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1

Lamsdell, James C. "The phylogeny and systematics of Xiphosura." PeerJ 8 (December 4, 2020): e10431. http://dx.doi.org/10.7717/peerj.10431.

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Xiphosurans are aquatic chelicerates with a fossil record extending into the Early Ordovician and known from a total of 88 described species, four of which are extant. Known for their apparent morphological conservatism, for which they have gained notoriety as supposed ‘living fossils’, recent analyses have demonstrated xiphosurans to have an ecologically diverse evolutionary history, with several groups moving into non-marine environments and developing morphologies markedly different from those of the modern species. The combination of their long evolutionary and complex ecological history a
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Shpinev, E. S., and D. V. Vasilenko. "First Fossil Xiphosuran (Chelicerata, Xiphosura) Egg Clutch from the Carboniferous of Khakassia." Paleontological Journal 52, no. 4 (2018): 400–404. http://dx.doi.org/10.1134/s0031030118040111.

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Shpinev, E. S. "New Data on Carboniferous Xiphosurans (Xiphosura, Chelicerata) of the Donets Coal Basin." Paleontological Journal 52, no. 3 (2018): 271–83. http://dx.doi.org/10.1134/s0031030118030127.

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Ballesteros, Jesús A., and Prashant P. Sharma. "A Critical Appraisal of the Placement of Xiphosura (Chelicerata) with Account of Known Sources of Phylogenetic Error." Systematic Biology 68, no. 6 (2019): 896–917. http://dx.doi.org/10.1093/sysbio/syz011.

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AbstractHorseshoe crabs (Xiphosura) are traditionally regarded as sister group to the clade of terrestrial chelicerates (Arachnida). This hypothesis has been challenged by recent phylogenomic analyses, but the non-monophyly of Arachnida has consistently been disregarded as artifactual. We re-evaluated the placement of Xiphosura among chelicerates using the most complete phylogenetic data set to date, expanding outgroup sampling, and including data from whole genome sequencing projects. In spite of uncertainty in the placement of some arachnid clades, all analyses show Xiphosura consistently ne
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5

Anderson, Lyall I., and Rachel A. Moore. "Bembicosoma re-examined: a xiphosuran from the Silurian of the North Esk Inlier, Pentland Hills, Scotland." Transactions of the Royal Society of Edinburgh: Earth Sciences 94, no. 3 (2003): 199–206. http://dx.doi.org/10.1017/s0263593300000614.

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ABSTRACTThe Silurian arthropod Bembicosoma pomphicus Laurie, 1899 is re-studied in relation to other Palaeozoic chelicerate taxa. All three known specimens of Bembicosoma originate from the Silurian (late Llandovery) Eurypterid Bed of the Gutterford Burn Flagstones, Reservoir Formation, Pentland Hills, Scotland. Bembicosoma is removed from its previous tentative assignment to Eurypterida and re-assigned to Xiphosura. A morphological reconstruction of this taxon is presented for the first time. This work continues a reappraisal of the systematics of a number of taxa that belong within the synzi
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Lamsdell, James C. "Revised systematics of Palaeozoic 'horseshoe crabs' and the myth of monophyletic Xiphosura." Zoological Journal of the Linnean Society 167, no. 1 (2012): 1–27. https://doi.org/10.1111/j.1096-3642.2012.00874.x.

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Lamsdell, James C. (2013): Revised systematics of Palaeozoic 'horseshoe crabs' and the myth of monophyletic Xiphosura. Zoological Journal of the Linnean Society 167 (1): 1-27, DOI: 10.1111/j.1096-3642.2012.00874.x, URL: http://dx.doi.org/10.1111/j.1096-3642.2012.00874.x
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7

Zhou, H., and Brian Morton. "The diets of juvenile horseshoe crabs, Tachypleus tridentatus and Carcinoscorpius rotundicauda (Xiphosura), from nursery beaches proposed for conservation in Hong Kong." Journal of Natural History 38, no. 15 (2004): 1915–25. https://doi.org/10.1080/0022293031000155377.

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Zhou, H., Morton, Brian (2004): The diets of juvenile horseshoe crabs, Tachypleus tridentatus and Carcinoscorpius rotundicauda (Xiphosura), from nursery beaches proposed for conservation in Hong Kong. Journal of Natural History 38 (15): 1915-1925, DOI: 10.1080/0022293031000155377, URL: http://dx.doi.org/10.1080/0022293031000155377
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8

ANDERSON, LYALL I., and PAUL A. SELDEN. "Opisthosomal fusion and phylogeny of Palaeozoic Xiphosura." Lethaia 30, no. 1 (2007): 19–31. http://dx.doi.org/10.1111/j.1502-3931.1997.tb00440.x.

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9

Chiu, Helen M. C., and Brian Morton. "The morphological differentiation of two horseshoe crab species, Tachypleus tridentatus and Carcinoscorpius rotundicauda (Xiphosura), in Hong Kong with a regional Asian comparison." Journal of Natural History 37, no. 19 (2003): 2369–82. https://doi.org/10.1080/00222930210149753.

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Chiu, Helen M. C., Morton, Brian (2003): The morphological differentiation of two horseshoe crab species, Tachypleus tridentatus and Carcinoscorpius rotundicauda (Xiphosura), in Hong Kong with a regional Asian comparison. Journal of Natural History 37 (19): 2369-2382, DOI: 10.1080/00222930210149753, URL: http://www.tandfonline.com/doi/abs/10.1080/00222930210149753
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10

Moore, Rachel A., Derek E. G. Briggs, Simon J. Braddy, and Jeffrey W. Shultz. "Synziphosurines (Xiphosura: Chelicerata) from the Silurian of Iowa." Journal of Paleontology 85, no. 1 (2011): 83–91. http://dx.doi.org/10.1666/10-057.1.

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A new genus and species of synziphosurine (Chelicerata) is described from the Silurian (Wenlock) Scotch Grove Formation Konservat-Lagerstätte in Clinton County, Iowa. Camanchia grovensis n. gen. n. sp. is characterized by a sub-triangular carapace, ten opisthosomal segments divided into a preabdomen of seven and a postabdomen of three, and a tuberculate ornament on the carapace and pleural margins. A single new specimen from the same location is assigned to Venustulus waukeshaensis, originally described from the Late Llandovery Waukesha Konservat-Lagerstätte of Wisconsin. A comparison of the m
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11

Bicknell, Russell D. C., Lisa Amati, and Javier Ortega-Hernández. "New insights into the evolution of lateral compound eyes in Palaeozoic horseshoe crabs." Zoological Journal of the Linnean Society 187, no. 4 (2019): 1061–77. http://dx.doi.org/10.1093/zoolinnean/zlz065.

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Abstract Vision allows animals to interact with their environment. Aquatic chelicerates dominate the early record of lateral compound eyes among non-biomineralizing crown-group euarthropods. Although the conservative morphology of lateral eyes in Xiphosura is potentially plesiomorphic for Euarthropoda, synziphosurine eye organization has received little attention despite their early diverging phylogenetic position. Here, we re-evaluate the fossil evidence for lateral compound eyes in the synziphosurines Bunodes sp., Cyamocephalus loganensis, Legrandella lombardii, Limuloides limuloides, Pseudo
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12

Шпинёв, Е. С., та Д. В. Василенко. "Первая ископаемая яйцекладка мечехвостов (Chelicerata, Xiphosura) из карбона Хакасии". Палеонтологический журнал, № 4 (2018): 48–52. http://dx.doi.org/10.1134/s0031031x18040116.

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13

Alberti, G., and H. H. Janssen. "On the fine structure of spermatozoa ofTachypleus gigas(Xiphosura, Merostomata)." International Journal of Invertebrate Reproduction and Development 9, no. 3 (1986): 309–19. http://dx.doi.org/10.1080/01688170.1986.10510207.

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14

MOHR, J. C. VAN DER MEER. "A NOTE ON TWO SPECIES OF MALAYSIAN KING-CRABS (XIPHOSURA)." Treubia 18, no. 1 (2025): 201–5. https://doi.org/10.14203/treubia.v18i1.2599.

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15

Anderson, Lyall I., and Carl Horrocks. "Valloisella lievinensis Racheboeuf, 1992 (Chelicerata; Xiphosura) from the Westphalian B of England." Neues Jahrbuch für Geologie und Paläontologie - Monatshefte 1995, no. 11 (1995): 647–58. http://dx.doi.org/10.1127/njgpm/1995/1995/647.

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16

MOORE, RACHEL A., SCOTT C. McKENZIE, and BRUCE S. LIEBERMAN. "A CARBONIFEROUS SYNZIPHOSURINE (XIPHOSURA) FROM THE BEAR GULCH LIMESTONE, MONTANA, USA." Palaeontology 50, no. 4 (2007): 1013–19. http://dx.doi.org/10.1111/j.1475-4983.2007.00685.x.

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17

Göpel, Torben, and Christian S. Wirkner. "An “ancient” complexity? Evolutionary morphology of the circulatory system in Xiphosura." Zoology 118, no. 4 (2015): 221–38. http://dx.doi.org/10.1016/j.zool.2014.12.004.

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18

Noah, Katherine E., Jiasheng Hao, Luyan Li, et al. "Major Revisions in Arthropod Phylogeny Through Improved Supermatrix, With Support for Two Possible Waves of Land Invasion by Chelicerates." Evolutionary Bioinformatics 16 (January 2020): 117693432090373. http://dx.doi.org/10.1177/1176934320903735.

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Deep phylogeny involving arthropod lineages is difficult to recover because the erosion of phylogenetic signals over time leads to unreliable multiple sequence alignment (MSA) and subsequent phylogenetic reconstruction. One way to alleviate the problem is to assemble a large number of gene sequences to compensate for the weakness in each individual gene. Such an approach has led to many robustly supported but contradictory phylogenies. A close examination shows that the supermatrix approach often suffers from two shortcomings. The first is that MSA is rarely checked for reliability and, as wil
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19

Lamsdell, James C. "Revised systematics of Palaeozoic ‘horseshoe crabs’ and the myth of monophyletic Xiphosura." Zoological Journal of the Linnean Society 167, no. 1 (2012): 1–27. http://dx.doi.org/10.1111/j.1096-3642.2012.00874.x.

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20

Selden, Paul A., and Derek J. Siveter. "The status of Bellinuroopsis Chernyshev, 1933, and Neobelinuropsis Eller, 1938 (Xiphosura, Bellinuroidea)." Journal of Paleontology 61, no. 6 (1987): 1285. http://dx.doi.org/10.1017/s0022336000029693.

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21

Шпинёв, Е. С. "НОВЫЕ ДАННЫЕ О КАМЕННОУГОЛЬНЫХ МЕЧЕХВОСТАХ (XIPHOSURA, CHELICERATA) ДОНЕЦКОГО УГОЛЬНОГО БАССЕЙНА, "Палеонтологический журнал"". Палеонтологический журнал, № 3 (2018): 49–62. http://dx.doi.org/10.7868/s0031031x18030078.

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22

Battelle, B. A. "The eyes of Limulus polyphemus (Xiphosura, Chelicerata) and their afferent and efferent projections." Arthropod Structure & Development 35, no. 4 (2006): 261–74. http://dx.doi.org/10.1016/j.asd.2006.07.002.

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23

Bicknell, Russell D. C., Lorenzo Lustri, and Tom Brougham. "Revision of “Bellinurus” carteri (Chelicerata: Xiphosura) from the Late Devonian of Pennsylvania, USA." Comptes Rendus Palevol 18, no. 8 (2019): 967–76. http://dx.doi.org/10.1016/j.crpv.2019.08.002.

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24

Ewington, DL, MJ Clarke, and MR Banks. "A Late Permian fossil horseshoe crab (Paleolimulus: Xiphosura) from Poatina, Great Western Tiers, Tasmania." Papers and Proceedings of The Royal Society of Tasmania 123 (1989): 127–31. http://dx.doi.org/10.26749/rstpp.123.127.

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25

Błażejowski, Błażej, Piotr Gieszcz, Daniel Tyborowski, Robert Siuda, and Jerzy Kwiatek. "ŻYCIE I ŚMIERĆ PÓŹNOJURAJSKICH SKRZYPŁOCZY (XIPHOSURA) Z OWADOWA-BRZEZINEK W ŚWIETLE INTERDYSCYPLINARNYCH BADAŃ PALEOŚRODOWISKOWYCH." Kosmos 68, no. 4 (2020): 535–41. http://dx.doi.org/10.36921/kos.2019_2565.

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Skrzypłocze (Xiphosura) są jednymi z najdziwniejszych istot w królestwie zwierząt. Grupa ta, współcześnie obejmująca cztery reliktowe gatunki, przetrwała właściwie niezmieniona od ordowiku (480 milionów lat), i była świadkiem masowych wymierań, stąd powszechnie używana jest jako przykład „żywych skamieniałości” lub „stabilomorfów”. Niniejsza praca poświęcona jest opisowi nowych i doskonale zachowanych skamieniałości skrzypłoczy z górnej jury (tyton) kamieniołomu Owadów-Brzezinki, który jest jednym z najważniejszych stanowisk paleontologicznych opisanych w ostatnim czasie w Polsce. Kamieniołom
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Obst, Matthias, Søren Faurby, Somchai Bussarawit, and Peter Funch. "Molecular phylogeny of extant horseshoe crabs (Xiphosura, Limulidae) indicates Paleogene diversification of Asian species." Molecular Phylogenetics and Evolution 62, no. 1 (2012): 21–26. http://dx.doi.org/10.1016/j.ympev.2011.08.025.

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27

LAMSDELL, JAMES C., JINZHUANG XUE, and PAUL A. SELDEN. "A horseshoe crab (Arthropoda: Chelicerata: Xiphosura) from the Lower Devonian (Lochkovian) of Yunnan, China." Geological Magazine 150, no. 2 (2012): 367–70. http://dx.doi.org/10.1017/s0016756812000891.

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AbstractA single specimen of a new species of the synziphosurine Kasibelinurus Pickett, 1993 is described from the Lower Devonian (Lochkovian) Xiaxishancun Formation of Yunnan Province, China. The new species, K. yueya sp. nov., extends the geographic extent of the family Kasibelinuridae from the Australian palaeocontinent to the South China palaeocontinent, and the stratigraphic range back some 50 Ma from Late to Early Devonian.
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Moore, Rachel A., Derek E. G. Briggs, and Christoph Bartels. "A new specimen ofWeinbergina opitzi (Chelicerata: Xiphosura) from the Lower Devonian Hunsriick Slate, Germany." Paläontologische Zeitschrift 79, no. 3 (2005): 399–408. http://dx.doi.org/10.1007/bf02991931.

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MOORE, RACHEL A., DEREK E. G. BRIGGS, SIMON J. BRADDY, LYALL I. ANDERSON, DONALD G. MIKULIC, and JOANNE KLUESSENDORF. "A NEW SYNZIPHOSURINE (CHELICERATA: XIPHOSURA) FROM THE LATE LLANDOVERY (SILURIAN) WAUKESHA LAGERSTÄTTE, WISCONSIN, USA." Journal of Paleontology 79, no. 2 (2005): 242–50. http://dx.doi.org/10.1666/0022-3360(2005)079<0242:anscxf>2.0.co;2.

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30

Haug, Carolin, and Marie A. I. N. Rötzer. "The ontogeny of Limulus polyphemus (Xiphosura s. str., Euchelicerata) revised: looking “under the skin”." Development Genes and Evolution 228, no. 1 (2018): 49–61. http://dx.doi.org/10.1007/s00427-018-0603-1.

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31

Novack-Gottshall, Philip M., and Roy E. Plotnick. "Correcting a 135-year error: Limulidae Leach, 1819 (Chelicerata, Xiphosura) is the proper authority, not Limulidae Zittel, 1885." Journal of Paleontology 95, no. 4 (2021): 886–87. http://dx.doi.org/10.1017/jpa.2021.23.

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The horseshoe crab Limulus polyphemus (Linnaeus, 1758) is a famous species, renowned as a ‘living fossil’ (Owen, 1873; Barthel, 1974; Kin and Błażejowski, 2014) for its apparently little-changed morphology for many millions of years. The genus Limulus Müller, 1785 was used by Leach (1819, p. 536) as the basis of a new family Limulidae and synonymized it with Polyphemus Lamarck, 1801 (Lamarck's proposed but later unaccepted replacement for Limulus, as discussed by Van der Hoeven, 1838, p. 8) and Xyphotheca Gronovius, 1764 (later changed to Xiphosura Gronovius, 1764, another junior synonym of Li
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32

Dunlop, Jason A., Marlene S. Compton, and Anja Friederichs. "An annotated catalogue of the horseshoe crabs (Xiphosura) held in the Museum für Naturkunde Berlin." Zoosystematics and Evolution 88, no. 2 (2012): 215–22. http://dx.doi.org/10.1002/zoos.201200018.

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Kwan, Billy K. Y., Alice K. Y. Chan, Siu Gin Cheung, and Paul K. S. Shin. "Marine microalgae as dietary supplements in the culture of juvenile Chinese horseshoe crabs,Tachypleus tridentatus(Xiphosura)." Aquaculture Research 48, no. 7 (2016): 3910–24. http://dx.doi.org/10.1111/are.13218.

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Shultz, Jeffrey W. "GROSS MUSCULAR ANATOMY OF LIMULUS POLYPHEMUS (XIPHOSURA, CHELICERATA) AND ITS BEARING ON EVOLUTION IN THE ARACHNIDA." Journal of Arachnology 29, no. 3 (2001): 283–303. http://dx.doi.org/10.1636/0161-8202(2001)029[0283:gmaolp]2.0.co;2.

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Baek, Su Youn, Eun Hwa Choi, Kuem Hee Jang, et al. "Complete Mitochondrial Genomes of Carcinoscorpius rotundicauda and Tachypleus tridentatus (Xiphosura, Arthropoda) and Implications for Chelicerate Phylogenetic Studies." International Journal of Biological Sciences 10, no. 5 (2014): 479–89. http://dx.doi.org/10.7150/ijbs.8739.

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36

Fahrein, Kathrin, Susan E. Masta, and Lars Podsiadlowski. "The first complete mitochondrial genome sequences of Amblypygi (Chelicerata: Arachnida) reveal conservation of the ancestral arthropod gene order." Genome 52, no. 5 (2009): 456–66. http://dx.doi.org/10.1139/g09-023.

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Amblypygi (whip spiders) are terrestrial chelicerates inhabiting the subtropics and tropics. In morphological and rRNA-based phylogenetic analyses, Amblypygi cluster with Uropygi (whip scorpions) and Araneae (spiders) to form the taxon Tetrapulmonata, but there is controversy regarding the interrelationship of these three taxa. Mitochondrial genomes provide an additional large data set of phylogenetic information (sequences, gene order, RNA secondary structure), but in arachnids, mitochondrial genome data are missing for some of the major orders. In the course of an ongoing project concerning
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37

Błażejowski, Błażej, Grzegorz Niedźwiedzki, Kamel Boukhalfa, and Mohamed Soussi. "Limulitella tejraensis, a new species of limulid (Chelicerata, Xiphosura) from the Middle Triassic of southern Tunisia (Saharan Platform)." Journal of Paleontology 91, no. 5 (2017): 960–67. http://dx.doi.org/10.1017/jpa.2017.29.

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AbstractNumerous well-preserved remains of a new limulid species from the Anisian-lower Ladinian (Middle Triassic) of the Tejra section of southern Tunisia are described. Comparisons are made with limulids from the Triassic deposits of Europe and Australia. The new specimens are congeneric with the type species ofLimulitella, but show some morphological differences. Here we describeLimulitella tejraensisnew species, a small limulid with semicircular prosoma, small and triangular opisthosoma, well-defined axial ridge, and pleurae along both ridges of the opisthosoma. The TunisianLimulitellafoss
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Kwan, Billy K. Y., Alice K. Y. Chan, Siu Gin Cheung, and Paul K. S. Shin. "Hemolymph quality as indicator of health status in juvenile Chinese horseshoe crab Tachypleus tridentatus (Xiphosura) under laboratory culture." Journal of Experimental Marine Biology and Ecology 457 (August 2014): 135–42. http://dx.doi.org/10.1016/j.jembe.2014.04.011.

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Kaiser, Dorkas, and Sabine Schoppe. "Postembryonic development of the Tri-spine Horseshoe Crab Tachypleus tridentatus (Merostomata: Xiphosura) in a nursery habitat in the Philippines." Journal of Threatened Taxa 10, no. 15 (2018): 12916–32. http://dx.doi.org/10.11609/jott.4125.10.15.12916-12932.

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Populations of the Tri-spine Horseshoe Crab Tachypleus tridentatus have dramatically decreased over their distribution range and conservation efforts are now crucial. The implementation of appropriate management strategies and stock assessment rely on accurate growth-rate estimates. The postembryonic development of the species in the tropics, however, is not elucidated. To provide the information needed to assess the demographics of juvenile populations and to judge the status of T. tridentatus in the Philippines, we conducted a mark-recapture experiment in a nursery habitat on Palawan Island.
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40

Chiu, Helen M. C., and Brian Morton. "The Behaviour of Juvenile Horseshoe Crabs, Tachypleus Tridentatus (Xiphosura), on a Nursery Beach at Shui Hau Wan, Hong Kong." Hydrobiologia 523, no. 1-3 (2004): 29–35. http://dx.doi.org/10.1023/b:hydr.0000033085.71861.63.

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Farley, Roger D. "Book gill development in embryos and first and second instars of the horseshoe crab Limulus polyphemus L. (Chelicerata, Xiphosura)." Arthropod Structure & Development 39, no. 5 (2010): 369–81. http://dx.doi.org/10.1016/j.asd.2010.04.001.

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Kwan, Billy K. Y., Alice K. Y. Chan, Siu Gin Cheung, and Paul K. S. Shin. "Responses of growth and hemolymph quality in juvenile Chinese horseshoe crab Tachypleus tridentatus (Xiphosura) to sublethal tributyltin and cadmium." Ecotoxicology 24, no. 9 (2015): 1880–95. http://dx.doi.org/10.1007/s10646-015-1524-7.

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43

Farley, Roger D. "Ultrastructure of book gill development in embryos and first instars of the horseshoe crab Limulus polyphemus L. (Chelicerata, Xiphosura)." Frontiers in Zoology 9, no. 1 (2012): 4. http://dx.doi.org/10.1186/1742-9994-9-4.

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44

Zhou, H., and Brian Morton. "The diets of juvenile horseshoe crabs,Tachypleus tridentatusandCarcinoscorpius rotundicauda(Xiphosura), from nursery beaches proposed for conservation in Hong Kong." Journal of Natural History 38, no. 15 (2004): 1915–25. http://dx.doi.org/10.1080/0022293031000155377.

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45

Lee, Christine N., and Brian Morton. "Experimentally derived estimates of growth by juvenile Tachypleus tridentatus and Carcinoscorpius rotundicauda (Xiphosura) from nursery beaches in Hong Kong." Journal of Experimental Marine Biology and Ecology 318, no. 1 (2005): 39–49. http://dx.doi.org/10.1016/j.jembe.2004.12.010.

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46

Aini, Naila Khuril, Erwyansyah Erwyansyah, Rahmat Kurnia, et al. "Two morphotypes of tri-spine horseshoe crab, Tachypleus tridentatus (Leach, 1819) (Xiphosura Limulidae) in Indonesia and implications for species identification." Biodiversity Journal 11, no. 2 (2020): 535–40. http://dx.doi.org/10.31396/biodiv.jour.2020.11.2.535.540.

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Hu, Menghong, Youji Wang, Sau Ting Tsang, Siu Gin Cheung, and Paul K. S. Shin. "Effect of starvation on the energy budget of two Asian horseshoe crab species: Tachypleus tridentatus and Carcinoscorpius rotundicauda (Chelicerata: Xiphosura)." Marine Biology 158, no. 7 (2011): 1591–600. http://dx.doi.org/10.1007/s00227-011-1672-0.

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Chiu, Helen M. C., and Brian Morton. "The morphological differentiation of two horseshoe crab species,Tachypleus tridentatusandCarcinoscorpius rotundicauda(Xiphosura), in Hong Kong with a regional Asian comparison." Journal of Natural History 37, no. 19 (2003): 2369–82. http://dx.doi.org/10.1080/00222930210149753.

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Chen, Y., CW Lau, SG Cheung, CH Ke, and PKS Shin. "Enhanced growth of juvenile Tachypleus tridentatus (Chelicerata: Xiphosura) in the laboratory: a step towards population restocking for conservation of the species." Aquatic Biology 11, no. 1 (2010): 37–46. http://dx.doi.org/10.3354/ab00289.

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Dolejš, Petr, and Kateřina Vaňousová. "A collection of horseshoe crabs (Chelicerata: Xiphosura) in the National Museum, Prague (Czech Republic) and a review of their immunological importance." Arachnologische Mitteilungen 49 (June 30, 2015): 1–9. http://dx.doi.org/10.5431/aramit4901.

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