soil microarthropods with altitude is concluded to be due to harsh abiotic conditions (e.g. lower temperatures, waterlogging, solar radiation). Low microbial biomass and resource quality presumably also contribute to the low abundances of soil microarthropods. The trophic structure of 32 species and potential basal food resources (litter of Graffenrieda emarginata) was investigated by analysing natural variations in stable isotope ratios (15N/14N; 13C/12C). The results indicate that the soil food web is similar to that of temperate forests and spans about four trophic levels. Primary decomposers, i.e. litter feeding species, were rare, potentially reflecting low litter quality. A large number of ‘decomposer’ animals were in fact predatory or necrophagous, suggesting that various putative decomposer soil animal species (especially in oribatid mites) presumably feed on other soil invertebrates, in particular nematodes or animal carcasses. (3) We studied leaf litter decomposition of two abundant tree species with higher and lower litter quality (Graffenrieda emarginata, Purdiaea nutans) and a mixture of both in a litterbag field experiment at two altitudes (1850 and 2280 m). Litter quality was measured as microbial biomass and N content. Decomposition rates at the studied tropical mountain rain forest were generally low (average of 32 % y-1). The slow decomposition rates may have been due to low litter quality (e.g. high C-to-N ratios) but also due to low temperatures. Litter generally decomposed slower at higher elevation supporting our assumption that temperature is a major driving force for litter decomposition at our study sites and indicating that organic matter accumulates at high altitudes. P. nutans litter after 2 and 6 months of exposure decomposed slower than that of G. emarginata, but not at the end of the experiment, after twelve months. We suggest that litter chemistry affect decomposition mainly at early stages of decomposition. After 12 months the mixture of G. emarginata and P. nutans litter decomposed significantly faster than both single litter types indicating that combining the two litter types accelerates decomposition processes ('non-additive effect'). Soil microarthropods contributed little to decomposition processes. Microbial biomass and density of microarthropods in the litterbags were higher at 1850 than at 2280 m indicating higher biological activity at lower altitudes. Species composition was similar in both litter types supporting previous findings that the structure of soil decomposer microarthropod communities is little affected by litter type.

碩士<br>國立臺灣大學<br>地理環境資源學研究所<br>106<br>Fine root, with diameter ≤ 2 mm, takes responsibility for water and nutrient transport. Besides being a carbon pool, it also provides a carbon flux pathway through plant canopy and soil, to contribute 30% to 70% of total net primary production (NPP) in the forest ecosystem, thus serving a dynamic role in its carbon cycling. Therefore, quantifying fine root productivity is necessary to study terrestrial carbon budget. However, compared to abundant studies on aboveground leaf and canopy, less is known about belowground root system due to its difficulty on observation and quantification. Based on the phenological correlation, we aimed to model fine root production by aboveground parameters. The study was conducted in Chilan Mountain in northeast Taiwan, a near tropical moist forest. A study plot was set, with 15 minirhizotron tubes installed and root densities measured. We took fine root photographs by an interval of three weeks; presence and absence of fine roots were delineated by image processing algorithms to derive fine-root production (g C ha-1 d-1) through time. As to the aboveground, we took biological and environmental factors into account, including litterfall production (branches and leaves), canopy density (leaf area index, LAI), meteorological (air temperature, air humidity, precipitation, solar radiation, wind speed), and soil physical conditions (soil temperature, soil moisture). As a result, the variables Leaf Area Index (LAI), △LAI, solar radiation, precipitation, wind speed, and soil temperature were particularly significant to root production, with adjusted R2 of 0.42 (n = 36, p-value < 0.005) by multiple linear regression. Synchrony existed in leaf flush and fine root emergence; in addition, environmental restrictions were more evident in soil than air conditions, where air temperature and precipitation were not limiting factors, which added the uniqueness to near tropical wet forests under global warming. This study demonstrated the feasibility of utilized aboveground variables to indirectly assess fine root growth. The models could reveal fine root dynamics through time, or be further applied on the regional scale mapping with aid of remote sensing, where more validation need to be done to fill the intra- and inter-annual, plot- and region-scale gaps.

This diploma thesis focuses on the implementation of the Czech approach of nature conservation management planning to the submontane rain forest in the Philippines. This work is based on own research supplemented by literary overview. Two main researched sites in Don Salvador Benedicto, Negros Occidental, as well as the comparative site in Patag, were researched during the September 2017. All the sites were completely inventoried and described the forest structure, tree species composition and other characteristics. The results of the work are a detailed description of both mentioned forest communities in Negros Occidental, the proposal of target forest community and forest management plan for the locality of Don Salvador Benedicto.

El sur de los Andes Ecuatorianos alberga una extraordinaria riqueza de especies. Muchos factores ambientales diferentes influyen entre sí en un espacio muy limitado y crean ecosistemas únicos y complejos. Sin embargo, esta área está altamente en peligro debido al creciente impacto humano a través de la intensificación del uso de la tierra y cambio global. Sólo poco se sabe acerca de la historia paleoecológica y dinámica del paisaje de esta zona. Sin embargo, la información acerca del por qué y cómo los ecosistemas han cambiado en el pasado es crucial para el desarrollo de estrategias innovadoras para la conservación y futuras predicciones del clima. En este estudio, presentamos los análisis palinológicos realizados en la región sur de los Andes Ecuatorianos, que ayudan aclarar los patrones y procesos de los ecosistemas presentes y pasados. El estudio paleoecológico de la cuenca volcánica Quimsacocha en la cresta oriental de la Cordillera Occidental revela cambios del clima, la vegetación y del régimen del fuego desde el período del Holoceno temprano. El Holoceno medio fue un período de severos cambios ambientales debido a un clima más seco y supuestamente más cálido en esta área. Durante el Holoceno tardío, varias fases cálidas y frías se indican en el registro. El fuego estuvo presente en la zona desde el Holoceno temprano y puede ser el primer signo del impacto humano. El análisis multivariado combinado con otros núcleos de sedimentos en el sur de los Andes Ecuatorianos reveló acontecimientos, en parte, contrastantes, los cuales supuestamente son debido a la heterogeneidad ambiental de los diferentes sitios. Por otra parte, un estudio de tres años entre la relación lluvia de polen actual – vegetación se llevó a cabo en la región del Parque Nacional Podocarpus, con el fin de comprender los patrones de dispersión de polen de los diferentes tipos de vegetación del bosque premontano, bosque montano bajo, bosque montano alto y páramo para crear una base más sólida para la interpretación de los datos de polen fósil. Una comparación entre la abundancia y los datos de presencia-ausencia a nivel de familia para el polen y la vegetación mostró que los patrones de diversidad, distribución y abundancia tienen una buena correspondencia el uno con el otro en ambos conjuntos de datos. Sin embargo, cantidades variables del polen transportado de larga distancia, la productividad desigual de polen de los diferentes taxones y los sistemas heterogéneos del viento impactan los patrones. Los análisis de las tasas de acumulación de polen indican poca variación interanual pero una alta variación espacial en los datos de polen. La evaluación de los dos diferentes tipos de trampa de polen utilizados con frecuencia en estudios palinológicos tropicales, la trampa Oldfield modificada y la trampa Behling, en comparación con muestras de suelo superficial y una trampa de referencia, mostró que en las muestras de suelo, los taxones con una exina frágil se representan en menor medida que en las trampas. Además, señaló que mientras que en el bosque, todos los tipos de trampas proporcionan resultados similares, la trampa Behling se debe preferir en el páramo, ya que resiste mejor a la alta radiación y periodos de sequía. Todos los análisis amplían el conocimiento preciso y completo de la dinámica de la vegetación de los Andes Tropicales en el espacio y el tiempo.