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1

CALVO, JOEL, MÉLICA MUÑOZ-SCHICK, and ANDRÉS MOREIRA-MUÑOZ. "Xenophyllum juniperinum (Compositae, Senecioneae), a new required combination." Phytotaxa 326, no. 3 (2017): 227. http://dx.doi.org/10.11646/phytotaxa.326.3.7.

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Due to ongoing studies on the austral species of Xenophyllum and Werneria, we noticed that some taxonomic and nomenclatural adjustments are required. Herein, we propose a new combination, Xenophyllum juniperinum.
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CALVO, JOEL, MÉLICA MUÑOZ-SCHICK, and ANDRÉS MOREIRA-MUÑOZ. "Towards a better understanding of Xenophyllum esquilachense (Senecioneae, Compositae), a poorly known Andean species." Phytotaxa 382, no. 3 (2018): 288. http://dx.doi.org/10.11646/phytotaxa.382.3.5.

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Xenophyllum esquilachense is a poorly known Andean species that was hitherto considered endemic to southern Peru. It is recorded for the first time from Bolivia and northern Chile, and the name Senecio pfisteri is synonymized with it. New insights on its global occurrence, ecology, and taxonomy are provided, as well as pictures of living plants and an updated dichotomous key to the Xenophyllum species from southern Andes.
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Calvo, Joel, and Andrés Moreira-Muñoz. "Taxonomic revision of the Andean genus Xenophyllum (Compositae, Senecioneae)." PhytoKeys 158 (September 3, 2020): 1–106. http://dx.doi.org/10.3897/phytokeys.158.50848.

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The Andean genus Xenophyllum (Compositae, Senecioneae) is distributed along the high-Andes from northeastern Colombia to northern Chile and northwestern Argentina, mainly thriving in the paramo and puna ecoregions. It comprises suffruticose plants forming dense mats, hummocks, or clumps of erect stems. They are characterized by displaying involucral bracts fused at the base, supplementary bracts absent, and mostly radiate capitula with white ray corollas, seldom yellow or pink (disciform in one species). Traditionally, Xenophyllum species were treated as members of the genus Werneria, a morphologically close genus that includes rosettiform or scapiform perennial herbs. As currently circumscribed, Xenophyllum mostly differs from Werneria in having elongate stems. Herein, the first modern and comprehensive revision of the genus recognizing twenty-two species and two subspecies is presented. Werneria decumbens is synonymized with X. weddellii, as well as X. fontii with X. humile and X. oscartovarii with X. dactylophyllum. Likewise, four varietal names and two sectional names are proposed as new synonyms. Seven names are lectotypified, the name X. sotarense is epitypified, W. decumbens neotypified, and the supraspecific name W. sect. Integrifoliae Rockh. is typified. The combination X. crassum subsp. orientalecomb. nov. is made. Descriptions and distribution maps are provided for all accepted species, in addition to an identification key. Ten species are illustrated, three of them for the first time.
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Calvo, Joel, and Andrés Moreira-Muñoz. "Taxonomic revision of the Andean genus Xenophyllum (Compositae, Senecioneae)." PhytoKeys 158 (September 3, 2020): 1–106. https://doi.org/10.3897/phytokeys.158.50848.

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The Andean genus Xenophyllum (Compositae, Senecioneae) is distributed along the high-Andes from northeastern Colombia to northern Chile and northwestern Argentina, mainly thriving in the paramo and puna ecoregions. It comprises suffruticose plants forming dense mats, hummocks, or clumps of erect stems. They are characterized by displaying involucral bracts fused at the base, supplementary bracts absent, and mostly radiate capitula with white ray corollas, seldom yellow or pink (disciform in one species). Traditionally, Xenophyllum species were treated as members of the genus Werneria, a morphologically close genus that includes rosettiform or scapiform perennial herbs. As currently circumscribed, Xenophyllum mostly differs from Werneria in having elongate stems. Herein, the first modern and comprehensive revision of the genus recognizing twenty-two species and two subspecies is presented. Werneria decumbens is synonymized with X. weddellii, as well as X. fontii with X. humile and X. oscartovarii with X. dactylophyllum. Likewise, four varietal names and two sectional names are proposed as new synonyms. Seven names are lectotypified, the name X. sotarense is epitypified, W. decumbens neotypified, and the supraspecific name W. sect. Integrifoliae Rockh. is typified. The combination X. crassum subsp. orientale comb. nov. is made. Descriptions and distribution maps are provided for all accepted species, in addition to an identification key. Ten species are illustrated, three of them for the first time.
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Calvo, Joel, Mélica Muñoz-Schick, and Andrés Moreira-Muñoz. "Xenophyllum juniperinum (Compositae, Senecioneae), a new required combination." Phytotaxa 326, no. 3 (2017): 227–29. https://doi.org/10.11646/phytotaxa.326.3.7.

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Calvo, Joel, Muñoz-Schick, Mélica, Moreira-Muñoz, Andrés (2017): Xenophyllum juniperinum (Compositae, Senecioneae), a new required combination. Phytotaxa 326 (3): 227-229, DOI: 10.11646/phytotaxa.326.3.7, URL: http://dx.doi.org/10.11646/phytotaxa.326.3.7
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6

Calvo, Joel, Mélica Muñoz-Schick, and Andrés Moreira-Muñoz. "Towards a better understanding of Xenophyllum esquilachense (Senecioneae, Compositae), a poorly known Andean species." Phytotaxa 382, no. 3 (2018): 288–92. https://doi.org/10.11646/phytotaxa.382.3.5.

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Calvo, Joel, Muñoz-Schick, Mélica, Moreira-Muñoz, Andrés (2018): Towards a better understanding of Xenophyllum esquilachense (Senecioneae, Compositae), a poorly known Andean species. Phytotaxa 382 (3): 288-292, DOI: 10.11646/phytotaxa.382.3.5, URL: http://dx.doi.org/10.11646/phytotaxa.382.3.5
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7

Calvo, Joel, and Vicki A. Funk. "Two new species of the Andean genus Xenophyllum (Senecioneae, Compositae)." PhytoKeys 139 (January 27, 2020): 29–38. http://dx.doi.org/10.3897/phytokeys.139.47872.

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Two new species of Xenophyllum are described from the Andes, X. funkianumsp. nov. from Ecuador and X. lorochaquisp. nov. from northwestern Argentina. Both species are compared with the morphologically closest taxa and useful characters for their proper identification are provided. Detailed illustrations and distribution maps are also presented, as well as pictures of living plants when available.
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8

Calvo, Joel, and Vicki A. Funk. "Two new species of the Andean genus Xenophyllum (Senecioneae, Compositae)." PhytoKeys 139 (January 27, 2020): 29–38. https://doi.org/10.3897/phytokeys.139.47872.

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Two new species of Xenophyllum are described from the Andes, X. funkianum sp. nov. from Ecuador and X. lorochaqui sp. nov. from northwestern Argentina. Both species are compared with the morphologically closest taxa and useful characters for their proper identification are provided. Detailed illustrations and distribution maps are also presented, as well as pictures of living plants when available.
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9

Benites, J., J. Lopez, L. E. Rojo, et al. "Chemical composition of the essential oil of Xenophyllum poposum." Chemistry of Natural Compounds 46, no. 6 (2011): 988–89. http://dx.doi.org/10.1007/s10600-011-9806-7.

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10

Sklenar, Petr, and Katya Romoleroux. "Werneria spathulata, nuevo registro para la flora del Ecuador, y redescubrimiento de Xenophyllum acerosum." Caldasia 43, no. 1 (2020): 214–17. http://dx.doi.org/10.15446/caldasia.v43n1.81909.

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Se presenta un nuevo reporte de Werneria (Asteraceae) para Ecuador, es W. spathulata, una pequeña hierba rizomatosa encontrada en los páramos del volcán Antisana. Además, se documenta el redescubrimiento de Xenophyllum acerosum en el Ecuador después de su primera colección hace 73 años en la provincia de Azuay. Se proveen fotografías del hábito de las dos especies, así como del hábitat donde fueron halladas.
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11

Espinoza Calderón, Gustavo Adolfo, Carmen Taipe Lucas, and Magdalena Taipe Yalli. "Modelamiento matemático del proceso de deshidratado de china konoca (Xenophyllum poposum) por aire caliente." Revista Alfa 5, no. 13 (2021): 98–108. http://dx.doi.org/10.33996/revistaalfa.v5i13.102.

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Durante mucho tiempo las plantas medicinales fueron el principal recurso disponible para tratar enfermedades. Por esta razón fueron desarrollándose numerosos estudios sobre esas plantas e investigaciones sobre el uso de sus productos y derivados. La china konoca (Xenophyllum poposum) es una planta nativa que crece de manera silvestre en los Andes peruanos a más de 4400 metros sobre el nivel del mar, es muy utilizada como una infusión natural para la digestión. El propósito de este trabajo fue realizar un modelado matemático durante el proceso de deshidratado de la china konoca (Xenophyllum poposum). Para ello, se efectuó la evaluación de cinética de deshidratado de esta planta a 60°C y 80°C, durante 480 y 600 minutos, en donde se hizo las mediciones de pérdida de humedad. Se obtuvieron gráficas de velocidad de deshidratado en función del tiempo, temperatura y la humedad, con estos datos se realizó un modelado matemático de cada tratamiento para describir la tendencia en las curvas de deshidratado. Resultando un modelo polinómico de segundo grado el que mostró mejor bondad de ajuste en la cinética de secado por aire caliente, %H = a – b ? +c ? 2. Además, se hizo el modelamiento para predecir el tiempo de equilibrio en el decrecimiento de humedad, derivando la ecuación anterior, ?=b/2c.
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12

Funk, V. A. "Xenophyllum, a New Andean Genus Extracted from Werneria s.l. (Compositae: Senecioneae)." Novon 7, no. 3 (1997): 235. http://dx.doi.org/10.2307/3391934.

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13

de Marchese, María J. A., Carola S. de Heluani, César A. N. Catalán, Claire A. Griffin, Joseph B. Vaughn, and Werner Herz. "Incisol, an alcohol with a novel sesquiterpene skeleton from Xenophyllum incisum." Biochemical Systematics and Ecology 35, no. 3 (2007): 169–75. http://dx.doi.org/10.1016/j.bse.2006.10.006.

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14

Romano, E., A. B. Raschi, A. M. Benavente, A. M. Gonzalez, and O. E. Piro. "Characterization and crystal structure of 2R-(−)-6-hydroxytremetone from Xenophyllum poposum (Phil.) V.A. Funk." Natural Product Research 22, no. 2 (2008): 124–35. http://dx.doi.org/10.1080/14786410701589899.

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15

González, Ana M., María I. Tracanna, Sara M. Amani, et al. "Chemical Composition, Antimicrobial and Antioxidant Properties of the Volatile Oil and Methanol Extract of Xenophyllum Poposum." Natural Product Communications 7, no. 12 (2012): 1934578X1200701. http://dx.doi.org/10.1177/1934578x1200701230.

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The essential oil and methanol extract of northwestern Argentina medicinal plant Xenophyllum poposum, collected in Catamarca province, were investigated. GC and GC-MS analyses of the essential oil identified 56 compounds accounting for 92.9%. The main components of the oil were δ-cadinene (16.5%), 6-hydroxytremetone (14.7%), epi-α-cadinol (12.0%), α-cadinol (8.8%), γ-cadinene (7.5%), 1- epi-cubenol (4.2%) and α-muurolene (3.0%). The essential oil exhibited antibacterial activities against five pathogenic strains as well as antifungal activities against two pathogenic fungi. The methanol extract showed antibacterial activity against two strains of Staphylococcus aureus and two pathogenic fungal strains. The main components isolated from the methanol extract were the antifungal 4-hydroxy-3-(isopenten-2-yl)-acetophenone, 6-hydroxytremetone, and tremetone. 6-Hydroxytremetone showed activity against all the fungal strains and one of the S. aureus strains assayed. Antioxidant and radical-scavenging properties of the methanol extract and essential oil were determined using the 2,2′-diphenyl-1-picrylhydrazyl assay and β-carotene bleaching (BCB) test. The methanol extract and the essential oil showed, respectively, moderate and weak antioxidant activity when compared to butylated hydroxytoluene.
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16

Cifuentes, Fredi, Javier Palacios, Jovan Kuzmicic, et al. "Vasodilator and hypotensive effects of pure compounds and hydroalcoholic extract of Xenophyllum poposum (Phil) V.A Funk (Compositae) on rats." Phytomedicine 50 (November 2018): 99–108. http://dx.doi.org/10.1016/j.phymed.2018.09.226.

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17

Calvo, Joel, Andrés Moreira-Muñoz, and Vicki A. Funk. "Taxonomic Revision of the Neotropical Genus Werneria (Compositae, Senecioneae)." Smithsonian Contributions to Botany, no. 111 (July 28, 2020): iv—123. http://dx.doi.org/10.5479/si.12728264.v1.

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Werneria is a neotropical genus belonging to the tribe Senecioneae (Compositae) that is mainly distributed through the highlands of the Andes. Only a single species, W. nubigena, has a trans-Andean distribution and reaches the Tacaná Volcano in southern Chiapas (Mexico). The traditional concept of Werneria was significantly narrowed at the end of the twentieth century after segregating several species and accommodating these mostly in the genus Xenophyllum. As currently circumscribed, it includes rosettiform or scapiform species characterized by usually displaying involucral bracts that are fused at the base, supplementary bracts commonly absent, usually radiate but sometimes discoid or disciform capitula, white or yellow ray corollas (when present), a balusterform filament collar, and style branches that are truncate and have a crown of sweeping hairs. The first modern, comprehensive revision of this genus recognizing 27 species is presented here. Werneria heteroloba and W. obtusiloba are synonymized with W. pinnatifida, and W. acerosifolia is synonymized with W. staticifolia. Moreover, five infraspecific names and three supraspecific names are synonymized. Detailed descriptions, distribution maps, photographs of living plants (except for W. cochlearis), and a list of all specimens examined are provided for each accepted species. An identification key to the species and a list of exsiccatae are also given. Eight species are illustrated, two of them for the first time. The following names are lectotypified: Werneria brachypappa Sch. Bip., W. caespitosa Wedd., W. calyculata Turcz., W. canaliculata Sch. Bip., W. carnulosa A. Gray, W. densa Benth., W. disticha Kunth, W. glaberrima Phil., W. glandulosa Wedd., W. heteroloba f. microcephala Rockh., W. mandoniana Wedd. ex Klatt, W. nubigena Kunth, W. nubigena var. caulescens Wedd., W. nubigena var. dombeyana Wedd., W. orbignyana Wedd., W. orbignyana var. breviradiata A. Gray, W. orbignyana var. longifolia Rockh., W. pectinata Lingelsh., W. plantaginifolia Wedd. ex Klatt, W. pygmaea var. caespitosa Wedd., W. rhizoma J. Rémy, W. solivifolia Sch. Bip., W. staticifolia Sch. Bip., W. staticifolia var. celmisioides Wedd., and W. villosa A. Gray. The names Werneria knocheae Perkins, W. minima Walp., W. minima var. pygmaea Walp., and W. stuebelii Hieron. are neotypified, and an epitype is designated for W. glandulosa. At the supraspecific level, the name Werneria sect. Integrifoliae Rockh. is typified.
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HAGBORG, ANDERS, LARS SÖDERSTRÖM, and MATT VON KONRAT. "Notes on Early Land Plants Today. 42. Validation of Nanomarsupella (Gymnomitraceae, Marchantiophyta)." Phytotaxa 112, no. 1 (2013): 16. http://dx.doi.org/10.11646/phytotaxa.112.1.3.

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Schuster (1978) described Marsupella xenophylla together with his new subgenus Nanomarsupella stating they were valid as generico-specific descriptions (ICN Art. 38.5; McNeill et al. 2012). However, as it was not described with a new genus but instead with new subgenus it is not validly described. However, the species is regarded as validly described here. Schuster (1996) then intended to elevate the subgenus to a genus but as the proposed basionym is not valid and no description was provided, his new genus is invalid as well as his proposed new combination of type species.
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Galán de Mera, Antonio, César Cáceres, and Adolfo González. "La vegetación de la alta montaña andina del sur del Perú." Acta Botanica Malacitana 28 (January 1, 2003): 121–47. http://dx.doi.org/10.24310/abm.v28i0.7271.

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RESUMEN. La vegetación de la alta montaña andina del sur del Perú. El presente artículo es un estudio fitosociológico de la vegetación de la alta montaña del sur del Perú (departamentos de Arequipa, Moquegua, Puno y Tacna). En base a la relación existente entre geomorfología, litología, altitud, pisos bioclimáticos, intervalos de precipitación, descripción bioclimática, y la combinación florística en determinados biotopos, fueron diferenciadas las comunidades de 9 clases (Anthochloo-Dielsiochloetea, Calamagrostietea vicunarum, Lantano-Chusqueetea, Lemnetea minoris, Not holaenetea niveae, Phragmito- Magnocaricetea, Plantaginetea australis, Planta gini rigidae-Di.slichietea muscoidis y Potametea). Como resultado, fueron descritos 2 nuevos órdenes (Calamagrostietalia nitidulae y Salpichroetalia glandulosae), 7 alianzas (Azorello-Festucion, Belloo-Salpichroion, Cha etantherion sphaeroidalis, Cortaderion jubatae, Fabianimi stephanii, Hypsello-Plantaginion y Nototrich ion obcuneatae), 10 asociaciones (Belloo- Dissanthelietum, Chersodomo-Valerianetum, Cortaderietuin jubatae, Diplostephio-Parastrephietum, Dunalio-Baccharidetum, Nototricho-Xenophylletum, Parastrephio-Festucetum, Ranunculetum limoselloidis, Stuckenietum punensis y Wernerio-Puccinellietum) y 16 subasociaciones y comunidades fragmentarias (comunidades basales, BC y comunidades derivadas. DC).Palabras clave. Vegetación, fitosociología, Sur Perú, Andes.ABSTRACT. The vegetation of the high Andean mountain of Southern Peru. In this paper, a phytosociological study about the vegetation of the mountain of Southern Peru (Arequipa, Moquegua, Puno and Tacna departments) is presented. In base of the relation between geomorphology, lithology, altitude, bioclimatic belts, precipitation intervals, bioclimatic description, and the floristic combination in specific biotopes, communities of 9 classes (Anthochloo-Dielsiochloetea, Calamagrostietea vicunarum, Lantano-Chusqueetea, Lemnetea minoris, Not holaenetea niveae, Phragmito-Magnocaricetea, Plantaginetea australis, Plantagini rigidae-Distichietea muscoidis and Potametea) were differenced. As result, 2 new orders (Calamagrostietalia nitidulae and Salpichroetalia glandulosae), 7 alliances (Azorello-Festucion, Belloo-Salpichroion, Chaetantherion sphaeroidalis, Cortaderion jubatae, Fabianion stephan ii, Hypsello-Plantaginion and Nototrichion obcuneatae), 10 associations (Belloo-Dissanthelietum, Che rsodomo-Valerianetum, Cortaderietumjubatae, Diplostephio-Parastrephietum, Dunalio-Baccharidetum, Nototricho-Xenophylletum, Paras. trephio-Fe.s. tucetum, Ranunculetum 1 imoselloidi s, Stuckenietum punensis and Wernerio- Puccinellienon) and 16 subassociations and phragmentary communities (basal communities, BC and derived communities, DC) are described.Key words. Vegetation, phytosociology, Southern Peru, Andes.
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Hind, D. J. Nicholas. "A new genus, Rockhausenia (Compositae: Senecioneae: Senecioninae)." Kew Bulletin, September 13, 2022. http://dx.doi.org/10.1007/s12225-022-10040-5.

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SummaryWerneria Kunth (Compositae: Senecioneae: Senecioninae) was effectively lectotypified by Rockhausen based on Werneria rigida. Two later lectotypifications have apparently been made, firstly (presumably by Cabrera) W. graminifolia and secondly (presumably by Funk) W. nubigena, yet the Index Nominum Genericorum website has no type stated. In removing W. rigida to the then newly recognised Xenophylllum, Funk moved Rockhausen’s type thereby creating a superfluous generic synonym. All species that Funk recognised in Xenophyllum are thus wernerias in the nomenclatural sense. Two species of Xenophyllum described since Funk’s revision are transferred to Werneria: X. funkianum and X. lorochaqui. The remainder of the species of Werneria are here placed in the new genus Rockhausenia (Compositae: Senecioneae: Senecioninae), a generitype is selected, and the corresponding 27 combinations are made at species level; the relevant subgeneric and sectional combinations are also made with two subgenera recognised, one with three sections. The confusion surrounding the generic name Oresigonia is commented on. A nomenclator of all relevant names in Rockhausenia, Werneria and Xenophyllum, is provided.
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Kelloff, Carol, Vicki A. Funk, Raymund Chan, and Joel Calvo. "A phylogenetic analysis and new delimitation of the Neotropical genus Werneria (Compositae, Senecioneae)." TAXON, December 5, 2024. https://doi.org/10.1002/tax.13294.

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AbstractThe genus Werneria was first described from six species found in the northern Andes in 1818 by Karl S. Kunth. By 1970 the number of species increased to around 50. Werneria is mostly distributed in the Andean highlands with a single species, W. nubigena, reaching as far as southern Chiapas, Mexico. Today, the genus Werneria encompasses 28 species with its center of diversity located in Peru and Bolivia. It is defined, in part, by having involucral bracts fused at least halfway and lacking a well‐developed calyculus at the base of the head. Further research indicated that there was considerable variability between Werneria species, and several species were transferred into the related genera Anticona, Misbrookea, Senecio, and Xenophyllum. Recently, it has been brought to light that the type of the genus Werneria is W. rigida, an effective lectotypification made by Rockhausen that previous authors working on this group overlooked. Since W. rigida has been accepted so far as a member of the genus Xenophyllum, Rockhausen's typification has resulted in: (1) Xenophyllum a superfluous name for Werneria; (2) Werneria a name to be applied to the species treated thus far as Xenophyllum; and (3) describing the new genus Rockhausenia to place the “traditional” Werneria species. Instead of proposing to conserve the name Werneria to avoid the aforementioned undesirable disruptions in the interest of nomenclatural stability, 30 new combinations have been made. Herein, we present the first comprehensive phylogenetic study based on ITS, trnL, ndhF, matK, and psbA sequence data involving 29 out of the 51 known species of Werneria and Xenophyllum (including the monotypic genus Misbrookea). The results indicate that: (1) the treatment of Misbrookea as a distinct genus is well supported; and (2) Werneria and Xenophyllum are each clearly not monophyletic. This, in addition to the fact that there are no diagnostic morphological synapomorphies to support either of the two genera, suggests treating them as members of a single genus. Moreover, the type of W. subg. Anactis appears to be nested in the genus Senecio. The three species belonging to this subgenus are accordingly transferred to Senecio. Finally, two subgenera are here proposed for facilitating species identification: (1) W. subg. Werneria, including species previously treated under Xenophyllum; (2) W. subg. Rockhausenia comb. & stat. nov., including the “traditional” Werneria species centered around W. nubigena.
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Cavalli, Daniele, Michelle Chang, Alasdair Milne, and William Morgan. "Xenophylum: Toward a Synthetic Cambrian Explosion." Antikythera Digital Journal, May 10, 2025. https://doi.org/10.1162/anti.5czl.

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This paper argues for a conceptual shift from biomimicry to xenomorphology in design, proposing a “synthetic Cambrian explosion” driven by techniques such as machine learning, robotics, and synthetic biology. Building on theoretical foundations from Bernard Stiegler’s notion of exosomatic evolution, mimetic theories, and assembly theory (developed by Michael Levin, Lee Cronin, and Sara Walker), we show how design has historically aligned with natural forms—a trend we term generalized biomimesis. While this biomimetic paradigm has yielded significant innovations, it constrains creativity by reinforcing nature as a universal model and moral ideal. By contrast, xenomorphology invites designers to explore genuinely alien morphologies unbound by terrestrial adaptation. Drawing on exemplars from the field of evolutionary computing, we argue that computational platforms and modular assembly enable vast new “morphospaces” decoupled from Earth’s evolutionary constraints. Ultimately, such a shift paves the way for new forms of anti-fragile design, where emergent resilience and novel behaviors come together to formulate new conceptions of intelligence and adaptation. Embracing xenomorphology opens a radical reimagining of design practice—one with the potential to shape the future of lifelike systems and our evolving relationship with technology.
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